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Biology (Undergraduate)

The Undergraduate Program in Biology offers courses at all levels of biological organization: molecular, cellular, organismal and population, and concentration programs with varying degrees of multidisciplinary or specialty foci.

There is a vast range of collaborative opportunities for independent study and research under the mentorship of the faculty. Faculty maintain research programs at the cutting edge of their fields including biochemistry, biophysics, cell biology, developmental biology, ecology, evolution, genetics, immunology, molecular biology, neurobiology, physiology, anatomy, artificial organs, microbiology, virology, experimental pathology, and pharmacology. The close administrative relationship with the Alpert Medical School offers additional opportunities for blending fundamental biology and clinical sciences in research projects at the Brown-affiliated hospitals.

The Office of Biology Undergraduate Education is a full service academic resource center, serving a key role for students of the life sciences here. The Office provides individualized advising for students at all levels; oversees student- faculty research collaborations, holds special events and maintains a variety of programs that support and enrich student life.

For further information, please visit: http://www.brown.edu/academics/biology/undergraduate-education/

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BIOL 0030. Principles of Nutrition.

Introduces the basic principles of human nutrition, and the application of these principles to the specific needs of humans, and the role of nutrition in chronic diseases. Provides an overview of the nutrients and their use by the human body. Also examines the role of nutrients in specific functions and disease states of the body. Not for biology concentration credit. Enrollment limited to 100.

Fall BIOL0030 S01 17086 Arranged (M. Flynn)
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BIOL 0040. Nutrition for Fitness and Physical Activity.

Reviews the role of nutrition in physical activity and health. It is designed to provide the student with the information and skills needed to translate nutrition and physical activity recommendations into guidelines for both the athlete for maximal performance and the non-athlete to improve both health and body weight. Students will learn the use of the energy yielding nutrition in physical activity and how food choices can influence both athletic performance and long-term health through the effect on risk factors for chronic diseases. Prerequisite: BIOL 0030. Enrollment limited to 20. Instructor permission required.

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BIOL 0060. Introduction to Human Physiology.

An introduction to human physiology aimed primarily at undergraduates who have minimal to no Biology background or who are not concentrating in biology. Acquire a basic understanding of the physiological mechanisms that allow for the running of each major organ systems. Topics include basic cardiovascular, respiratory, urinary, digestive, endocrine, and neuromuscular function, as well as aspects of reproduction and exercise physiology. Not for biology concentration credit. Lab.

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BIOL 0080. Biotechnology Management.

An examination of the pharmaceutical, biotechnology, and medical product industries: what they are, how they function, whence they originate, and various perspectives on why some succeed and others fail. Pathways from lab-bench to marketplace are described as are the pervasive influences of the FDA, patent office, and courts. Extensive reading; emphasis on oral presentation. Primarily intended for students planning a career in biomedical industry. Not for biology concentration credit. Students MUST register for the lecture section and the conference. Enrollment limited to 20.

Spr BIOL0080 S01 25530 T 4:00-6:30(16) (B. Bready)
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BIOL 0100. Living Biology at Brown and Beyond.

BIOL0100 is a first year seminar designed for aspiring biology program students. The course introduces students to the people and opportunities in Brown’s Biology Program and fosters academic and soft skill development. Students will practice a range of laboratory methods used in molecular biology while examining the ways in which those methods are used in research and in the development of medical treatments. Students will hone their writing skills, learn to effectively read primary STEM literature, and consider how various disciplines in Biology come together to tackle critical global problems such as global environmental change and emerging infectious disease. BIOL 0100 is best suited for students with minimal experience in the biological sciences.

Fall BIOL0100 S01 17052 W 3:00-5:30(10) (K. Monteiro)
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BIOL 0110. Fake Math: Analyzing the Misuse of Quantitative Concepts.

From the Scarecrow in the Wizard of Oz to the White House, quantitative errors are made and put forth. This course will present, discuss, and challenge students to analyze misused quantitative concepts. These errors will be taken from misapplied logic; inappropriate use of numbers; graphical inaccuracies and manipulations; cognitive illusions; self-fulfilling prophecies; the wrong-but-often-relied-upon Law of Small Numbers; conditional probability; correlation, confounding, and causality; hazards of data mining; circular reasoning; and basic misuses of statistics. Introductory concepts of classical logic, probability, and statistics will be presented and reviewed, and examples of their misuse will be discussed and evaluated.

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BIOL 0140A. Topics in Science Communications: Science Journalism Practicum.

Participants will understand how to read scientific research papers to interpret their findings and communicate these to a broader lay audience; analyze and understand best practices in science writing and the challenges of covering science for mass media; interviewing; fair and balanced coverage in reporting; give and receive peer feedback. Not for concentration credit in Biological Sciences programs. Enrollment limited to 10. Instructor permission required. S/NC

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BIOL 0140B. Communicating Science: Biological Illustration.

This Sophomore Seminar is an immersion practicum that conjoins art and science. Employing a series of techniques, students will learn the protocols of scientific rendering in an intensive hands-on approach. Field trips will include the RISD Nature Lab, the Brown Greenhouse, and John Hay collection of biomedical and botanical folios. Media will include graphite/ carbon dust; pen and ink (stipple, line); coquille board; scratch-board, colored pencil, watercolor and polymer clay. Course will be driven by project presentations focused on communicating science through illustration. Not for concentration credit in Biological Sciences programs. Instructor permission required; enrollment 10 students. 1/2 credit.

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BIOL 0140K. Conservation Medicine.

How have fruit bats contributed to the emergence of Nipah virus in Malaysia? Is an infectious cancer going to drive the Tasmanian Devil to extinction? Will a warmer world be a sicker world? We will consider these and additional topics at the intersection of global change biology and infectious disease emergence in this course. The course should be of interest to pre-med, general biology and environmental studies concentrators seeking interdisciplinary learning classroom experience. This will satisfy "Area 3" organismal biology concentration requirement for Biology/Health-Human Biology. Expected background: BIOL 0200 or equivalent placement. Enrollment limited to 12 sophomores. Instructor permission required.

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BIOL 0140T. Communicating Science Through Visual Media.

This class, offered jointly by professors at RISD and Brown, will explore the pedagogy and practice of using visual media to convey scientific concepts. The goal of this course will be to assess examples of existing material and create new animations/video that fill an educational need and make science engaging and accessible. Class time will be comprised of lectures, labs, screenings, discussions, critiques and exercises. After an introduction to teaching pedagogy and the basics of animation and visual design, small student teams will be paired with science faculty mentors to create videos and animations that explain scientific concepts.

Fall BIOL0140T S01 17055 W 1:00-6:00(08) (J. Stein)
Fall BIOL0140T L01 17056 Arranged (J. Stein)
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BIOL 0150A. Techniques and Analyses using DNA-Based Biotechnology.

Students will study and practice a range of methods used in molecular biology while examining the ways in which those tools are used in research and in the development of medical treatments. This experience, combined with the reading and discussion of selected papers from the primary literature, fosters development of a skill set critically important for the modern day biology student. Expected background: high school Biology course. Enrollment limited to 10 first year students. Instructor permission required. Half-credit course. S/NC.

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BIOL 0150B. Statistical Computing for Biology.

Modern biological research is a data rich endeavor, necessitating strong quantitative and computational skills to interpret the results of experimental and observational studies. In this course we will explore the application of statistics and modeling in biological research and environmental science. Topics covered will include basic probability, experimental design, sampling, hypothesis testing and mathematical models for prediction. No prior statistics knowledge is assumed. Enrollment limited to 10 first year and sophomore students. Instructor permission, based on a portfolio review. This is a half-credit course. S/NC

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BIOL 0150C. Introduction to Ethnopharmacology.

Plant secondary metabolites are currently the subject of much research interest when investigating new target compounds for potential medicine from natural products. New leads for drugs and phytomedicines from plants and plant parts have been increasing at a rapid rate especially by the pharmaceutical industry. Many plants have been selected and collected for their specific secondary compounds and healing powers by ethnobotanists in the field. Students will gain hands on experience identifying medicinal plants, laboratory equipment, sampling procedures, field tests and extraction techniques of secondary metabolites by high throughput screening methods. Enrollment limited to 10 FYS. Instructor permission required.

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BIOL 0150D. Techniques in Regenerative Medicine: Cells, Scaffolds and Staining.

Regenerative Medicine, also known as Tissue Engineering, is the process of creating living, functional tissues to repair or replace native tissue or organ functions that have been lost due to disease or congenital defects. As such, it is a prominent scientific discipline that can either "stand alone" or complement material-based research efforts in the areas of device design, drug delivery, diagnostics and pharmaceuticals. Students will develop proficiencies in basic cell culture techniques, early stage tissue regeneration strategies and histochemical characterization of mammalian cell constructs. Enrollment limited to 10 first year students. Instructor permission required. Half-credit course. S/NC.

Spr BIOL0150D S01 26543 M 10:00-12:00(03) (T. Achilli)
Spr BIOL0150D L01 26544 W 10:00-12:00 (T. Achilli)
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BIOL 0160. Plants, Food, and People.

Examines the selection, breeding, cultivation and uses of food plants. Discusses the effects on agriculture of pathogens, climate change, and loss of biodiversity. Considers whether enough food can be produced for a world population of potentially 10 billion, while sustaining biodiversity and environmental quality. Enrollment limited to 50.

Spr BIOL0160 S01 25375 TTh 10:30-11:50(09) (P. Heywood)
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BIOL 0170. Biotechnology in Medicine.

Introduces undergraduates to the main technological advances currently dominating the practice of medicine. Provides an overview of the objectives, techniques, and problems related to the application of biomedical technology to the diagnosis and treatment of disease and the contemporary health care industry. Topics include: pharmaceutical development and formulation; organ replacement by prosthesis and transplantation; medical imaging; tissue engineering, therapeutic cloning, regenerative medicine; stem cells; societal, economic, and ethical issues. This course does carry Biology concentration credit.

Fall BIOL0170 S01 18272 MWF 1:00-1:50(08) (T. Achilli)
Fall BIOL0170 S02 19277 Arranged (T. Achilli)
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BIOL 0180. The Biology of AIDS.

AIDS represents an example of the vulnerability of humans to new infectious agents. We will review some human infectious diseases including small pox yellow fever and influenza, and then explore AIDS/HIV. First characterized in 1981, AIDS became the leading cause of death in U.S. males aged 25-44 within a decade. We will examine what factors make HIV such a potent pathogen. The course is intended for students beginning in biology. Expected: BIOL 0200, or equivalent placement. This course does carry Biology concentration credit.

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BIOL 0190E. Botanical Roots of Modern Medicine.

This course will explore a variety of medicinal plants found throughout the world, the diverse cultures that use them in their daily lives and the scientific underpinnings of their medicinal uses. In conjunction with readings, students will gain a hands-on approach in lab, observing, identifying and growing these plants. Enrollment limited to 19. Students MUST register for the lecture section and the lab.

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BIOL 0190F. Darwinian Medicine.

Explores evolutionary explanations of why we get sick, and how this can shape, or misshape, our interpretations of medicine. Draws on evolutionary genetics, population biology, molecular biology and physiology. This course will build on evolutionary biology and then focus on disease processes such as infection, aging, cancer, allergy, diabetes, and obesity. Enrollment limited to 19 first year students.

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BIOL 0190P. Pride and Prejudice in the Development of Scientific Theories.

We will examine how the pace and shape of scientific progress is affected by the social/cultural context and the "personality" of the individual. We will look into how the interplay between society and the individual affects how scientific theories arise, are presented, are debated and are accepted. The course will initially focus on Charles Darwin and his theory of Natural Selection using the biography of Adrian Desmond and James Moore, "Darwin: The Life of a Tormented Evolutionist." Enrollment limited to 19 first year students.

Fall BIOL0190P S01 16625 TTh 2:30-3:50(12) (S. Helfand)
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BIOL 0190Q. Climate Change and Species Extinction.

In this course students will go beyond the headlines and delve into the science to explore the impact of climate change on species extinction. Students will explore the integration of science and technology through traditional textbooks, primary literature, open source databases, simulations, and discussions. Students will investigate the impact of climate change on species distribution, ecology, and behavior through interactive, inquiry-based, collaborative classroom investigations. Students will learn to integrate information from a variety of sources and disciplines and share their ideas through classroom discussion, written assignments, and oral presentations. Enrollment limited to 19 first year students.

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BIOL 0190R. Phage Hunters, Part I.

A research-based lab class for freshmen; both semesters are required in the sequence. Students will isolate and characterize a bacteriophage viruses found in the soil. Lab work includes isolation and purification of your own phage, DNA isolation and restriction mapping, and EM characterization of your phage. Several phages will be selected for genome sequencing, and will be annotated during the second semester. Instructor permission required. Admittance based on review of applications in the first class. Limited to 19 freshmen.

Fall BIOL0190R S01 16546 T 4:00-6:30(07) (S. Taylor)
Fall BIOL0190R L01 16547 M 1:00-4:00 (S. Taylor)
Fall BIOL0190R L02 16548 W 1:00-4:00 (S. Taylor)
Fall BIOL0190R L03 16549 F 1:00-4:00 (S. Taylor)
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BIOL 0190S. Phage Hunters, Part II.

A research-based laboratory/class for freshmen; both semesters are required. Students will isolate and characterize a bacteriophage viruses found in the soil. Lab work includes isolation and purification of your own phage, DNA isolation and restriction mapping, and EM characterization of your phage. Several phages will be selected for genome sequencing over Winter Recess, and annotated in the spring. One hour of lecture/discussion, and 3 hours lab per week. Expected: AP Biology or equivalent, HS chemistry, and permission of the instructor. Students are expected to take fall and spring courses in the sequence. Enrollment limited to 19 first-year students. Instructor permission.

Spr BIOL0190S S01 25125 Th 4:00-6:30(16) (S. Taylor)
Spr BIOL0190S S01 25125 T 4:00-6:30(16) (S. Taylor)
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BIOL 0190T. Bioinformatics: A Practical Introduction.

The amount of biological sequence data has grown at an exponential pace and spurred the development of computational tools that allow biologists to use this information. Students will become familiar with useful bioinformatics tools used by researchers. The course will introduce concepts of information transfer in biological molecules, storage in public databases, and how to use tools to access this information and organize it meaningfully. We will explore tools for studying whole genomes, including high-throughput sequencing data to assemble genomes and mapping subsets. Students will gain hands-on experience using these tools. Expected: AP credit or equivalent placement for BIOL 0200.

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BIOL 0190U. The Lives of Plants.

This course examines the lives of plants through their development, structure, function, reproduction, and responses to environmental conditions. Enrollment limited to 19 first year students.

Spr BIOL0190U S01 25376 F 3:00-5:30(15) (P. Heywood)
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BIOL 0200. The Foundation of Living Systems.

A broad overview of biological systems, emphasizing patterns and processes that form the basis of life. Explores essentials of biochemistry, molecular, and cellular biology and their relationship to the larger issues of ecology, evolution, and development. Examines current research trends in biology and their influence on culture. Appropriate for all students interested in biology. Serves as a gateway course to much of the intermediate and advanced curriculum. Placement tests are offered (contact Jody_Hall@brown.edu); AP scores of 4 or 5 are equivalent to BIOL 0200, and place a student out of this course. Students will be assigned to a lab time during the second week of class.

Spr BIOL0200 S01 25377 MWF 11:00-11:50(04) (J. Stein)
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BIOL 0210. Diversity of Life.

This course will explore biological diversity – the number of taxa, and the functions, and processes that support life – from the perspectives of ecology and evolutionary biology. It will draw on examples and case studies from the geological record, functional morphology, the evolution of organ systems in vertebrates, genomics, behavior and sexual selection in birds and invertebrates. Overarching themes will emphasize that taxonomic diversity is an emergent property of complex life on Earth, and the importance of diversity of biological functions and processes in generating and maintaining taxonomic diversity. The course is open to all students.

Fall BIOL0210 S01 16870 MWF 11:00-11:50(16) (J. Kellner)
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BIOL 0280. Biochemistry.

Lectures and recitation sections explore the mechanisms involved in the principles of macromolecular structure and function, the organization and regulation of pathways for intermediary metabolism, and the transfer of information from genes to proteins.

Spr BIOL0280 S01 25378 TTh 1:00-2:20(08) (A. Deaconescu)
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BIOL 0285. Inquiry in Biochemistry: From Gene to Protein Function.

In this inquiry-based research course, students work in teams to formulate and test a hypothesis about how a change in genetic sequence affects enzyme function. Students will cultivate skills in scientific visualization, experimental design, data analysis, and laboratory techniques in molecular biology and biochemistry. In discussion, students will learn scientific writing through peer editing and iterative revisions to write a full scientific paper. This course is WRIT designated and will prepare students for writing an honors thesis. Expected: Students have previously taken or be concurrently enrolled in BIOL 0280. Enrollment in one lab section and one discussion section is required.

Spr BIOL0285 S01 25391 M 1:00-5:00(06) (A. Sobieraj)
Spr BIOL0285 S02 25393 Th 2:30-6:30(11) (A. Sobieraj)
Spr BIOL0285 C01 25392 T 4:00-5:00 (A. Sobieraj)
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BIOL 0300. Endocrinology.

A basic examination of endocrinology with emphasis on hormone biosynthesis, mechanism of action, physiological roles, and endocrine pathology. Topics include: mechanism of action of steroid, amine, and peptide hormones; neuroendocrinology; reproductive endocrinology; and endocrinology of metabolism and calcium homeostasis. It is expected that students have taken BIOL 0200 (or equivalent) and CHEM 0350.

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BIOL 0320. Vertebrate Embryology.

Introduction to the developmental anatomy of vertebrate embryos, including humans, in an evolutionary context, through lecture, discussion and microscope slide study. Gametogenesis through germ layers and their organ system derivatives. Expected: BIOL 0200, or equivalent placement, or AP Biology score of 4 or 5. Limited to 18 freshmen and 18 sophomores. Students MUST register for the lecture section and the lab.

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BIOL 0350. The Fossil Record: Life through Time on Earth.

Course is designed for students with prior background in geology or evolutionary biology and who want to learn more about the fossil record, the origins of modern biodiversity and ecosystem structure, and interaction between organisms, and the geological and chemical cycles on the Earth. Lectures will cover major time periods during which animals and plants lived, as well as focusing on major transitions in the evolution of life on Earth. This course will fulfill requirements in both the geology/biology and evolutionary biology concentrations. Expected: BIOL 0210, GEOL 0240 or equivalent. Instructor permission, enrollment limited to 20 sophomores/juniors; register for course/lab.

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BIOL 0380. The Ecology and Evolution of Infectious Disease.

Infectious diseases remain among the leading causes of death worldwide, and this burden is disproportionately borne by children living in low- and middle-income countries. Thus management of infectious disease remains a critical intellectual challenge in the 21st century. This course will develop and apply ecological and evolutionary theory to infectious microbes (and their hosts) via the detailed examination of a number of case studies. This will be accomplished by a combination of lectures, discussions, and readings drawn mainly from the primary literature. Assessment will be based on biweekly problem sets, two midterms and one final exam. Expected: BIOL 0200, BIOL 0210 or equivalent.

Fall BIOL0380 S01 16659 MWF 10:00-10:50(14) (K. Witt Dillon)
Fall BIOL0380 C01 16660 M 6:00-6:50 (K. Witt Dillon)
Fall BIOL0380 C02 16661 M 7:00-7:50 (K. Witt Dillon)
Fall BIOL0380 C03 16662 T 8:00PM-8:50PM (K. Witt Dillon)
Fall BIOL0380 C04 16663 W 7:00-7:50 (K. Witt Dillon)
Fall BIOL0380 C05 16664 Th 8:00PM-8:50PM (K. Witt Dillon)
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BIOL 0390. Vertebrate Evolution and Diversity.

An overview of vertebrate evolution that not only covers historical events, but also introduces various scientific concepts and modes of thought. Topics include past and present biodiversity, convergent evolution, biogeography, competition, continental drift, climatic change over time, the notion of evolution as progress, and a whole-animal approach to understanding evolutionary events. Enrollment limited to 50.

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BIOL 0400. Biological Design: Structural Architecture of Organisms.

Many questions about the workings of living creatures can be answered by joining math, physics, and biology. We will identify basic physical science concepts that help biologists understand the structure and function of animals, plants, and microorganisms, and use these to study how the physical world constrains and facilitates the evolution of the extraordinary design and diversity of organisms. For first and second year students; others by permission. Recommended background: BIOL 0200, or equivalent. Enrollment limited to 40. Instructor permission required.

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BIOL 0410. Invertebrate Zoology.

A survey of invertebrate animals emphasizing evolutionary patterns and ecological relationships. Functional morphology, physiology, reproduction, development, and behavior of invertebrates will be examined. Laboratory exercises and two separate day-long field trips provide firsthand experience with the animals. Expected: BIOL 0200 or equivalent. Enrollment limited to 44. Students MUST register for the lecture section and a lab.

Fall BIOL0410 S01 16876 TTh 9:00-10:20(05) (P. Ewanchuk)
Fall BIOL0410 L01 16877 W 1:00-3:50 (P. Ewanchuk)
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BIOL 0415. Microbes in the Environment.

Examines the diversity of microbial life in the environment. Surveys key services that microbes perform on land and sea, including biodegradation of contaminants in the environment and ecosystem processes related to climate change. Examines biological interactions of symbioses, quorum sensing, and antibiotic production in a ecological context. Explores the genomic mechanisms explaining phylogeny and life history strategies in microbes. Demonstrate knowledge of the diversity of microbes in the environment and benefits in an ecological/evolutionary context. Lecture based, two fieldtrips to expand appreciation for microbial ecology. BIOL 0200 or equivalent placement; CHEM 0330. Enrollment limited to 20 sophomores, juniors and seniors.

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BIOL 0420. Principles of Ecology.

The principles, concepts, and controversies involved in the study of the distribution and abundance of plant and animal populations and their integration into natural communities. Emphasizes interactions among organisms and the hierarchical nature of ecological processes affecting individuals, populations, and communities. Expected: BIOL 0200 (or equivalent) and MATH 0090. Lectures and weekly discussion.

Spr BIOL0420 S01 25199 TTh 9:00-10:20(05) (J. Witman)
Spr BIOL0420 C01 25200 T 5:30-7:00 (J. Witman)
Spr BIOL0420 C02 25201 W 1:00-2:30 (J. Witman)
Spr BIOL0420 C03 25202 W 4:30-6:00 (J. Witman)
Spr BIOL0420 C04 25203 Th 4:30-6:00 (J. Witman)
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BIOL 0430. The Evolution of Plant Diversity.

Examines the evolutionary history of plants from a phylogenetic perspective. Introduces the science of phylogenetics - how to infer phylogenies and how to use them to understand organismal evolution. Highlights major trends in plant evolution over the past 400 million years. Lectures survey major plant lineages, with special focus on flowering plants. Weekly labs, field trips, and assignments stress basic plant anatomy and morphology, identification, and learning the local flora. Expected: BIOL 0200 (or equivalent placement).

Spr BIOL0430 S01 25204 TTh 9:00-10:20(05) (R. Kartzinel)
Spr BIOL0430 L01 25206 Th 1:00-4:00 (R. Kartzinel)
Spr BIOL0430 L02 25207 Arranged (R. Kartzinel)
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BIOL 0440. Inquiry in Plant Biology: Analysis of Plant Growth, Reproduction and Adaptive Responses.

This course focuses on what plants do and how they do it. Introduces the biology of plants, their growth and development, structural features, and their cellular and organismal responses to key stimuli. Examines physiological, reproductive and developmental strategies throughout the plant life cycle and in relation to environmental challenges. During laboratory section meetings, students pursue inquiry-based group research projects addressing novel questions about mechanisms that control plant growth and development. Laboratory section is required. Prerequisites: One Brown course with laboratory section in either Biology or Chemistry. Enrollment limited to 24 students.

Spr BIOL0440 S01 25394 TTh 10:30-11:50(09) (A. DeLong)
Spr BIOL0440 L01 25395 W 2:00-4:50 (A. DeLong)
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BIOL 0450. Evolutionary Behavioral Ecology.

An exploration of the ecological and evolutionary principles that define animal behavior in the natural world. We will focus mostly on the field of behavioral ecology, including important phenomena like avoiding predators, obtaining food, finding mates, producing offspring, living in groups, and solving problems. In doing so, we will combine ideas and principles from the disciplines of ecology, systematics, physiology, and economics.

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BIOL 0455. Coastal Ecology and Conservation.

Will enable to students to master fundamental ecological concepts and understand how this knowledge can be used to inform coastal conservation and management. Case studies from New England and elsewhere, field trips to rocky shores, salt marshes and coastal ecosystems enable students to develop scientific skills and experience the challenges of coastal conservation science. The course is aimed at freshmen and sophomores. Expected background: BIOL 0200 or equivalent placement. Enrollment limited to 10 students, and written permission required. Email (Mark_Bertness@brown.edu) to receive course application (due May 1). Admitted students register for the course in September.

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BIOL 0460. Insect Biology.

Focuses on characteristics that make insects unique and why more insect species have been described than all other organisms combined; the opportunity to investigate diversity and adaptation; their abundance, small size, and short lifespans; their importance as agents of biocontrol pollination, agricultural pests, and disease vectors. Expected: BIOL 0200 or equivalent. Enrollment limited to 20. Students MUST register for lecture AND lab. Primarily for freshmen and sophomores.

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BIOL 0470. Genetics.

Genetic phenomena at the molecular, cellular, organismal, and population levels. Topics include transmission of genes and chromosomes, mutation, structure and regulation of the expression of the genetic material, elements of genetic engineering, and evolutionary genetics. One laboratory session and one discussion session per week. (Students should not plan to take BIOL 0470 after 1540.) Expected: BIOL 0200 (or equivalent placement). Students will be assigned to Lab sections the first week of class. If you plan to take BIOL 470 but are also considering registering for a course that conflicts with the BIOL 470 lecture period, you may be eligible to do so by enrolling in S02, an alternative section of BIOL 470. Enrollment in this section allows students asynchronous engagement with lectures which are recorded and posted to the course Canvas page at the end of each class period. In-person participation in laboratory meetings and exams, however, are still required.

Fall BIOL0470 S01 16626 TTh 10:30-11:50(13) (M. Johnson)
Fall BIOL0470 S02 19154 Arranged (M. Johnson)
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BIOL 0480. Evolutionary Biology.

A broad introduction to the patterns and processes of evolution at diverse levels of biological organization. Topics covered include natural selection, adaptation, speciation, systematics, macroevolution, mass extinction events, and human evolution. Students will be given the opportunity to do their 23andMe ancestry analyses as a means of integrating the topics that span genomics to human variation. Weekly discussion sections involve active learning simulations and discussions of papers from the primary literature. Occasional problem sets involve computer exercises with population genetics and phylogeny reconstruction. Expected: BIOL 0200 (or equivalent placement).

Fall BIOL0480 S01 16665 MWF 9:00-9:50(09) (D. Rand)
Fall BIOL0480 S02 19278 Arranged (D. Rand)
Fall BIOL0480 C01 16666 T 4:00-5:20 (D. Rand)
Fall BIOL0480 C02 16667 T 5:30-6:50 (D. Rand)
Fall BIOL0480 C03 16668 W 1:00-2:20 (D. Rand)
Fall BIOL0480 C04 16669 W 2:30-3:50 (D. Rand)
Fall BIOL0480 C05 16670 Th 3:00-4:20 (D. Rand)
Fall BIOL0480 C06 16671 Th 5:00-6:20 (D. Rand)
Fall BIOL0480 C07 19187 Th 10:30-11:50 (D. Rand)
Fall BIOL0480 C08 19188 Arranged (D. Rand)
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BIOL 0495. Statistical Analysis of Biological Data.

A first course in probability distributions and the use of statistical methods for biological data. Topics covered will include describing data, statistical inference (hypothesis tests and confidence intervals), analyzing associations, and methods for categorical data (contingency tables and odds ratios). Methods will be applied to data drawn from areas of biological inquiry. For statistics or related science credit in Biology programs. Expected background: BIOL 0200 or equivalent, math equivalent to MATH 0100. This course is for related science credit only in Biological Sciences concentration programs. Enrollment limited: 40 undergraduates-20 juniors and 20 sophomores. Registration for seniors requires permission from the instructor.

Spr BIOL0495 S01 25346 TTh 2:30-3:50(11) (S. Ramachandran)
Spr BIOL0495 C01 25347 M 6:00-8:00PM (S. Ramachandran)
Spr BIOL0495 C02 25348 T 8:00PM-10:00PM (S. Ramachandran)
Spr BIOL0495 C03 25349 W 6:00-8:00PM (S. Ramachandran)
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BIOL 0500. Cell and Molecular Biology.

This course examines the structure and function of the basic unit of an organism, the cell. An experimental approach is used to examine cellular functions, ranging from gene transcription, cell division and protein secretion, to cell motility, and signal transduction. Relevance to health and disease will be considered. Expected: BIOL 0200 (or equivalent advanced placement biology 4/5).

Spr BIOL0500 S01 25396 MW 3:00-4:20(10) (G. Valdez)
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BIOL 0510. Introductory Microbiology.

Introduces role of microbes in our understanding of biology at the cellular and molecular level. Focuses on microbial significance for infectious disease, public health, genetics, biotechnology, and biogeochemical cycles. Laboratory involves basic microbiological techniques and selection and manipulation of microbes. Expected: BIOL 0200 (or equivalent placement). Students MUST register for the lecture section, conference, and the lab. Enrollment limited to 108.

Spr BIOL0510 S01 25129 MWF 1:00-1:50(06) (P. Belenky)
Spr BIOL0510 C01 25132 MWF 2:00-2:50 (P. Belenky)
Spr BIOL0510 L01 25140 T 9:00-11:50 (A. Dugan)
Spr BIOL0510 L02 25141 T 1:00-3:50 (A. Dugan)
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BIOL 0530. Principles of Immunology.

Introduction to experimental and theoretical foundations of immunology. Focuses on concepts, landmark experiments and recent advances. Topics include innate and adaptive immunity; structure/function of antibody molecules and T cell receptors; regulation of immune responses through cellular interactions. Applications of concepts to medically significant issues (vaccines, transplantation, inflammation, autoimmunity, cancer, HIV/AIDS) are discussed. Interpretative analysis of experimental data is emphasized. Expected background: BIOL 0200 or equivalent placement credit.

Fall BIOL0530 S01 16550 TTh 2:30-3:50(12) (A. Dugan)
Fall BIOL0530 C01 16551 Su 2:00-2:50 (A. Dugan)
Fall BIOL0530 C02 16552 M 6:00-6:50 (A. Dugan)
Fall BIOL0530 C03 16553 M 7:00-7:50 (A. Dugan)
Fall BIOL0530 C04 16554 M 8:00PM-8:50PM (A. Dugan)
Fall BIOL0530 C05 16555 T 8:30PM-9:20PM (A. Dugan)
Fall BIOL0530 C06 16556 W 6:00-6:50 (A. Dugan)
Fall BIOL0530 C07 16557 W 7:00-7:50 (A. Dugan)
Fall BIOL0530 C08 16558 F 6:00-6:50 (A. Dugan)
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BIOL 0600. Genetic Screening in Model Organisms.

Using gene silencing (RNAi) in the nemotode C. elegans, students will identify genetic modifiers of proteins with roles in aging by reverse genetics. Analyzing the effect of knocking down genes on the level of aging-related proteins tagged with fluorophores (GFP, RFP, etc.). Students will use function-specific RNAi libraries (transcription factors, kinases, etc) established in our lab. Students will evaluate the effect of genetic modifiers on proteostasis and lifespan, also familiarize C. elegans work and appreciating the use of model organisms, the students will master microscopy, genetic crosses, gene silencing, and molecular and biochemical readout assays such as qPCR and immunoblotting. Expected: BIOL 0200 or equivalent.

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BIOL 0800. Principles of Physiology.

Introduction to the function and integration of organ systems with an emphasis on human physiology. Includes basic concepts in cell and organ system physiology as well as fundamentals of modern trends in physiological science. Emphasizes the application of physical and chemical principles to organ function at both the cellular and systemic levels. Expected: BIOL 0200 or equivalent.
If you have a class conflict during the lecture time, there is an option to take the lecture asynchronously and the labs and exams synchronously. Please review the requirements for Section 02 and if you have a class conflict and are able to satisfy the synchronous requirements you may be able to take the course. The requirements are in-person lab offered at various times during the week and taking the 3 exams on the same day they are scheduled.

Fall BIOL0800 S01 16957 TTh 10:30-11:50(13) (J. Stein)
Fall BIOL0800 S02 18943 Arranged (J. Stein)
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BIOL 0860. Diet and Chronic Disease.

This course addresses the relationship of food to the development and treatment of chronic diseases. Chronic diseases discussed are obesity, dyslipidemia/heart disease, diabetes mellitus, cancers and osteoporosis. Dietary recommendations for these diseases are critically assessed. Geared toward students interested in nutrition, medicine, and public health. Prerequisites: BIOL 0030, plus permission of the instructor. Enrollment limited to 20.

Spr BIOL0860 S01 25508 T 4:00-6:30(16) (M. Flynn)
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BIOL 0920A. Controversies in Medicine.

Why and how do controversies in medicine emerge at specific moments in time? Why do scientists come to different conclusions based on the same data? Does it matter how we interpret controversies? This sophomore-level seminar critically analyzes contemporary controversies in medicine and public health. Using a case study approach, we will examine the social and political assumptions that inform important controversies. Questions related to the relationship between science, media, activism, and health inequality will be woven into the case studies. Enrollment limited to 20 sophomores. (For theme, not biology, credit in Health and Human Health and Biology only.)

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BIOL 0940A. Viral Epidemics.

This sophomore seminar will examine epidemics (outbreaks) of viral infections from a historical perspective. We will also cover current literature and up to the minute news accounts of infectious disease related outbreaks occurring around the globe. The major focus will be on virus related diseases but any microbial outbreak in the news will be explored. The seminar will cover basic aspects of microbial pathogenesis so students can gain an appreciation of microbial host interactions. Essential writing skills will also be developed. Enrollment limited to 20 sophomore students.

Fall BIOL0940A S01 16649 Th 4:00-6:30(04) (W. Atwood)
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BIOL 0940B. Sophomore Seminars in Biology: Life in a Shell.

This Sophomore seminar is an examination of broad themes in whole animal physiology with an emphasis on environmental adaptations. The foundation of the course will be the instructor’s recent book “Life in a Shell: A Physiologist’s View of Turtle.” A consideration of this iconic animal’s novel biological traits will lead into comparisons with our own biology and that of other animals. Topics: respiration, circulation, metabolic rate, buoyancy control, overwintering, migration, reproduction, and bone structure and function. Relevant original research papers will be used. Mandatory S/NC; enrollment of 20 students; override required. Expected: BIOL 0200 or equivalent placement credit.

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BIOL 0940C. Sophomore Seminar: Insect Biology.

Focuses on characteristics that make insects unique and why more insect species have been described than all other organisms combined; the opportunity to investigate diversity and adaptation; their abundance, small size, and short lifespans; their importance as agents of biocontrol pollination, agricultural pests, and disease vectors. Expected: BIOL 0200 or equivalent. Enrollment limited to 12 sophomores only. Students MUST register for lecture AND lab.

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BIOL 0940D. Rhode Island Flora: Understanding and Documenting Local Plant Diversity.

This Sophomore Seminar focuses on species level identification of plants in Rhode Island and will cover the dominant plant species in each of the state’s main habitats including coastal wetlands and uplands, freshwater wetlands, peatlands, upland forests, and disturbed areas. Students will learn to identify plants using online interactive keys as well as more technical dichotomous keys and will also cover basic ecological processes in each habitat including the interaction of soils, geology, and hydrology. Materials related to plant morphology, plant taxonomy, plant evolution, understanding phylogenetic trees, and botanical illustration. Instructor permission required.

Fall BIOL0940D S01 16871 F 1:00-5:00(11) (R. Kartzinel)
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BIOL 0940E. Precision Medicine or Privileged Medicine? Addressing Disparities in Biomedical Research.

This course examines the biomedical research behind precision medicine, disparities in the inclusiveness of this research, and implications of these disparities for the relevance of precision medicine innovations for people and places in Rhode Island. We will focus on these four questions: What new knowledge is making precision medicine possible? Who has been the focus of the biomedical research generating this knowledge, and why? How might inclusiveness of this research impact healthcare disparities in Rhode Island? What is needed to improve the design and outcomes of precision medicine research so that it provides benefits and mitigates harms for all?

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BIOL 0940G. Antibiotic Drug Discovery: Identifying Novel Soil Microbes to Combat Antibiotic Resistance.

This is a Course-based Undergraduate Research Experience (CURE) class that will provide students with the chance to propose, design and conduct their own research projects. Antibiotic resistance is a major global health threat. Pharmaceutical companies are less likely to fund research and development of new antibiotics due to their relative low profitability. As bacteria become more resistant to antibiotics, it is critical that we have a robust pipeline ready to combat these pathogens. The main focus of the course will be for students to discover new antibiotics in soil bacteria that can be used to treat infectious disease.

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BIOL 0945. Toolbox for Scientific Research.

Why is scientific research important? What is the scientific method? What are hypotheses and theories? How do scientists identify research questions, design experiments, fund research, and communicate results? This sophomore seminar is designed for students who want to understand and engage in scientific research in biology. Through active learning seminars, group discussions, and meetings with scientists, students will gain a deeper understanding and an appreciation for the principles, practice, and culture of scientific research. The course will also help develop practical and transferable skills to succeed in research and give students an opportunity to connect with research groups at Brown.

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BIOL 0946. Research Design + Quantitative Methods for the Health Sciences.

This is a new course offered for students in one of Brown University's Biology concentrations that provides the skills necessary to identify a clinical research question from a case scenario, design a research design to answer a research question, identify the appropriate quantitative methods for the design, and interpret the results. These skills translate to almost any research experience, including honors theses and future endeavors. Students will also learn how to publish in oral and written formats across the health sciences disciplines.

Spr BIOL0946 S01 26808 W 3:00-5:30(10) (K. Monteiro)
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BIOL 0960. Independent Study in Science Writing.

BIOL 0960 (fall/spring) is a half credit Independent Study in Science Writing course incorporating a nontechnical science journalism component into the Biology curriculum. Assignments may include investigative or analytical reviews, or feature articles on ethical or social impacts of new discoveries in the biological sciences. BIOL 0960 requires the submission of a formal project proposal completed collaboratively by the student and faculty mentor (see the Biology Undergraduate Education research page for details). BIOL 0960 is not for concentration credit in the biological sciences programs.

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BIOL 1000. Quality Improvement Science and Application.

Have you ever set a goal, tried to accomplish it, failed, adjusted your routine, and tried again? If so, you have already engaged in quality improvement (QI)! QI is focused on how change is implemented in healthcare and in other settings (such as aerospace) with the goal of standardizing processes and reducing variation. QI is achieved through setting specific, measurable goals and making small, iterative changes; all while implementing and evaluating change. If you want to learn how to implement a change, sustain that change over time, and measure success, this course is for you. The project provides an opportunity to engage in QI in your life to implement and evaluate progress on a goal that is important to you (e.g., drinking more water, practicing better time management skills).

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BIOL 1040. Ultrastructure/Bioimaging.

This course examines microscopy and image analysis in the life sciences. Theoretical and practical aspects of microscopy will be discussed. Students will obtain hands-on experience with electron microscopy, light microscopy, fluorescence microscopy, and confocal microscopy. Students will learn to display images in 3D. Advanced undergraduates. Instructor permission required.

Spr BIOL1040 S01 25397 M 2:00-5:00(13) (G. Williams)
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BIOL 1050. Biology of the Eukaryotic Cell.

Examines organelles and macromolecular complexes of eukaryotic cells with respect to structural and functional roles in major cellular activities. Emphasizes experimental basis for knowledge in modern cell biology using original literature, and discusses validity of current concepts. For advanced undergraduates and beginning graduate students. Complementary to BIOL 1270 and 1540. Prerequisites: BIOL 0280 or 0470 or 0500, or instructor permission. Graduate students register for BIOL 2050.

Fall BIOL1050 S01 16652 TTh 1:00-2:20(06) (S. Gerbi)
Fall BIOL1050 C01 16653 Th 12:00-12:50 (S. Gerbi)
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BIOL 1070. Biotechnology and Global Health.

This course examines contemporary biotechnologies used to combat the predominant, worldwide problems in human health. Global health will be addressed from the scientific and engineering perspectives while integrating public health policy, health systems and economics, medical and research ethics, and technology regulation and management. This course is intended for graduate and advanced undergraduate students in biology, engineering, or related fields who have an interest in global health initiatives. Expected background: BIOL 0200 and BIOL 0800, or equivalents. Preference will be granted to graduate students in the Biotechnology and Biomedical Engineering programs. Only for related course credit in Biology. Enrollment limited to 20. Instructor permission required.

Fall BIOL1070 S01 16952 TTh 9:00-10:20(05) (J. Schell)
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BIOL 1090. Polymer Science for Biomaterials.

Basic principles of polymer science and its application in medicine. Topics include basic polymerization chemistry, kinetics of polymerization and depolymerization with emphasis on bioerodible polymers, characterization of polymers by physical methods, bulk and surface properties, behavior of polymers in solutions, crystallization, gelation, and liquid crystals. Hands-on experience with polymer characterization. Expected: CHEM 0350. Enrollment limited to 25.

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BIOL 1100. Cell Physiology and Biophysics.

Current topics in cell physiology, with an emphasis on membrane-mediated interactions between cells and their environment. Topics may include: ion channel structure, function and regulation; intracellular regulatory molecules; mechanisms of sensory transduction; membrane receptors and second messenger systems; vesicle secretion; and cytoskeletal regulation of cell function. Lectures, discussion, and student presentations of the current literature. Expected: BIOL 0800 or NEUR 0010. Instructor permission required. Enrollment limited to 30, and admission is based on seniority -- graduate students, seniors, then juniors. (Not for first and second-year undergraduates.) Students must request an override code through C@B.

Spr BIOL1100 S01 25183 M 3:00-5:30(13) (D. Horrigan)
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BIOL 1110. Topics in Signal Transduction.

Signal transduction is one of the most rapidly developing fields in biomedical sciences. Defects in signaling pathways can be responsible for diseases such as cancer, diabetes, cardiovascular disorders and psychoses. This course offers students an overview of the molecular pathways that allow cells to receive and process signals from their external environment, with an emphasis on the emerging state-of-the-art techniques used in their study. Expected background: BIOL 0200, 0280, 0470, or 0500. Enrollment limited to 20 juniors and seniors.

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BIOL 1120. Biomaterials.

A biomaterial is defined as a material suitable for use in medical implants that come in direct contact with patients' tissues. These include polymers, metals, and ceramics, and materials obtained from biological sources or through recombinant biotechnology. Goal: to provide comprehensive coverage of biomaterial science and technology. Emphasizes the transition from replacement to repair strategies. For advanced undergraduates and graduate students. Prerequisite: BIOL 0800 or instructor permission.

Spr BIOL1120 S01 26848 Th 4:00-6:30(17) (Y. Dingle)
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BIOL 1140. Tissue Engineering.

Tissue engineering is an interdisciplinary field that incorporates progress in cellular and molecular biology, materials science, and engineering, to advance the goal of replacing or regenerating compromised tissue function. Using an integrative approach, we will examine tissue design and development, manipulation of the tissue microenvironment, and current strategies for functional reconstruction of injured tissues. Expected: CHEM 0330, plus BIOL 0500 or 0800. Enrollment limited to 20. Instructor permission required.

Fall BIOL1140 S01 16956 W 3:00-5:30(10) (D. Hoffman-Kim)
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BIOL 1150. Stem Cell Engineering.

Stem cell engineering focuses on using adult, embryonic, and induced pluripotent stem cells to repair damaged or diseased tissues. This course will examine the role of stem cells in development, tissue homeostasis, and wound healing, as well as how they can be used for tissue engineering and cell-based regenerative therapies. We will also discuss the ethical, legal, and regulatory issues that accompany current and emerging stem cell engineering endeavors. The course will use an inverted lecture and classroom discussion format to effectively deliver relevant information. Emphasis is placed on oral and written communication skills applied to assignments, tests, and individual projects. As an additional part of this course, students will receive hands-on training in how to culture cells and assess samples for stemness characteristics in a group laboratory setting.

Spr BIOL1150 S01 25161 Th 3:00-5:30(17) (E. Darling)
Spr BIOL1150 L01 25162 F 9:00-12:00 (E. Darling)
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BIOL 1155. Hormones and Behavior.

This class will explore the hormonal basis of animal behavior. We will assess this relationship at the molecular, cellular, physiological, and evolutionary levels, focusing on a wide range of species beyond humans. Our goal is to understand the diverse mechanisms by which hormones act throughout the animal body to mediate what individuals do in their natural environment. We will explore how selective forces shape these mechanisms to not only arrive at common behavior traits, but also unique and unusual traits that allow species to thrive in harsh or extreme environments.

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BIOL 1160. Principles of Exercise Physiology.

Application of the basic principles of physiology to the study of the response mechanisms of the human body during exercise. Topics include muscle and neural control, energy metabolism, cardiovascular and respiratory effects, endocrinology, principles of training, and special topics (e.g., diving, high altitude, and microgravity). Student presentations based on scientific articles are included. Expected: BIOL 0800 or written permission of the instructor.

Spr BIOL1160 S01 25438 MWF 12:00-12:50(01) (A. Sobieraj)
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BIOL 1180. Comparative Animal Physiology.

Comparative approach to the function and regulation of animal systems with an emphasis on vertebrates. Topics include circulation, gas exchange, neuromuscular function, excretion, acid-base and ion regulation, and temperature regulation. Considers the unity and diversity of physiological processes in animals differing both in phylogeny and environmental adaptation. Original papers are discussed. Expected: BIOL 0800 or equivalent.

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BIOL 1190. Synaptic Transmission and Plasticity.

Synapses are the means by which the nervous system communicates. In this seminar-style course, we will explore the molecular and physiological underpinnings of synaptic transmission. We will then examine ways in which synapses can modulate their strength during development, learning, after addictive drugs, and other adaptive processes. Readings are ONLY from primary literature. Course recommended for juniors and seniors. Required: NEUR 1020. Enrollment limited to 20. Instructor's permission required, attendance at class on the first day is mandatory.

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BIOL 1200. Protein Biophysics and Structure.

Structural Biology is the science to determine 3-dimensional structures of biomacromolecules (i.e. proteins, RNA, and DNA). These structures enable biologists to understand and explore their function. Since proteins, RNA, and DNA are the primary molecules of life, structural biology enables us to understand and influence these molecular machineries which form the basis of all biological processes. Throughout the class, the students will see examples of biologically important proteins and protein complexes that will allow them to correlate structure and biological function. Prerequisite: BIOL 0280.

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BIOL 1210. Synthetic Biological Systems.

A multidisciplinary course that combines science and engineering providing a solid foundation in a cutting edge field of biological engineering. Synthetic biology is a mixture of biology, chemistry, engineering, genetic engineering and biophysics. It builds on recent work in systems biology which involves the modeling of biological systems, but goes further in that it involves the construction and standardization of biological parts, that fit together to form more complex systems. Expected: at least four courses beyond BIOL 0200, CHEM 0330, PHYS 0300, ENGN 0300, MATH 0900, or CSCI 0040.

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BIOL 1220. Synthetic Biological Systems in Theory and Practice.

A multidisciplinary laboratory, lecture, and discussion based course that combines several areas of science and engineering providing a foundation in the cutting edge field of synthetic biological engineering. The field of synthetic biology is centered around trying to make biology easier to engineer. It builds on recent work in systems biology which involves the modeling of biological systems, but goes further in that it involves the construction and standardization of biological parts that fit together making complex systems. This course will combine classes, guest lectures and discussion lab visits to give students the best possible tools for understanding and applying research in synthetic biology. Expected: at least two courses in any of the key disciplines (biology, chemistry, physics, math, engineering, computer science) beyond the introductory level, and permission of the instructor.

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BIOL 1222A. Current Topics in Functional Genomics.

A technological revolution in genomics has exponentially increased our ability to gather biological data. A host of new methods and types of analysis has arisen to accommodate this dramatic shift in data collection. The broad scope of inquiry has ushered in an era of “system-wide” approaches and brute-force strategies where rare signals can be detected and studied. In this seminar we will cover papers that embody this new approach. Students typically have taken an advanced undergraduate-level course in biology.

Spr BIOL1222A S01 25399 W 3:00-5:30(10) (W. Fairbrother)
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BIOL 1250. Host-microbiome Interactions in Health and Disease.

Will focus on current understanding of how various microbiomes communicate and interact with the host and the factors that influence these interactions. We will discuss how the new technologies such as metagenomics and metabolomics have enhanced our understanding of host-microbiome interactions in health and disease. Students will have the opportunity to participate in discussions on how to apply recent discoveries to disease processes, health restoration and maintenance. The course will help students develop skills in critical thinking and in reading and evaluating original scientific literature. Expected: students with a background in basic microbiology (BIOL 0530 or its equivalent). 20 enrollment.

Spr BIOL1250 S01 25143 Th 2:00-4:30(11) (P. Belenky)
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BIOL 1260. Physiological Pharmacology.

Covers the physiology of human disease (e.g., Heart failure and arrhythmia; cancer signaling pathways with a focus on breast cancer; neurological disorders such as schizophrenia and Parkinson's disease) and discusses the pharmacology of the drugs used to treat disease. A group of the most commonly prescribed drugs is discussed in terms of their fundamental modes of action and clinical importance. Expected: BIOL 0800.

Fall BIOL1260 S01 16650 TTh 10:30-11:50(13) (J. Marshall)
Fall BIOL1260 S02 19309 Arranged (J. Marshall)
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BIOL 1270. Advanced Biochemistry.

An advanced course in biochemistry, biochemical methods, and reading of the primary literature, featuring systematic coverage of the biochemistry of the central dogma, including DNA (replication, repair, recombination), RNA (regulation and mechanism of transcription, processing, turnover), and proteins structure, synthesis, modification, degradation, mechanisms of action, function). Expected: BIOL 0280, CHEM 0350, 0360. Graduate students register for BIOL 2270.

Fall BIOL1270 S01 16656 TTh 2:30-3:50(12) (G. Lisi)
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BIOL 1290. Cancer Biology.

Provides a conceptual understanding of molecular events underlying development of human cancer. Focused on genetic changes leading to malignant transformation of cells. Covers cell cycle control, DNA damage, mutagenesis, cancer predisposition syndromes, oncogenic viruses, tumor immunology, metastasis, cancer chemotherapy and drug resistance. Lecture plus discussion of primary literature. Prerequisites: BIOL 0280 OR BIOL 0470 OR BIOL 0500.

Fall BIOL1290 S01 16633 MW 3:00-4:20(10) (A. Zhitkovich)
Fall BIOL1290 S02 18903 W 3:00-6:00(10) (P. Dubielecka-Szczerba)
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BIOL 1300. Biomolecular Interactions: Health, Disease and Drug Design.

Interactions between the molecules of life-proteins, RNA, DNA, membrane components-underlie all functions necessary for life. This course focuses on how nature controls these interactions, how these interactions can go awry in disease, and how we can learn the rules of these interactions to design drugs to treat disease. Students will review the physical basis of molecular interactions, learn classic and state-of-the-art high-resolution and high-throughput tools used to measure interaction, and survey the experimental and computational strategies to harness these interactions using a case study in rational drug design. Prerequisite: Introductory Biochemistry (BIOL 0280). Enrollment limited to 20; instructor permission.

Fall BIOL1300 S01 16777 M 3:00-5:30(03) (N. Fawzi)
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BIOL 1310. Developmental Biology.

Covers the molecular and cellular events of development from fertilized egg to adult. Genetic basis of body form, cell fate specification and differentiation, processes controlling morphogenesis, growth, stem cells and regeneration will be examined. Differential gene regulation, intercellular signaling and their evolutionary conservation will be central to discussion of mechanisms governing developmental processes. Additional topics: developmental plasticity, impact of epigenetic and environmental factors, and basis of disease gleaned from developmental biology research. Live embryos will complement and reinforce concepts covered in class. Enrollment limited to 36. Expected: BIOL 0200 (or equivalent), and one course in genetics, cell biology or embryology.

Spr BIOL1310 S01 26862 TTh 9:00-10:20(05) (K. Wharton)
Spr BIOL1310 L01 26863 W 1:00-5:00 (K. Wharton)
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BIOL 1330. Biology of Reproduction.

This course is an advanced, seminar-based course. Primary literature is emphasized to complement the format of extensive student seminar presentations. It is essential that students have a strong background in biology in order to gain the most from this course. The emphasis of the course is student seminar presentation and extensive discussion on the material. This is often the first opportunity for students to present/discuss science in a seminar format. Expected background: a course in Cell Biology (e.g. BIOL 0500 or 1050), and two additional Biology courses above the introductory (BIOL 0200) level. Enrollment limited to 20.

Spr BIOL1330 S01 25401 M 3:00-5:30(13) (G. Wessel)
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BIOL 1410. Evolutionary Genetics.

This course will focus on selected topics in molecular population genetics, molecular evolution, and comparative genomics. Classic and current primary literature at the interface of evolution and genetics will be discussed in a seminar format. The laboratory involves wet-lab exercises (allozymes, PCR- RFLP, sequencing), plus computer labs using DNA analysis packages. Students will prepare a final grant proposal on specific research interests. Expected: BIOL 0470 or 0480. Students MUST register for the lecture section and the lab. Enrollment limited to 20.

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BIOL 1420. Experimental Design in Ecology.

An overview and discussion of the basic principles used to design lab and field experiments in ecology and environmental science. Topics include: replication and statistical power, appropriate use of factorial designs, nonparametric methods, post hoc texts, natural versus manipulative experiments, experimental artifacts and impact study design. Discussions based on primary literature and a new text. Expected: BIOL 0420.

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BIOL 1425. Phylogenetic Biology.

This course is the study of the evolutionary relationships between organisms, and the use of evolutionary relationships to understand other aspects of organism biology. This course will provide a detailed picture of the statistical, mathematical, and computational tools for building phylogenies and using them to study evolution. Enrollment is by instructor permission. Students will present scientific papers in class and complete a final project consisting of their own phylogenetic analysis. Expected Background: Evolutionary Biology and quantitative methods (such as statistics, computation, or math). Open to juniors, seniors, and graduate students. Enrollment limited to 16.

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BIOL 1430. Foundations of Population Genetics.

Population genetics considers the genetic basis of evolution: temporal changes in the genetic composition of populations in response to processes such as mutation, natural selection and random sampling effects. Starting from first principles, this course will develop a theoretical understanding of these dynamics. We will also explore the application of these tools to genomic-scale data in order to quantify the influence of various evolutionary processes at work in natural populations. Assessments will be based on problem sets, two midterm exams and one final exam. Prerequisites: MATH 0100 and one of BIOL 0470 or 0480, or permission.

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BIOL 1435. Computational Methods for Studying Demographic History with Molecular Data.

This course broadly covers the field of population genetics and genomics, and focuses on how inferences about demographic history can be made from genetic variation observed across populations today. The main question we will endeavor to answer in class is “How can we infer demographic history in a population using next-generation sequencing data?” Students will also learn how to apply computational tools/methods to infer demographic history using both simulated and real DNA sequencing data.

Spr BIOL1435 S01 25351 TTh 2:30-3:50(11) 'To Be Arranged'
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BIOL 1440. Marine Biology.

An examination of current topics in the ecology of marine organisms and communities. Current literature and ideas are analyzed in a seminar format (5hr/week). A class research project provides hands-on experience with designing and interpreting experimental field work. Prerequisites: BIOL 0410 and 0420. Instructor's permission required.

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BIOL 1445. Medicine in Austere Settings.

This course examines the practice of medicine in austere settings and will include content related to tropical medicine/neglected tropical diseases, humanitarian relief, disaster management and wilderness medicine. Students will explore the history of these fields and the various challenges that medical professionals face in these environments. The course will particularly focus on the multitude of ethical dilemmas faced when providing care in austere settings. Students should expect to think critically about challenging questions. As part of the course, students will also learn various skills useful to the practice of medicine in austere settings, including basic epidemiology, public health coordination, disaster management and practical (hands-on) wilderness medicine techniques. This course is particularly well-suited for students who are interested in public health, international health policy or aim to pursue a career as a medical professional.

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BIOL 1450. Community Ecology.

This course will explore foundational concepts in community ecology, and will draw on examples and case studies from marine and terrestrial ecosystems, including species-rich tropical rain forests and coral reefs, the marine intertidal and benthic environments, and species-poor forests and grasslands of the temperate zone. Overarching themes will emphasize theoretical frameworks to understand the evolutionary origins and maintenance of this biological diversity. This will be accomplished using traditional lectures, weekly student-led discussions, readings of the primary literature, and class activities. Expected background: BIOL 200 or equivalent placement; and BIOL 0420; OR instructor permission.

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BIOL 1455. Planetary Health: Global Environmental Change and Emerging Infectious Disease.

Will a warmer world be a sicker world? What is it about New England that supports the proliferation of Lyme Disease? How are wildlife trade and species invasions contributors to emerging diseases like covid-19? We will explore these and related questions in Planetary Health. Planetary health is a timely field focused on understanding the human health implications of human-caused disruptions to Earth's natural systems. The facet of ‘health’ that we focus on is infectious disease. Students will learn how/when/where/why infectious diseases emerge in association with anthropogenic environmental impacts. Students will learn through engagement with an array of materials and experiences including lectures, primary/secondary literature, expert-based resources, peer and instructor interaction, virtual discussion, interest driven projects, reflections, and David Quammen’s ‘Spillover’. The course is open to juniors/seniors and designed for those with interests in environmental science, conservation biology and human health.

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BIOL 1465. Human Population Genomics.

An introduction to human genomics and the evolutionary forces that shape observed genetic variation across humans today. Topics will include the relationship among humans and other primates, human population genetics and genomics, and examples of the concomitant evolution of both cultural traits and domesticated organisms. Assignments include a class presentation and reviewing papers on a selected topic. Expected background: BIOL 0470 or 0480, and BIOL 0495, PHP 2500, or equivalent. Enrollment limited to 25. Instructor permission required.

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BIOL 1470. Conservation Biology.

Conservation Biology is the scientific study of the phenomena that affect the maintenance, loss, and restoration of biological diversity. Topics covered include: 1) the impacts of global warming, species invasions, and habitat destruction on biodiversity, 2) strategies developed to combat these threats, and 3) a consideration of key economic and ethical tradeoffs. Special attention will be paid to current debate and controversy within this rapidly emerging field of study. Readings will include the primary literature. Prerequisite: BIOL 0420 or instructor permission. Enrollment limited to 30.

Fall BIOL1470 S01 16872 TTh 9:00-10:20(05) (D. Sax)
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BIOL 1475. Biogeography.

Will provide an overview of the field of biogeography--the study of geography of living organisms. Class meetings will be split between lectures and discussions. Each discussion will expose students to foundational papers, which set the context for the field's development, and more recent papers, which show where the field is headed. Each student will conduct a short (but time consuming) original research project on some topic in biogeography. Prerequisites: BIOL 0420 and 0480. Expected: one taxonomy-based course (e.g., BIOL 0410, 0430, or 0460). Enrollment limited to 15 juniors, seniors, and graduate students. Instructor permission required.

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BIOL 1480. Terrestrial Biogeochemistry and the Functioning of Ecosystems.

Three fundamental multidisciplinary questions will be addressed: How do ecosystems work? What limits the growth of life on Earth? How are humans altering the framework in which all life exists? Earth is basically a closed chemical system, and the reactions that support life are fueled by sunlight. But added to this chemistry and physics is the tremendous influence of life. Life created an oxygen atmosphere; the evolution of biological nitrogen fixation exponentially increased how many organisms could exist, and the soils that support human food production developed only by biologically-mediated processes. Throughout Earth's 4.5 billion-year history changes in Earth's basic biogeochemical processes have been fairly slow. Under our inattentive stewardship, we have almost instantaneously altered all of the major element cycles. We will focus heavily on what these changes mean for life on Earth. Instructor permission required.

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BIOL 1485. Climate Change Health and Ecology.

Explore the linkages between climate change and health. Students will come to appreciate the topic through the foundations of the primary disciplines relevant to the field including global health, environmental change, disease ecology, and others. Climate-health linkages will be learned through weekly case studies addressed collectively through student-centered lectures, discussion of the primary literature, groups activities and guest lectures from campus faculty on topics ranging from climate migration to infectious disease range shifts. Expected background: BIOL 0475, or BIOL 1470, or PHP 1070, or PHP 1920, or equivalent experience with instructor's permission. Enrollment limited to 12 juniors and seniors.

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BIOL 1495. 500 Million Years of Land Plants.

Explores the evolution of terrestrial plants and the ecosystems they structure. Introduces the fossil record of plants and basic patterns of plant diversification on land. Highlights major trends in the evolution of plant morphology, anatomy, and ecology. Lectures survey the diversity and community structure of different geological time periods. Weekly discussion sections, field trips, and assignments examine major evolutionary trends, particularly with regard to climatic changes over time. Expected: BIOL 0400, BIOL 0430, (or equivalent placement). Enrollment limited to 15 students; instructor permission; register for section and conference.

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BIOL 1500. Plant Physiological Ecology.

An in-depth look at plant ecological strategy, focusing on the anatomical and physiological adaptations of plants to particular environments. Additional topics include plant-animal interactions, historical biogeography, and community assembly processes. A comparative, phylogenetic approach is emphasized. Lectures present a broad overview of topics, and discussions focus on current outstanding problems. Lab exercises provide hands-on experience in designing experiments, measuring plant performance, and scientific writing. Required laboratory hours to be arranged by the instructor. Expected: BIOL 0430 or BIOL 0440. Enrollment limited to 15.

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BIOL 1515. Conservation in the Genomics Age.

The course will introduce students to the rapidly developing field of molecular ecology, emphasizing its importance for conservation biology. Students will explore key principles in evolutionary ecology based on readings, lectures, and discussions. Participants will also gain practical experience with ecological, genomic, and computational methods in the lab. This course is intended for advanced undergraduate and graduate students. Suggested prerequisites include Principles of Ecology (0420); Evolutionary Biology (0480) or Genetics (0470); the Lab Techniques Workshop for Biology Students provided by MDL; or similar with permission. Students will obtain permission from the professor to enroll.

Fall BIOL1515 S01 16735 MWF 9:00-9:50(09) (T. Kartzinel)
Fall BIOL1515 L01 16736 W 1:00-4:00 (T. Kartzinel)
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BIOL 1520. Innate Immunity.

Innate immunity is the initial response to microbes that prevents infection of the host. It acts within minutes to hours, allowing the development of the adaptive response in vertebrates. It is the sole mechanism of defense in invertebrates such as insects. The components and mechanisms dictating this response are explored. Prerequisite: BIOL 0530. Enrollment limited to 30. Graduate students must obtain instructor permission.

Fall BIOL1520 S01 16631 MW 8:30-9:50(09) (L. Brossay)
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BIOL 1530. Emergency Medical Systems: An Anatomy of Critical Performance.

Students taking BIOL 1530 will explore Emergency Medicine and the many subjects that make up the practice of providing care to patients at the most critical and dire moments of their lives. We will read, discuss, and write about Trauma, Medical Malpractice, Death and Dying, Refugee and Disaster Medicine, Interpersonal Violence, and Crisis focused Medical Ethics, among others, all viewed through the lens of Emergency Medicine. You will shadow in the Emergency Department of a Trauma Center and debrief with your TA’s in weekly meetings, where the class topics will be discussed in open forums. This class, formerly PHP1520, has created a transformative experience for more than 30 years and will continue to foster new perspectives and ideas for every student who participates.

Spr BIOL1530 S01 26236 W 3:00-5:30(10) (B. Becker)
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BIOL 1540. Molecular Genetics.

Even in this era when whole genome DNA sequencing has become routine, there are still thousands of eukaryotic genes with unknown functions. Genetic screens for mutations that alter pathways of interest remain the premier approach to understanding gene function in the context of the organism. In Molecular Genetics students will learn the key concepts involved in designing and interpreting genetic screens using the powerful tools available in model animal, plant, and fungal organisms. Students will also learn how to understand and analyze results presented in the primary scientific literature. Furthermore, students will gain an appreciation of how the field of genetics has changed through discoveries and technological advances made over the past 50 years. Graduate students should register for BIOL 2540.

Spr BIOL1540 S01 25402 TTh 2:30-3:50(11) (E. Larschan)
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BIOL 1545. Human Genetics and Genomics.

This course will exemplify the power of genetically informed approaches to understanding human biology. It is intended for advanced undergraduate students and graduate students; prerequisites include BIOL0470 or equivalent. The course is based in lectures, reading material (textbook and primary literature), and in-class discussions. Course topics include: medical genetics and genomics; methods to study human genotypes and related phenotypes; industry-related topics; and ethical and societal implications of genome science. It will benefit students with career interests in basic science, medicine, biotechnology, or science policy. Enrollment is limited to 20 students; selection will be based on seniority, prerequisites, and registration order.

Spr BIOL1545 S01 25404 W 3:00-5:30(05) (E. Morrow)
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BIOL 1550. Parasitism: Biology and Disease.

Protozoan parasites influence the health of human populations in less developed countries, leading to severe human suffering and loss of economic development. Understanding the fundamental biology of these pathogens and how they are transmitted is essential for developing treatments and mitigation strategies to improve human health. Focuses on biochemical, genetic, cellular, and immunological aspects of parasite and host responses. Expected: BIOL 0470 or BIOL 0530.

This course was formerly "Biology of Emerging Microbial Diseases"

Spr BIOL1550 S01 25144 MWF 11:00-11:50(04) (C. de Graffenried)
Spr BIOL1550 C01 25145 M 6:00-6:50 (C. de Graffenried)
Spr BIOL1550 C02 25146 W 6:00-6:50 (C. de Graffenried)
Spr BIOL1550 C03 25147 T 12:00-12:50 (C. de Graffenried)
Spr BIOL1550 C04 25148 Th 12:00-12:50 (C. de Graffenried)
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BIOL 1555. Methods in Informatics and Data Science for Health.

The goal of this course is for students to develop a solution that uses data science and informatics approaches to address a biomedical or health challenge. This course will teach informatics and data science skills needed for public health and biomedicine research. Emphasis will be given to algorithms used within the context of biomedical research and health care, including those used in biomolecular sequence analysis, electronic health records, clinical decision support, and public health surveillance. This course has been developed as a Course-based Undergraduate Research Experience (CURE), where students will gain experience with the scientific method, its application, and presentation.

Spr BIOL1555 S01 25179 M 3:00-5:30(13) (E. Chen)
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BIOL 1560. Virology.

Emphasizes the understanding of molecular mechanisms of viral pathogenesis. Begins with a general introduction to the field of virology and then focuses on the molecular biology of specific viruses that are associated with human disease. Lectures based on current literature. Prerequisite: BIOL 0280, 0470, or 0530, or instructor permission.

Fall BIOL1560 S01 16559 MWF 9:00-9:50(09) (S. Lee)
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BIOL 1565. Survey of Biomedical Informatics.

Survey course provides overview of field of biomedical informatics. Topics include computer science, healthcare, biology, social science. This course is designed to be complementary to BIOL 1555. Emphasis given to understanding the organization of biomedical information, effective management of information using computer technology, impact of such technology on biomedical research, education, patient care. Major aim explores the process of developing and applying computational and information science techniques for assessing current information practices, determining information needs of health care providers and patients, developing interventions or supporting clinical practice using informatics, and evaluating the impact of informatics solutions from a biomedical perspective.

Fall BIOL1565 S01 16641 M 3:00-5:30(03) (N. Sarkar)
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BIOL 1575. Evaluation of Health Information Systems.

This course covers the field of evaluation of health information systems (HIS) in a range of roles and environments, in the US and worldwide. It includes topics in health information system (HIS) design and deployment, healthcare workflow, quantitative and qualitative evaluation methods and socio-technical environment for HIS. Emphasis is given to understanding the range of evaluation questions that can be asked, identifying the key stakeholders, understanding available evaluation techniques, and designing rigorous but achievable studies. Examples will include Open Source systems, medical Apps, and economic evaluation, the role of evaluation frameworks and theories, and notable HIS successes and failures. Recommended: past or concurrent enrollment BIOL 1565 or a public health course covering clinical research.

Fall BIOL1575 S01 16642 TTh 1:00-2:20(06) (H. Fraser)
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BIOL 1595. Artificial Intelligence in Biomedicine.

This course will teach the fundamental theory and methods of artificial intelligence (AI) alongside their application to the biomedical domain. It will give a representative overview of traditional methods as well as modern developments in the areas of (deep) machine learning, natural language processing and information retrieval. The course is designed to be accessible to non-computer science audiences and will not require extensive prior programming experience. The course will be accompanied by practical assignments applying the discussed techniques in a biomedical context. Understanding of formal theoretical knowledge will be assessed in a final exam. The course is designed for students concentrating in domains such as Computational Biology and Applied Mathematics-Biology; or Neuroscience concentrators who have completed a course in introductory statistics (e.g., BIOL 0495).

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BIOL 1600. Development of Vaccines to Infectious Diseases.

Provides background steps involved in vaccine development, from conceptualization to production to deployment. Considers infectious diseases and associated vaccines in context of community health. Appropriate for students wanting to gain an understanding of vaccine science. Provides a foundation for advanced courses in immunology and infectious disease, biomedical research, or medical/graduate studies. Activities include a weekly section meeting for discussion of relevant primary literature, and a final project of the student’s choice in the form of an in–class presentation, a research paper or an approved alternative format. Expected: BIOL 0200 or equivalent placement; BIOL 0530, and at least one additional biology course.

Spr BIOL1600 S01 25149 MW 3:00-4:20(10) (S. Vaishnava)
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BIOL 1610. Experimental Neurobiology.

Intensive laboratory experience in neuroscience appropriate for students with basic background in Neurobiology. Learn and employ the classical neurophysiological techniques of extracellular recording, intracellular recording and receptive field mapping using a variety of animal species. Experiments will include recording of sensory signals in the cockroach leg; frog sciatic nerve and sciatic nerve/muscle preparation; intracellular recording of neurons in Aplysia; receptive field mapping in frog skin; and visual field mapping in the frog tectum. Instruction on and practice of effective science writing is another component to this course. Labs are supplemented by informal lectures. Enrollment limited to 18. Please request override through C@B. Overrides will not be given until after the first course meeting.

Spr BIOL1610 S01 26845 W 1:00-6:00(06) (J. Stein)
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BIOL 1650. Structure of the Nervous System.

Combined lecture and laboratory course on the anatomy of the central nervous system. Lectures survey the circuitry of the major neural systems for sensation, movement, cognition, and emotion. Laboratory exercises (Mon. 10:30-12:30) include brain dissections, microscopy of neural tissue, and discussion of clinical cases. Prerequisites: NEUR 0010, NEUR 1020, and NEUR 1030. Please request an override through C@B. Please keep in mind that decisions on overrides may not be made until the first meeting of the course.

Fall BIOL1650 S01 18583 TTh 2:30-3:50(12) (D. Berson)
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BIOL 1800. Animal Locomotion.

How and why do animals run, jump, swim and fly? Physiology, anatomy, ecology, and evolutionary history all influence, and are influenced by, the way animals move around. We will integrate analyses from many levels of biological organization - from molecular motors, through bone-muscle systems, to biogeography - with methods and approaches from mechanics, fluid dynamics, and robotics. Expected: BIOL 0800 and PHYS 0030. Instructor permission required.

Fall BIOL1800 S01 16739 TTh 2:30-3:50(12) (S. Swartz)
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BIOL 1810. 21st Century Applications in Cell and Molecular Biology.

Twenty-first century applications in cell and molecular biology focuses on the structure and function of macromolecules and cells and how they are altered in disease and therapy. This course will explore physical principles underlying cell function, along with biophysical approaches for solving problems of cell and molecular biology. Cutting-edge molecular and cellular-based therapeutics will be discussed throughout this course; this includes viral gene delivery constructs, novel platforms for tissue engineering, CRISPR genome editing, and immune checkpoint therapy. This course is particularly suitable for undergraduate students interested in basic medical research, graduate school, or research-based careers in biotechnology or pharmaceutical industry.

Spr BIOL1810 S01 25406 MWF 10:00-10:50(03) (M. Dawson)
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BIOL 1820. Environmental Health and Disease.

Humans live, work, and play in complex chemical environments. BIOL1820 examines how environmental exposures impact human health and contribute to disease. The course covers basic concepts in toxicology, epidemiology, and safety assessment, and is divided into 4 sections: radiation, lead, perfluorinated chemicals, and endocrine disruptors. For each section, students will examine the molecular mechanisms that mediate toxicity, learn how toxicant exposure impacts physiology, evaluate exposure risk, and discuss issues of environmental justice. Prerequisites: introductory level biology and chemistry. BIOL 1820 is designed for junior and senior undergraduates, and is open to others with permission.

Spr BIOL1820 S01 25169 TTh 10:30-11:50(09) (J. Plavicki)
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BIOL 1850. Environmental and Genetic Toxicology.

Human disease is produced by complex interactions between inherited genetic predisposition and environmental exposures. These interactions will be explored at the molecular, cellular, and systemic levels. Prototype diseases will include hereditary disorders of hemoglobin, hypercholesterolemia, birth defects, and cancer. Expected: Cell Biology.

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BIOL 1865. Toxicology.

Toxicology is the science that describes the adverse biological effects of exogenous chemical and physical stressors, including environmental, industrial, and agricultural chemicals and pharmaceuticals. This course will introduce the principal biological processes that determine an organism’s response to a toxicant, including absorption, distribution through a biological system, metabolism, elimination, and effects at the site(s) of action. We will discuss modern challenges in toxicology, such as assessing toxicity of mixtures and testing some of the thousands of untested chemicals in commerce. The material will be presented in lecture and student-led discussions, with readings from the toxicology literature.

Spr BIOL1865 S01 25170 TTh 1:00-2:20(08) (D. Spade)
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BIOL 1870. Techniques and Clinical Applications in Pathobiology.

A methodology course featuring laboratory and lecture instruction in established and leading-edge technologies. Examples: flow cytometry (multi-parameter analysis, cell sorting); molecular biology (PCR, real time PCR, in situ hybridization, microarrays, DNA sequencing, bioinformatics); digital imaging (image acquisition, processing and analysis); confocal microscopy ; histology and immunohistochemistry(confocal, immuno-histochemistry).

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BIOL 1880. Comparative Biology of the Vertebrates.

The biology, structure, and evolutionary history of the vertebrates considered phylogenetically, emphasizing evolution of the major body systems. Stresses an evolutionary approach to the correlation of structure and function with environment and mode of life. Labs include dissection of several different vertebrates and comparative osteological material. Emphasis of course is on critical thinking rather than memorization of material. Recommended: BIOL 0320 or 0800. First year students must obtain instructor permission to register. Enrollment limited to 32. Students MUST register for the lecture section and the lab.

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BIOL 1885. Human Anatomy and Biomechanics.

Human Anatomy and Biomechanics examines the structure of the human body from the perspectives of biomechanics, evolution, and development, focusing on the musculoskeletal system, sensory systems, and the complex, multifunctional organ of the human skin. The course considers biological form and function at multiple levels of organization and in diverse contexts, and offers students opportunities to synthesize, question, and write about ideas presented in class. The course includes a weekly, 2-hr laboratory section in which students will use kinesthetic learning to develop deep knowledge of anatomico-medical terminology, study osteology and develop powers of observation working with human and non-human animal bones, dissect a preserved mink specimen to learn the soft-tissue anatomy of a mammal and visit the human anatomy dissection lab at the Alpert Medical School.

Spr BIOL1885 S01 25352 TTh 1:00-2:20(08) (S. Swartz)
Spr BIOL1885 L01 25353 M 3:00-5:00 (S. Swartz)
Spr BIOL1885 L02 25354 T 3:00-5:00 (S. Swartz)
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BIOL 1890. Human Histology.

This course will provide an in-depth treatment of the “stuff we are made of” and the wonderful logic of its organization. This course focuses fi rst on the biology of the four basic tissues (epithelium, connective tissue, muscle and nerve) and second, how they contribute to the functional anatomy of all organs and systems. For Pfizer students only.

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BIOL 1920C. Social Contexts of Disease.

What shapes our understandings of disease, and what makes a disease real? How might we explain the demise of formerly prevalent diseases and the arrival of others? How do politics, technologies, and institutions affect conceptions of disease and structure their treatment? Will examine the impact of social context on patients' experiences of disease, including clinical, scientific, and public health approaches. Will consider disease in relation to social relationships, power of the state to regulate disease, and cultural care of the body. Enrollment limited to 20 students; instructor permission required; serves as Capstone in Health and Human Biology. Not for concentration credit.

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BIOL 1941A. Plants in a Changing Planet.

Plants are the foundation of Earth's ecosystems and essential to human survival and civilization. This seminar will examine the physiological, ecological, and evolutionary responses of plants to rapid environmental change, and the consequences for agriculture and the structure and function of natural systems. Expected background: at least one of the following courses - BIOL 0420, 0430, 0440, 0480, or ENVS 0490.

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BIOL 1945. Outbreaks and Infections: Harnessing History and Preventing Future Epidemics.

This senior capstone course will investigate epidemic diseases that have plagued mankind in the past and explore the factors that contribute to the continued spread of diseases today. This course will emphasize the biology of both infectious and noninfectious diseases by exploring mechanisms of spread, pathogenesis, and prevention. Attention will also be given to environmental, economic, geographical, and social contributors to disease. Interactive class discussions, student presentations, analysis of primary literature, and multimedia homework will develop student skills culminating in an independent research project that dissects the life history of a disease and provides a forward-thinking strategy to combat it.

Fall BIOL1945 S01 18928 TTh 1:00-2:20(06) (A. Dugan)
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BIOL 1950. Directed Research/Independent Study.

Directed research/independent study in biological sciences: basic science, social studies of biomedical science, and clinically-oriented projects, mentored by individual faculty members in the Division of Biology and Medicine. Sites include campus and hospital based facilities. Projects can serve as the basis for Honors theses, or to fulfill research requirements in a Bio-Med concentration program. Students planning to use 1950/1960 to fulfill a concentration requirement must receive approval from the concentration advisor. No more than two (2) semesters of BIOL 1950/1960 may be used toward a concentration program in the biological sciences. Faculty from outside the Division may supervise projects for bio-med program concentrators, but should do so using their Department's own Independent Study course number.

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BIOL 1960. Directed Research/Independent Study.

Directed research/independent study in biological sciences: basic science, social studies of biomedical science, and clinically-oriented projects, mentored by individual faculty members in the Division of Biology and Medicine. Sites include campus and hospital based facilities. Projects can serve as the basis for Honors theses, or to fulfill research requirements in a Bio-Med concentration program. Students planning to use 1950/1960 to fulfill a concentration requirement must receive approval from the concentration advisor. No more than two (2) semesters of BIOL 1950/1960 may be used toward a concentration program in the biological sciences. Faculty from outside the Division may supervise projects for bio-med program concentrators, but should do so using their Department's own Independent Study course number.

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BIOL 1970A. Stem Cell Biology.

Senior seminar course will provide an interactive forum by which up to twenty seniors (and qualified juniors with permission) will explore the biology of stem cells from their humble beginnings in the embryo to their potential use in regenerative medicine. The potency and regulation of embryonic and adult stem cell populations derived from diverse organisms will be contrasted with laboratory-derived human stem-like cells for biomedical applications. Critical reading of classical and modern literature in the field of stem cell biology will form the basis of student-led presentations, papers and ethical forums. Expected: biochemistry, genetics and/or cell biology. Instructor permission; 20 students.

Fall BIOL1970A S01 16779 M 2:00-4:30(10) (R. Freiman)
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BIOL 1980. HIV/AIDS in Diverse Settings: Focus on Israel.

Participants in this winter session course will explore HIV/AIDS within the context of Israel’s diverse society, unique demographics and universal healthcare. While in Israel, students will visit clinics, hospitals, and universities, engage with health care providers, experts in the field and populations with HIV. By the end of the course students will gain research skills and an understanding of this pandemic, its management and challenges in Israel, and how this important disease is modulated through risk factors, healthcare systems, medical innovations, and socio-economic factors. There is no need for prior experience in any associated discipline or any knowledge of Hebrew.
Course schedule: January 2-6, 2023 - daily seminars at Brown (including student and guest lectures); January 7-16 - tour in Israel; January 17 - rest day; January 18-24 - daily seminars at Brown.

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BIOL XLIST. Courses of Interest to Biology Concentrators.

Fall 2022
The following courses may be taken for concentration credit. Please see the sponsoring department for the time and location of each course.

Africana Studies
AFRI 1920 Health Inequality in Historical Perspective
Cognitive, Linguistics, Psychological Sciences
CLPS 1195 Life Under Water in the Anthropocene
Neuroscience
NEUR 1530 Communication In the Brain: What We Know and How We Know It
Spring 2023
The following courses may be taken for concentration credit. Please see the sponsoring department for the time and location of each course.

Environmental Studies
ENVS 0490 Environmental Science in a Changing World

Senior Associate Dean for the Program in Biology and Chair

Edward Hawrot

Senior Associate Dean for Biology and Curricular Affairs

Katherine F. Smith

Professor

Roy K. Aaron
Professor of Orthopaedics

Abbas E. Abbas
Professor of Surgery

J. Dawn Abbott
Professor of Medicine

Ana M. Abrantes
Professor of Psychiatry and Human Behavior

Charles A. Adams
Professor of Surgery

Eli Y. Adashi
Professor of Medical Science

Kailash C. Agarwal
Professor Emeritus of Medical Science (Research)

Carlos Aizenman
Royce Family Professor of Teaching Excellence in Neuroscience

Edward Akelman
Vincent Zecchino, MD Professor of Orthopaedic Surgery

Jorge E. Albina
Professor of Surgery

Joseph F. Amaral
Professor Emeritus of Surgery

Diane J. Angelini
Professor Emerita of Obstetrics and Gynecology, Clinician Educator

Douglas Anthony
Professor of Pathology and Laboratory Medicine; Professor of Neurology

James A. Arrighi
Professor of Medicine; Professor of Diagnostic Imaging; Professor of Medical Science

Michael K. Atalay
Professor of Diagnostic Imaging; Professor of Medicine

Walter J. Atwood
Professor of Medical Science; Professor of Neurology

Gilad Barnea
Sidney A. Fox and Dorothea Doctors Fox Professor of Ophthalmology and Visual Science, Professor of Neuroscience

Christine E. Barron
Professor of Pediatrics

Jay M. Baruch
Professor of Emergency Medicine

Cynthia Battle
Professor of Psychiatry and Human Behavior

Samuel I. Beale
Professor Emeritus of Biology

Curt G. Beckwith
Professor of Medicine

Michael D. Beland
Professor of Diagnostic Imaging

Judith L. Bender
Professor of Biology

Richard J. Bennett
Professor of Biology

Lionel Bercovitch
Professor of Dermatology

David M. Berson
Sidney A. Fox and Dorothea Doctors Fox Professor of Ophthalmology and Visual Science

Mark D. Bertness
Professor Emeritus of Ecology and Evolutionary Biology

Richard W. Besdine
Professor of Medicine; Professor of Health Services, Policy and Practice

Piero Biancani
Professor Emeritus of Medicine (Research)

Joseph M. Bliss
Professor of Pediatrics

Andrew S. Blum
Professor of Neurology

Beth C. Bock
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Kim Boekelheide
Professor Emeritus of Pathology and Laboratory Medicine, Professor of Pathology and Laboratory Medicine (Research)

Jeffrey Borkan
Professor of Family Medicine; Professor of Medical Science

Christopher T. Born
Intrepid Heroes Professor of Orthopedic Surgery

Ghada Bourjeily
Professor of Medicine

Wayne D. Bowen
Upjohn Professor of Pharmacology, Professor Emeritus of Neuroscience

Jerrold L. Boxerman
Professor of Diagnostic Imaging

Elizabeth Brainerd
Robert P. Brown Professor of Biology, Professor of Medical Science

Lundy Braun
Professor of Medical Science; Professor of Africana Studies

Andrew S. Brem
Professor Emeritus of Pediatrics

Laurent Brossay
Charles A. and Helen B. Stuart Professor of Medical Science, Professor of Molecular Microbiology and Immunology

Larry K. Brown
Professor of Psychiatry and Human Behavior

Linda L. Brown
Professor of Emergency Medicine; Professor of Pediatrics

John E. Buster
Professor Emeritus of Obstetrics and Gynecology

Blake Cady
Professor Emeritus of Surgery

Anthony Angelo Caldamone
Professor of Surgery; Professor of Pediatrics

Angela Caliendo
Warren Alpert Foundation Professor of Medicine

Andrew G. Campbell
Professor of Medical Science

Michael Carey
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Linda L. Carpenter
Professor of Psychiatry and Human Behavior

Stephen R. Carr
Professor of Obstetrics and Gynecology

Mary A. Carskadon
Professor of Psychiatry and Human Behavior

E. Jane Carter
Professor of Medicine

Angelina C. A. Carvalho
Professor Emerita of Medicine

William J. Cashore
Professor Emeritus of Pediatrics

Kimberle C. Chapin
Professor of Pathology and Laboratory Medicine; Professor of Medicine

Qian Chen
Michael G. Ehrlich, MD Professor of Orthopedic Research, Professor of Medical Science

Gaurav Choudhary
Ruth and Paul Levinger Professor of Cardiology

Ming-Yu Wang Chu
Professor Emerita of Medicine (Research)

Shih-Hsi Chu
Professor Emeritus of Molecular Pharmacology, Physiology and Biotechnology (Research)

Thomas H. Chun
Professor of Emergency Medicine; Professor of Pediatrics

William Cioffi
J. Murray Beardsley Professor of Surgery

Brian Clyne
Professor of Emergency Medicine; Professor of Medical Science

Andrew Cohen
Professor of Medicine

Suzanne M. Colby
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Annette W. Coleman
Stephen T Olney Professor Emerita of Natural History

Barry W. Connors
L. Herbert Ballou University Professor Emeritus of Neuroscience, Professor Emeritus of Medical Science

Donald R. Coustan
Professor Emeritus of Obstetrics and Gynecology

Joseph Crisco
Henry F. Lippitt Professor of Orthopedics, Professor of Engineering (Research)

John J. Cronan
Charles and Elfriede Collis-Frances Weeden Gibson Professor of Diagnostic Imaging

James P. Crowley
Professor Emeritus of Medicine

Susan Cu-Uvin
Professor of Obstetrics and Gynecology; Professor of Health Services, Policy and Practice; Professor of Medicine

Michele G. Cyr
Professor of Medicine; Professor of Medical Science

Erika D'Agata
Professor of Medicine

Albert E. Dahlberg
Professor Emeritus of Medical Science

Robert Paul Davis
Professor Emeritus of Medical Science

Ivana Delalle
Professor of Pathology and Laboratory Medicine

Suzanne M. De La Monte
Professor of Pathology and Laboratory Medicine; Professor of Neurosurgery

Ronald A. De Lellis
Professor Emeritus of Pathology and Laboratory Medicine

Penelope Hill Dennehy
Professor of Pediatrics

Phyllis A. Dennery
Sylvia Kay Hassenfeld Professor of Pediatrics, Professor of Molecular Biology, Cell Biology and Biochemistry

Monique E. De Paepe
Professor of Pathology and Laboratory Medicine

Daniel P. Dickstein
Professor of Psychiatry and Human Behavior; Professor of Pediatrics

Paul A. Disilvestro
Professor of Obstetrics and Gynecology

Don S. Dizon
Professor of Medicine; Professor of Surgery

John P. Donoghue
Henry Merritt Wriston Professor of Neuroscience, Professor of Engineering

Gary S. Dorfman
Professor Emeritus of Diagnostic Imaging

Susan Duffy
Professor of Emergency Medicine; Professor of Pediatrics

Raymond G. Dufresne
Professor of Dermatology

J. Donald Easton
Professor Emeritus of Neurology

Charles B. Eaton
Professor of Family Medicine; Professor of Epidemiology

Craig P. Eberson
Professor of Orthopaedics

Lee Edstrom
Professor Emeritus of Surgery

Afshin Ehsan
Professor of Surgery

Wafik S. El-Deiry
Mencoff Family University Professor of Medical Science, Professor of Pathology and Laboratory Medicine

Jack A. Elias
Warren Alpert Foundation Professor of Translational Science, Professor of Molecular Microbiology and Immunology, Professor of Medicine, Professor of Molecular Biology, Cell Biology and Biochemistry

Anashua Rani Ghose Elwy
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Mel H. Epstein
Professor Emeritus of Neurosurgery

Paul Fadale
Professor of Orthopaedics

Mark J. Fagan
Professor Emeritus of Medicine

William G. Fairbrother
Professor of Biology

Justin R. Fallon
Professor of Medical Science; Professor of Psychiatry and Human Behavior

Anne Fausto-Sterling
Nancy Duke Lewis Professor Emerita of Biology

Patricia Jeanne Flanagan
Professor of Pediatrics

Timothy P. Flanigan
Professor of Medicine; Professor of Health Services, Policy and Practice

Braden C. Fleming
Lucy Lippitt Professor of Orthopaedics, Professor of Engineering

Edwin N. Forman
Professor Emeritus of Pediatrics

Richard N. Freiman
Professor of Molecular Biology, Cell Biology and Biochemistry

Jennifer F. Friedman
Professor of Pediatrics; Professor of Epidemiology

Joseph H. Friedman
Professor of Neurology

Gregory K. Fritz
Professor Emeritus of Psychiatry and Human Behavior

Generoso G. Gascon
Professor Emeritus of Pediatrics and Professor Emeritus of Neurology

Jennifer S. Gass
Professor of Surgery; Professor of Obstetrics and Gynecology

Stephen M. Gatesy
Professor of Biology; Professor of Medical Science

Brandon A. Gaudiano
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Susan A. Gerbi
George D. Eggleston Professor of Biochemistry

Reginald Y. Gohh
Professor of Medicine

Ziya L. Gokaslan
Professor of Neurosurgery

Amy P. Goldberg
Professor of Pediatrics

Richard J. Goldberg
Professor Emeritus of Psychiatry and Human Behavior

George E. Goslow
Professor Emeritus of Biology

Janet Grace
Professor Emerita of Psychiatry and Human Behavior

David J. Grand
Professor of Diagnostic Imaging

Stefan Gravenstein
David S. Greer Professor of Geriatric Medicine, Professor of Health Services, Policy and Practice

Andrew Green
Professor of Orthopaedics

Benjamin D. Greenberg
Professor of Psychiatry and Human Behavior

Paul B. Greenberg
Professor of Surgery

Samuel H. Greenblatt
Professor Emeritus of Neurosurgery

Philip A. Gruppuso
Professor of Pediatrics; Professor of Medical Science; Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Fusun Gundogan
Professor of Pathology and Laboratory Medicine

Kate M. Guthrie
Professor of Psychiatry and Human Behavior

James E. Haddow
Professor Emeritus of Pathology and Laboratory Medicine

Chi-Ming Hai
Professor Emeritus of Molecular Pharmacology, Physiology and Biotechnology

Brittany Star Hampton
Professor of Obstetrics and Gynecology; Professor of Medical Science

David T. Harrington
Professor of Surgery

Elizabeth O. Harrington
Professor of Medicine

Anne C. Hart
Professor of Neuroscience

Edward Hawrot
Alva O. Way Professor of Medical Science

Roman A. Hayda
Professor of Orthopaedics

Stephen L. Helfand
George D. Eggleston Professor of Biochemistry

Peter Heywood
Professor of Biology

Douglas C. Hixson
Professor Emeritus of Medicine (Research) and Professor Emeritus of Pathology and Laboratory Medicine (Research)

Frederic G. Hoppin
Professor Emeritus of Medicine and Professor Emeritus of Physiology

Michael Jan Hulstyn
Professor of Orthopaedics

Jeffrey I. Hunt
Professor of Psychiatry and Human Behavior

Vincent R. Hunt
Professor Emeritus of Family Medicine

Cynthia L. Jackson
Professor of Pathology and Laboratory Medicine

Ivor M. D. Jackson
Professor Emeritus of Medicine

Mukesh Jain
Frank L. Day Professor of Biology, Professor of Molecular Biology, Cell Biology and Biochemistry, Professor of Medicine

Christine Marie Janis
Professor Emerita of Ecology and Evolutionary Biology

Gregory D. Jay
John and Mary Panton and Emergency Medicine Professor of Translational Science, Professor of Medicine, Professor of Orthopaedics

Mahesh V. Jayaraman
Professor of Diagnostic Imaging; Professor of Neurology; Professor of Neurosurgery

Elissa Jelalian
Professor of Psychiatry and Human Behavior; Professor of Pediatrics

Conrad E. Johanson
Professor Emeritus of Neurosurgery

Lenworth N. Johnson
Professor of Surgery

Rich Jones
Professor of Psychiatry and Human Behavior; Professor of Neurology

Agnes B. Kane
Professor Emerita of Pathology and Laboratory Medicine

Rami Kantor
Professor of Medicine

Julia A. Katarincic
Professor of Orthopaedics

William E. Kaye
Professor Emeritus of Surgery and Professor Emeritus of Medicine

Gabor I. Keitner
Professor of Psychiatry and Human Behavior

Martin B. Keller
Professor Emeritus of Psychiatry and Human Behavior

Martin Keszler
Professor of Pediatrics

Nabil Y. Khoury
Professor Emeritus of Obstetrics and Gynecology

Louise Sadler Kiessling
Professor Emerita of Family Medicine and Professor Emerita of Pediatrics

Peter Kim
Professor of Surgery

Timothy Kinsella
Professor Emeritus of Radiation Oncology

Petra Klinge
Professor of Neurosurgery

James R. Klinger
Professor of Medicine

Leo Kobayashi
Professor of Emergency Medicine

Gideon Koren
Professor of Medicine

Jonathan David Kurtis
Professor of Pathology and Laboratory Medicine

W. Curt LaFrance
Professor of Psychiatry and Human Behavior; Professor of Neurology

Edward V. Lally
Professor of Medicine

Arthur Landy
University Professor Emeritus of Medical Science

Abbot Laptook
Professor of Pediatrics

W. Dwayne Lawrence
Professor Emeritus of Pathology and Laboratory Medicine

Neal S. Leleiko
Professor Emeritus of Pediatrics

Louis A. Leone
Professor Emeritus of Medicine

Barry M. Lester
Professor of Psychiatry and Human Behavior; Professor of Pediatrics

Adam Levine
Professor of Emergency Medicine

Mitchell M. Levy
Professor of Medicine

William J. Lewander
Professor of Emergency Medicine; Professor of Pediatrics

James G. Linakis
Professor Emeritus of Emergency Medicine

Diane Lipscombe
Thomas J. Watson, Sr. Professor of Science, Professor of Neuroscience

Paul Liu
Professor of Surgery

Debra J. Lobato
Professor Emerita of Psychiatry and Human Behavior

Richard H. Longabaugh
Professor Emeritus of Psychiatry and Human Behavior

Sydney Louis
Professor Emeritus of Neurology

Ana P. Lourenco
Professor of Diagnostic Imaging

Francois I. Luks
Professor of Surgery; Professor of Obstetrics and Gynecology; Professor of Pediatrics

Martha B. Mainiero
Professor of Diagnostic Imaging

Paul F. Malloy
Professor Emeritus of Psychiatry and Human Behavior

David E. Mandelbaum
Professor Emeritus of Pediatrics

Anthony Lynn Mansell
Professor Emeritus of Pediatrics

Jill L. Maron
Professor of Pediatrics

John Marshall
Professor of Medical Science

Edith Mathiowitz
Professor of Pathology and Laboratory Medicine; Professor of Engineering

Kathleen M. McCarten
Professor Emerita of Diagnostic Imaging, Clinician Educator and Professor Emerita of Pediatrics, Clinician Educator

F. Dennis McCool
Professor of Medicine

Kelly Ann McGarry
Professor of Medicine

James T. McIlwain
Sidney A. and Dorothea Doctors Fox Professor Emeritus of Ophthalmology and Visual Science, Professor Emeritus of Surgery, Professor Emeritus of Neuroscience

Elizabeth L. McQuaid
Professor of Psychiatry and Human Behavior; Professor of Pediatrics

Antone A. Medeiros
Professor Emeritus of Medicine

Michael J. Mello
Professor of Emergency Medicine; Professor of Health Services, Policy and Practice; Professor of Medical Science

Ulrike Mende
Professor of Medicine

Leonard A. Mermel
Professor of Medicine

Geralyn M. L. Messerlian
Professor of Pathology and Laboratory Medicine; Professor of Obstetrics and Gynecology

Ivan W. Miller
Professor of Psychiatry and Human Behavior

Kenneth Raymond Miller
Professor Emeritus of Molecular Biology, Cell Biology and Biochemistry

Richard P. Millman
Professor of Medicine; Professor of Pediatrics

Thomas J. Miner
Professor of Surgery

Robert Miranda
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Christopher I. Moore
Professor of Neuroscience

Jeffrey R. Morgan
Professor of Pathology and Laboratory Medicine; Professor of Engineering

David J. Morris
Professor Emeritus of Pathology and Laboratory Medicine

Paul E. Morrissey
Professor of Surgery

Eric M. Morrow
Mencoff Family Professor of Biology, Professor of Neuroscience, Professor of Psychiatry and Human Behavior

Douglass H. Morse
Hermon Bumpus Professor Emeritus of Biology, Professor of Ecology and Evolutionary Biology (Research)

Steven F. Moss
Professor of Medicine

Albert S. Most
Professor Emeritus of Medicine

Kimberly L. Mowry
Robin Chemers Neustein Professor of Biomedicine

John B. Murphy
Professor Emeritus of Medicine and Professor Emeritus of Family Medicine

Deborah L. Myers
Professor of Obstetrics and Gynecology

Wade Myers
Professor of Psychiatry and Human Behavior

Eleftherios E. Mylonakis
Charles C.J. Carpenter, MD Professor of Infectious Diseases, Professor of Medicine, Professor of Molecular Microbiology and Immunology

Eduardo A. Nillni
Professor Emeritus of Medicine (Research) and Professor Emeritus of Molecular Biology, Cell Biology and Biochemistry (Research)

Georg Noren
Professor Emeritus of Neurosurgery

Bonnie O'Connor
Professor Emerita of Pediatrics, Clinician Educator

Thomas P. O'Toole
Professor of Medicine; Professor of Health Services, Policy and Practice

Elena Oancea
Professor of Medical Science

William Oh
Sylvia Kay Hassenfeld Professor Emeritus of Pediatrics

Frank L. Overly
Professor of Emergency Medicine; Professor of Pediatrics

Brett Douglas Owens
Professor of Orthopaedics; Professor of Epidemiology

Adetokunbo A. Oyelese
Professor of Neurosurgery

Bai-Chuan Pan
Professor Emeritus of Medicine (Research)

Michael A. Paradiso
Sidney A. Fox and Dorothea Doctors Fox Professor of Ophthalmology and Visual Science, Professor of Neuroscience

Gyan Pareek
Professor of Surgery; Professor of Medicine

Teri B. Pearlstein
Professor of Psychiatry and Human Behavior; Professor of Medicine

Georges Peter
Professor Emeritus of Pediatrics

Noah S. Philip
Professor of Psychiatry and Human Behavior

Maureen Phipps
Professor Emerita of Obstetrics and Gynecology

M. Halit Pinar
Professor of Pathology and Laboratory Medicine

Stephen Porder
Professor of Ecology, Evolution, and Organismal Biology and Environment and Society

Raymond O. Powrie
Professor of Medicine; Professor of Obstetrics and Gynecology

Lawrence H. Price
Professor of Psychiatry and Human Behavior

M. Quddus
Professor of Pathology and Laboratory Medicine

Peter J. Quesenberry
Paul Calabresi, M.D. Professor of Oncology, Professor of Medicine

Abrar Qureshi
Warren Alpert Foundation Professor of Dermatology, Professor of Epidemiology

Glenn Radice
Professor of Medicine

Sohini Ramachandran
Professor of Biology; Professor of Computer Science

David M. Rand
Stephen T. Olney Professor of Natural History

Charles R. Rardin
Professor of Obstetrics and Gynecology

Robert A. Reenan
Professor of Biology

Murray Resnick
Professor of Pathology and Laboratory Medicine

Louis B. Rice
Professor of Medicine; Professor of Molecular Microbiology and Immunology

Josiah D. Rich
Professor of Medicine; Professor of Epidemiology

Mark J. Rivard
Professor of Radiation Oncology

Thomas Roberts
Professor of Ecology, Evolution, and Organismal Biology

Leslie Robinson-Bostom
Professor of Dermatology

Randy Rockney
Professor Emeritus of Pediatrics, Professor Emeritus of Family Medicine, and Professor Emeritus of Medical Science

Karen Rosene Montella
Professor Emerita of Medicine

Frank G. Rothman
Professor Emeritus of Biology

Sharon I. Rounds
Professor of Medicine; Professor of Pathology and Laboratory Medicine

Dwight J. Rouse
Professor of Obstetrics and Gynecology; Professor of Epidemiology

Ranna Rozenfeld
Professor of Pediatrics

James L. Rudolph
Professor of Medicine; Professor of Health Services, Policy and Practice

Raymond Richard Russell
Professor of Medicine

Stephen P. Salloway
Martin M. Zucker Professor of Psychiatry and Human Behavior, Professor of Neurology

Arthur R. Salomon
Professor of Molecular Biology, Cell Biology, and Biochemistry; Professor of Chemistry

Jerome Sanes
Professor of Neuroscience

Dov F. Sax
Professor of Ecology, Evolution, and Organismal Biology; Professor of Environment and Society

Fred Jay Schiffman
Sigal Family Professor of Humanistic Medicine, Professor of Medicine

Jeremiah Schuur
Frances Weeden Gibson-Edward A. Iannuccilli, MD Professor of Emergency Medicine, Professor of Health Services, Policy and Practice

Robert Schwartz
Professor Emeritus of Pediatrics

H. Denman Scott
Professor Emeritus of Medicine and Professor Emeritus of Health Services, Policy and Practice

John M. Sedivy
Hermon C. Bumpus Professor of Biology

Frank W. Sellke
Karl E. Karlson, MD and Gloria A. Karlson Professor of Cardiothoracic Surgery, Professor of Surgery

Peter R. Shank
Professor Emeritus of Medical Science

M. Tracie Shea
Professor of Psychiatry and Human Behavior

David Sheinberg
Professor of Neuroscience

Mark Sigman
Krishnamurthi Family Professor of Urology, Professor of Surgery, Professor of Pathology and Laboratory Medicine

Fredric Joel Silverblatt
Professor Emeritus of Medicine

James A. Simmons
Professor of Biology

Don B. Singer
Professor Emeritus of Pathology and Laboratory Medicine

Alan D. Sirota
Professor Emeritus of Psychiatry and Human Behavior, Clinician Educator

Robert J. Smith
Professor Emeritus of Medicine

Stephen R. Smith
Professor Emeritus of Family Medicine

Peter Snyder
Professor of Neurology; Professor of Surgery

Anthony Spirito
Professor of Psychiatry and Human Behavior

Dale W. Steele
Professor of Emergency Medicine; Professor of Health Services, Policy and Practice; Professor of Pediatrics

Barry S. Stein
Professor Emeritus of Surgery

Margaret M. Steinhoff
Professor Emerita of Pathology and Laboratory Medicine

Edward S. Sternick
Professor Emeritus of Radiation Oncology

Barbara S. Stonestreet
Professor Emerita of Pediatrics

Laura Stroud
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences; Professor of Obstetrics and Gynecology

Selim Suner
Professor of Emergency Medicine; Professor of Surgery

C. James Sung
Professor of Pathology and Laboratory Medicine

Vivian W. Sung
Professor of Obstetrics and Gynecology

Sharon M. Swartz
Professor of Biology; Professor of Engineering

Joseph D. Sweeney
Professor of Pathology and Laboratory Medicine; Professor of Medicine

Patrick J. Sweeney
Professor Emeritus of Obstetrics and Gynecology

Robert M. Swift
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Dominick Tammaro
Professor of Medicine

Karen T. Tashima
Professor of Medicine

Marc Tatar
Professor of Biology

Nina Tatevian
Professor of Pathology and Laboratory Medicine

Richard M. Terek
Professor of Orthopaedics

Walter R. Thayer
Professor Emeritus of Medical Science

Nancy L. Thompson
Professor Emerita of Medicine (Research) and Professor Emerita of Pathology and Laboratory Medicine (Research)

Steven A. Toms
Professor of Neurosurgery; Professor of Medicine

Peter G. Trafton
Professor Emeritus of Orthopaedics

Stephen Traub
Professor of Emergency Medicine

Glenn A. Tung
Professor of Diagnostic Imaging

Allan R. Tunkel
Professor of Medicine; Professor of Medical Science

Methodius Tuuli
Chace-Joukowsky Professor of Obstetrics and Gynecology

Audrey R. Tyrka
Professor of Psychiatry and Human Behavior

Lisa A. Uebelacker
Professor of Psychiatry and Human Behavior

Jonathan Valente
Professor of Emergency Medicine; Professor of Pediatrics

Herman H. Vandenburgh
Professor Emeritus of Pathology and Laboratory Medicine and Professor Emeritus of Molecular Pharmacology, Physiology and Biotechnology

Rosalind M. Vaz
Professor Emerita of Pediatrics, Clinician Educator

Michael P. Vezeridis
Professor of Surgery

Betty R. Vohr
Professor of Pediatrics

Jonathan K. Waage
Professor Emeritus of Biology

Jack R. Wands
Jeffrey and Kimberly Greenberg-Artemis and Martha Joukowsky Professor of Gastroenterology, Professor of Medicine, Professor of Medical Science

Li Juan Wang
Professor of Pathology and Laboratory Medicine

Daniel M. Weinreich
Royce Family Professor of Teaching Excellence in Biology

Lauren M. Weinstock
Professor of Psychiatry and Human Behavior

Martin A. Weinstock
Professor of Dermatology; Professor of Epidemiology

Arnold-Peter Weiss
Professor of Orthopaedics

Gary M. Wessel
Professor of Biology

Conrad W. Wesselhoeft
Professor Emeritus of Surgery, Clinician Educator

Robert J. Westlake
Professor Emeritus of Psychiatry and Human Behavior

Kristi A. Wharton
Professor of Biology

Edward Wing
Professor Emeritus of Medical Science

Rena R. Wing
Professor of Psychiatry and Human Behavior

Marion F. Winkler
Professor of Surgery

Jon D. Witman
Professor of Biology

Eric T. Wong
Professor of Neurology; Professor of Medicine; Professor of Neurosurgery; Professor of Radiation Oncology

Wen-Chih Wu
Professor of Medicine; Professor of Epidemiology

Evgeny Yakirevich
Professor of Pathology and Laboratory Medicine

Ronald A. Yankee
Professor Emeritus of Medicine

Tao Zheng
Professor of Pediatrics; Professor of Medicine; Professor of Molecular Microbiology and Immunology (Research)

Anatoly Zhitkovich
Professor of Medical Science

Anita L. Zimmerman
Professor of Molecular Microbiology and Immunology

Mark Zimmerman
Professor of Psychiatry and Human Behavior

Caron Zlotnick
Professor of Psychiatry and Human Behavior; Professor of Medicine; Professor of Obstetrics and Gynecology

Mark Robert Zonfrillo
Professor of Emergency Medicine; Professor of Pediatrics

Professor Research

Adam Chodobski
Professor Emeritus of Emergency Medicine (Research)

Alfred Ayala
Professor of Surgery (Research)

Robbert J. Creton
Professor of Medical Science (Research); Professor of Pathology and Laboratory Medicine (Research)

Jennifer B. Freeman
Professor of Psychiatry and Human Behavior (Research)

Leslie B. Gordon
Professor of Pediatrics (Research)

Anthony Richard Hayward
Professor of Pediatrics (Research)

Marshall Kadin
Professor of Pathology and Laboratory Medicine (Research)

Daphne Koinis-Mitchell
Professor of Psychiatry and Human Behavior (Research); Professor of Pediatrics (Research)

Chun Geun Lee
Professor of Molecular Microbiology and Immunology (Research)

Richard L. Marsh
Professor of Ecology, Evolution, and Organismal Biology (Research)

Glenn E. Palomaki
Professor of Pathology and Laboratory Medicine (Research)

Susan Ramsey
Professor of Psychiatry and Human Behavior (Research); Professor of Medicine (Research)

Jonathan S. Reichner
Professor of Surgery (Research)

Carl Saab
Professor of Neurosurgery (Research); Professor of Neuroscience (Research)

Surendra Sharma
Professor of Pediatrics (Research); Professor of Pathology and Laboratory Medicine (Research)

Graham Thomas
Professor of Psychiatry and Human Behavior (Research)

Shuping Tong
Professor of Medicine (Research)

Ting Zhao
Professor of Surgery (Research)

Shougang Zhuang
Professor of Medicine (Research)

Clinical Professor

Scott David Berns
Clinical Professor of Pediatrics; Clinical Professor of Health Services, Policy and Practice

Rosemarie Bigsby
Clinical Professor Emeritus of Pediatrics

Andrew S. Blazar
Clinical Professor Emeritus of Obstetrics and Gynecology

Walter A. Brown
Clinical Professor Emeritus of Psychiatry and Human Behavior

Gary Bubly
Clinical Professor of Emergency Medicine; Clinical Professor of Medicine

Frederick W. Burgess
Clinical Professor Emeritus of Surgery

Marshall Carpenter
Clinical Professor of Obstetrics and Gynecology

Joseph Chazan
Clinical Professor Emeritus of Medicine

William M. Corrao
Clinical Professor of Medicine

Robert Crausman
Clinical Professor of Medicine

Catherine A. Cummings
Clinical Professor of Emergency Medicine

Karen Daigle
Clinical Professor of Pediatrics

David Egilman
Clinical Professor of Family Medicine

Yul D. Ejnes
Clinical Professor of Medicine

Lynn C. Epstein
Clinical Professor Emerita of Psychiatry and Human Behavior and Clinical Professor Emerita of Behavioral and Social Sciences

Edward R. Feller
Clinical Professor of Medical Science

Fred F. Ferri
Clinical Professor of Medicine

B. Allen Flaxman
Clinical Professor Emeritus of Dermatology

Gary N. Frishman
Clinical Professor of Obstetrics and Gynecology

Maria Luisa Garcia-Moliner
Clinical Professor of Pathology and Laboratory Medicine

Roberta E. Goldman
Clinical Professor of Family Medicine

Debra S. Herman
Clinical Professor of Psychiatry and Human Behavior

Kathleen C. Hittner
Clinical Professor Emerita of Surgery

Horacio B. Hojman
Clinical Professor of Psychiatry and Human Behavior

Karen A. Holler
Clinical Professor of Psychiatry and Human Behavior

Edward A. Iannuccilli
Clinical Professor Emeritus of Medicine

Julianne Ip
Clinical Professor Emerita of Family Medicine abd Clinical Professor Emerita of Medical Science

Charles B. Kahn
Clinical Professor Emeritus of Medicine

Achyut Kamat
Clinical Professor of Emergency Medicine

Stephen R. Kaplan
Clinical Professor Emeritus of Medicine

Joel M. Kaufman
Clinical Professor of Neurology

Peter D. Kramer
Clinical Professor Emeritus of Psychiatry and Human Behavior

Carol Landau
Clinical Professor Emerita of Psychiatry and Human Behavior and Clinical Professor Emerita of Medicine

Mary D. Lekas
Clinical Professor Emerita of Surgery

Daniel J. Levine
Clinical Professor of Medicine

Susanna R. Magee
Clinical Professor of Family Medicine

Hon Fong Louie Mark
Clinical Professor Emerita of Pediatrics

Martin M. Miner
Clinical Professor of Family Medicine; Clinical Professor of Surgery

Jack M. Monchik
Clinical Professor Emeritus of Surgery

James E. Monti
Clinical Professor of Emergency Medicine

Anthony Moulton
Clinical Professor of Medical Science

Andrew T. Nathanson
Clinical Professor of Emergency Medicine

Elizabeth M. Nestor
Clinical Professor Emerita of Emergency Medicine

Steven M. Opal
Clinical Professor of Medicine

John R. Parziale
Clinical Professor of Orthopaedics

Robert B. Patterson
Clinical Professor of Surgery

Wendy A. Plante
Clinical Professor of Psychiatry and Human Behavior

David C. Portelli
Clinical Professor of Emergency Medicine

Victor E. Pricolo
Clinical Professor of Medical Science

Herbert Rakatansky
Clinical Professor Emeritus of Medicine

Patricia R. Recupero
Clinical Professor of Psychiatry and Human Behavior; Clinical Professor of Medical Science

Michelle L. Rickerby
Clinical Professor of Psychiatry and Human Behavior

Stephen N. Rous
Clinical Professor Emeritus of Surgery

Steven Schechter
Clinical Professor of Surgery

Francis H. Scola
Clinical Professor Emeritus of Diagnostic Imaging

Thomas E. Sepe
Clinical Professor of Medicine

Russell Settipane
Clinical Professor of Medicine

Samir A. Shah
Clinical Professor of Medicine

Arun Singh
Clinical Professor Emeritus of Surgery

Boris Skurkovich
Clinical Professor of Pediatrics

Tracy O'Leary Tevyaw
Clinical Professor of Psychiatry and Human Behavior

King W. To
Clinical Professor of Surgery

William G. Tsiaras
Clinical Professor Emeritus of Surgery

Barbara Tylenda
Clinical Professor Emerita of Psychiatry and Human Behavior

Marguerite B. Vigliani
Clinical Professor Emerita of Obstetrics and Gynecology

Hedy S. Wald
Clinical Professor of Family Medicine

Barry W. Wall
Clinical Professor of Psychiatry and Human Behavior

Anne S. Walters
Clinical Professor of Psychiatry and Human Behavior

Marjorie E. Weishaar
Clinical Professor Emerita of Psychiatry and Human Behavior

Joel K. Weltman
Clinical Professor Emeritus of Medicine

Russell E. White
Clinical Professor of Surgery

John P. Wincze
Clinical Professor Emeritus of Psychiatry and Human Behavior

Therese Zink
Clinical Professor of Family Medicine; Clinical Professor of Behavioral and Social Sciences

Visiting Professor

Darlene R. Ketten
Visiting Professor of Neuroscience

Associate Professor

J. Gary Abuelo
Associate Professor Emeritus of Medicine

Sun Ho Ahn
Associate Professor of Diagnostic Imaging

Umer Akbar
Associate Professor of Neurology

Rebecca H. Allen
Associate Professor of Obstetrics and Gynecology; Associate Professor of Medical Science

Siraj Amanullah
Associate Professor of Emergency Medicine; Associate Professor of Health Services, Policy and Practice; Associate Professor of Pediatrics

Ali Amin
Associate Professor of Pathology and Laboratory Medicine

David C. Anthony
Associate Professor of Family Medicine

Valentin Antoci
Associate Professor of Orthopaedics

Herbert D. Aronow
Associate Professor of Medicine

Wael Asaad
Sidney A. Fox and Dorothea Doctors Fox Associate Professor of Ophthalmology and Visual Science, Associate Professor of Neuroscience

Subhashini Ayloo
Associate Professor of Surgery

Jeffrey A. Bailey
Mencoff Family Associate Professor of Translational Research, Associate Professor of Pathology and Laboratory Medicine

Grayson L. Baird
Associate Professor of Diagnostic Imaging

Barbara E. Barker
Associate Professor Emerita of Pathology and Laboratory Medicine

Thomas B. Bartnikas
Associate Professor of Pathology and Laboratory Medicine

George P. Bayliss
Associate Professor of Medicine

Peter A. Belenky
Associate Professor of Molecular Microbiology and Immunology

James M. Brindle
Associate Professor of Radiation Oncology

Willoughby B. Britton
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Behavioral and Social Sciences

Jonathan F. Cahill
Associate Professor of Neurology

Weibiao Cao
Associate Professor of Pathology and Laboratory Medicine; Associate Professor of Medicine

Chung-Ja M. Cha
Associate Professor Emerita of Pediatrics (Research)

Philip A. Chan
Associate Professor of Medicine; Associate Professor of Behavioral and Social Sciences

Elizabeth S. Chen
Associate Professor of Medical Science; Associate Professor of Health Services, Policy and Practice

Eunyoung Cho
Associate Professor of Dermatology; Associate Professor of Epidemiology

Paul P. Christopher
Associate Professor of Psychiatry and Human Behavior

Jennifer G. Clarke
Associate Professor of Medicine; Associate Professor of Obstetrics and Gynecology

Cheston B. Cunha
Associate Professor of Medicine

Shawna M. Cutting
Associate Professor of Neurology

Alan H. Daniels
Associate Professor of Orthopaedics

Eric M. Darling
Associate Professor of Medical Science; Associate Professor of Engineering; Associate Professor of Orthopaedics

Manuel F. Dasilva
Associate Professor of Orthopaedics

Christopher L. de Graffenried
Associate Professor of Molecular Microbiology and Immunology

Alison DeLong
Associate Professor of Biology

Joseph Anthony Diaz
Associate Professor of Medicine; Associate Professor of Medical Science

Thomas Dipetrillo
Associate Professor of Radiation Oncology

John E. Donahue
Associate Professor of Pathology and Laboratory Medicine; Associate Professor of Neurology

David M. Dosa
Associate Professor of Medicine; Associate Professor of Health Services, Policy and Practice

Luba Dumenco
Associate Professor of Medical Science; Associate Professor of Pathology and Laboratory Medicine

Jennifer Eaton
Associate Professor of Obstetrics and Gynecology

Thomas K. Egglin
Associate Professor of Diagnostic Imaging

Gary P. Epstein-Lubow
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Health Services, Policy and Practice; Associate Professor of Medical Science

Allan Erickson
Associate Professor Emeritus of Medicine

Dimitrios Farmakiotis
Associate Professor of Medicine

Loren D. Fast
Associate Professor Emeritus of Medicine

Nicolas Lux Fawzi
Associate Professor of Medical Science

Lloyd R. Feit
Associate Professor of Pediatrics

Jun Feng
Associate Professor of Surgery

Alexander Fleischmann
Provost's Associate Professor of Brain Science

Greta Francis
Associate Professor of Psychiatry and Human Behavior

Hamish S. Fraser
Associate Professor of Medical Science; Associate Professor of Health Services, Policy and Practice

Matthew Fuxjager
Associate Professor of Ecology, Evolution, and Organismal Biology

Rebekah L. Gardner
Associate Professor of Medicine

Eric J. Gartman
Associate Professor of Medicine

Elizabeth M. Goldberg
Associate Professor of Emergency Medicine; Associate Professor of Health Services, Policy and Practice

Dragan Golijanin
Associate Professor of Surgery

Christopher J. Got
Associate Professor of Orthopaedics

Rogers C. Griffith
Associate Professor of Pathology and Laboratory Medicine

Carolina L. Haass-Koffler
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Behavioral and Social Sciences

Katrine Hansen
Associate Professor of Pathology and Laboratory Medicine

Terrance T. Healey
Associate Professor of Diagnostic Imaging

Daithi J. Heffernan
Associate Professor of Surgery

Diane Hoffman-Kim
Associate Professor of Neuroscience; Associate Professor of Engineering

Andrew W. Holowinsky
Associate Professor Emeritus of Molecular Biology, Cell Biology and Biochemistry

Emilia Huerta-Sanchez
Associate Professor of Ecology, Evolution, and Organismal Biology

Amanda M. Jamieson
Associate Professor of Molecular Microbiology and Immunology

Matthew D. Jankowich
Associate Professor of Medicine

Alexander Jaworski
June G. Zimmerman Associate Professor of Brain Science

Ernestine G. Jennings
Associate Professor of Psychiatry and Human Behavior

Gerwald Jogl
Associate Professor of Biology

Mark A. Johnson
Associate Professor of Biology

Stephanie R. Jones
Associate Professor of Neuroscience

Steven C. Katz
Associate Professor of Surgery

Karla Kaun
Associate Professor of Neuroscience

James R. Kellner
Associate Professor of Ecology, Evolution, and Organismal Biology and Environment and Society

Mark Kendall
Associate Professor of Anesthesiology

Noubar Kessimian
Associate Professor Emeritus of Pathology and Laboratory Medicine, Clinician Educator

Kenneth S. Korr
Associate Professor Emeritus of Medicine

Michelle A. Lally
Associate Professor of Medicine; Associate Professor of Health Services, Policy and Practice

Robert E. Lambiase
Associate Professor Emeritus of Diagnostic Imaging

Erica N. Larschan
Associate Professor of Molecular Biology, Cell Biology and Biochemistry

Lucia Larson
Associate Professor of Medicine

Sean Lawler
Associate Professor of Pathology and Laboratory Medicine

Elizabeth Lazarus
Associate Professor of Diagnostic Imaging

Beatrice E. Lechner
Associate Professor of Pediatrics

Kara Lynne Leonard
Associate Professor of Radiation Oncology

Otto Liebmann
Associate Professor of Emergency Medicine

John R. Lonks
Associate Professor of Medicine; Associate Professor of Medical Science

Shaolei Lu
Associate Professor of Pathology and Laboratory Medicine

Rishi R. Lulla
Alan G. Hassenfeld Associate Professor of Pediatrics

Tracy E. Madsen
Associate Professor of Emergency Medicine; Associate Professor of Epidemiology

Katherine E. Mason
Associate Professor of Pediatrics

Cara A. Mathews
Associate Professor of Obstetrics and Gynecology

James R. McCartney
Associate Professor Emeritus of Psychiatry and Human Behavior

Patrick T. McGann
Associate Professor of Pediatrics; Associate Professor of Medicine

John E. McGeary
Associate Professor of Psychiatry and Human Behavior

Elisabeth C. McGowan
Associate Professor of Pediatrics

George R. McKendall
Associate Professor of Medicine

Nicole McLaughlin
Associate Professor of Psychiatry and Human Behavior

Suzanne McLaughlin
Associate Professor of Pediatrics and Medicine

Christopher H. Merritt
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

Maria D. Mileno
Associate Professor of Medicine

Margaret Miller
Associate Professor of Medicine

Ethan Moitra
Associate Professor of Psychiatry and Human Behavior

Christine E. Montross
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Medical Science

Alan R. Morrison
Associate Professor of Medicine

Aman Nanda
Associate Professor of Medicine

Anthony M. Napoli
Associate Professor of Emergency Medicine

Marguerite A. Neill
Associate Professor of Medicine

Nicola Neretti
Associate Professor of Molecular Biology, Cell Biology, and Biochemistry

Christian Parcher Nixon
Associate Professor of Pathology and Laboratory Medicine

Nicole R. Nugent
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Emergency Medicine

James A. O'Brien
Associate Professor Emeritus of Obstetrics and Gynecology

Kate O'Connor-Giles
Provost's Associate Professor of Brain Science

Hwamee Oh
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Cognitive, Linguistic and Psychological Sciences

Adam Olszewski
Associate Professor of Medicine

Joyce Ou
Associate Professor of Pathology and Laboratory Medicine; Associate Professor of Medical Science

Stephanie Parade
Associate Professor of Psychiatry and Human Behavior

Robert L. Patrick
Associate Professor Emeritus of Neuroscience

E. Scott Paxton
Associate Professor of Orthopaedics

Megan Pinkston-Camp
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Medicine

Patricia Poitevien
Associate Professor of Pediatrics

Kittichai Promrat
Associate Professor of Medicine

Bharat Ramratnam
Associate Professor of Medicine

John L. Reagan
Associate Professor of Medicine

Harlan G. Rich
Associate Professor of Medicine; Associate Professor of Medical Science

Alison J. Riese
Associate Professor of Pediatrics; Associate Professor of Medical Science

Abdalla Rifai
Associate Professor Emeritus of Pathology and Laboratory Medicine

Julie L. Roth
Associate Professor of Neurology; Associate Professor of Medical Science

Steven C. Rougas
Associate Professor of Emergency Medicine; Associate Professor of Medical Science

Elena Salmoirago-Blotcher
Associate Professor of Medicine; Associate Professor of Epidemiology; Associate Professor of Psychiatry and Human Behavior

Juan Sanchez-Esteban
Associate Professor of Pediatrics

Neil Sarkar
Associate Professor of Medical Science; Associate Professor of Health Services, Policy and Practice

Frank John Schaberg
Associate Professor Emeritus of Surgery, Clinician Educator

Jonathan R. Schiller
Associate Professor of Orthopaedics

Mark Schleinitz
Associate Professor of Medicine

Barry L. Sharaf
Associate Professor of Medicine

Katherine M. Sharkey
Associate Professor of Medicine; Associate Professor of Psychiatry and Human Behavior

Michael F. Sheff
Associate Professor Emeritus of Pathology and Laboratory Medicine (Research)

Douglas G. Shemin
Associate Professor of Medicine

Kamaljeet Singh
Associate Professor of Pathology and Laboratory Medicine

Katherine F. Smith
Associate Professor of Medical Science

Gregory M. Soares
Associate Professor of Diagnostic Imaging

Neel R. Sodha
Associate Professor of Surgery

Ashley R. Stuckey
Associate Professor of Obstetrics and Gynecology; Associate Professor of Medical Science

Patrick K. Sullivan
Associate Professor of Surgery

Lisa Michelle Swartz Topor
Associate Professor of Pediatrics

Daria Szkwarko
Associate Professor of Family Medicine

Joanna Szmydynger-Chodobska
Associate Professor Emerita of Emergency Medicine

Gladys H. Telang
Associate Professor of Dermatology

Simone Thavaseelan
Associate Professor of Surgery

Bradford B. Thompson
Associate Professor of Neurology; Associate Professor of Neurosurgery

Iris L. Tong
Associate Professor of Medicine

Diana O. Treaba
Associate Professor of Pathology and Laboratory Medicine

Geoffrey Tremont
Associate Professor of Psychiatry and Human Behavior

Wilson Truccolo
Pablo J. Salame Goldman Sachs Associate Professor of Computational Neuroscience

Robert J. Tubbs
Associate Professor of Emergency Medicine

Jennifer Unger
Associate Professor of Obstetrics and Gynecology

Shipra Vaishnava
Associate Professor of Molecular Microbiology and Immunology

Gregorio Valdez
GLF Translational Associate Professor of Molecular Biology, Cell Biology and Biochemistry

Corey E. Ventetuolo
Associate Professor of Medicine; Associate Professor of Health Services, Policy and Practice

Roxanne Vrees
Associate Professor of Obstetrics and Gynecology

Yihong Wang
Associate Professor of Pathology and Laboratory Medicine

Sarita S. Warrier
Associate Professor of Medicine; Associate Professor of Medical Science

Jennifer Greene Welch
Associate Professor of Pediatrics

Linda C. Wendell
Associate Professor of Neurology; Associate Professor of Medical Science; Associate Professor of Neurosurgery

Laura Brave Whiteley
Associate Professor of Psychiatry and Human Behavior

Joanne E. Wilkinson
Associate Professor of Family Medicine

Robyn Wing
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

Albert S. Woo
Associate Professor of Surgery; Associate Professor of Neurosurgery; Associate Professor of Pediatrics

Israel Yaar
Associate Professor Emeritus of Neurology

Shadi Yaghi
Associate Professor of Neurology

Wentian Yang
Associate Professor of Orthopaedics; Associate Professor of Medicine

Cunxian Zhang
Associate Professor of Pathology and Laboratory Medicine

Richard J. Zienowicz
Associate Professor of Surgery

Clinical Associate Professor

Khaja Ahmed
Clinical Associate Professor of Medicine

Paul E. Alexander
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior

Marla Cathleen Angermeier
Clinical Associate Professor Emerita of Dermatology

Naureen Attiullah
Clinical Associate Professor of Psychiatry and Human Behavior

Genie L. Bailey
Clinical Associate Professor of Psychiatry and Human Behavior

Mohammad Banki
Clinical Associate Professor of Surgery

Karyn K. Blane
Clinical Associate Professor of Psychiatry and Human Behavior

Thomas A. Bledsoe
Clinical Associate Professor of Medicine

Stanley Hoyt Block
Clinical Associate Professor Emeritus of Pediatrics

John R. Boekamp
Clinical Associate Professor of Psychiatry and Human Behavior

Kathleen Cote Bowling
Clinical Associate Professor of Obstetrics and Gynecology

Joanna D. Brown
Clinical Associate Professor of Family Medicine

Steven A. Brown
Clinical Associate Professor of Surgery

Maria L. Buckley
Clinical Associate Professor of Psychiatry and Human Behavior

Douglas M. Burtt
Clinical Associate Professor of Medicine

James Butera
Clinical Associate Professor of Medicine

Terrence F. Cahill
Clinical Associate Professor of Obstetrics and Gynecology

Keith Callahan
Clinical Associate Professor of Family Medicine

Christopher P. Campanile
Clinical Associate Professor Emeritus of Family Medicine

Lauralyn B. Cannistra
Clinical Associate Professor of Medicine

David A. Carcieri
Clinical Associate Professor of Obstetrics and Gynecology

Amos Charles
Clinical Associate Professor of Medicine

Wayne C. Clairborne
Clinical Associate Professor Emeritus of Obstetrics and Gynecology

David R. Cloutier
Clinical Associate Professor of Surgery

Bradley J. Collins
Clinical Associate Professor of Medicine

Francis Connor
Clinical Associate Professor of Surgery

Dominic Corrigan
Clinical Associate Professor of Medicine

Geanina O. Costea
Clinical Associate Professor of Psychiatry and Human Behavior

Richard A. Cottiero
Clinical Associate Professor of Medicine

Beth Cronin
Clinical Associate Professor of Obstetrics and Gynecology

Arthur D. Daily
Clinical Associate Professor Emeritus of Dermatology

Stephen W. Davis
Clinical Associate Professor Emeritus of Family Medicine

Thomas D. Denucci
Clinical Associate Professor of Medicine

Allan Mark Deutsch
Clinical Associate Professor Emeritus of Diagnostic Imaging

Christy Dibble
Clinical Associate Professor of Medicine

Elizabeth R. Didie
Clinical Associate Professor of Psychiatry and Human Behavior

Amy Bach Dilello
Clinical Associate Professor of Psychiatry and Human Behavior; Clinical Associate Professor of Medicine

Walter E. Donat
Clinical Associate Professor of Medicine

Joseph L. Dowling
Clinical Associate Professor Emeritus of Surgery

Thomas Drew
Clinical Associate Professor Emeritus of Medicine

Brian E. Duff
Clinical Associate Professor of Surgery

Charles Eil
Clinical Associate Professor of Medicine

Jonathan Elion
Clinical Associate Professor of Medicine

Timothy M. Empkie
Clinical Associate Professor of Family Medicine

Esther Entin
Clinical Associate Professor Emerita of Family Medicine

Yoash R. Enzer
Clinical Associate Professor of Surgery; Clinical Associate Professor of Dermatology

Stephen Falkenberry
Clinical Associate Professor of Obstetrics and Gynecology

David S. Farrell
Clinical Associate Professor Emeritus of Dermatology

Mary Anne Fenton
Clinical Associate Professor of Medicine

Ellen Flynn
Clinical Associate Professor of Psychiatry and Human Behavior; Clinical Associate Professor of Medicine

Glenn G. Fort
Clinical Associate Professor of Medicine

David J. Fortunato
Clinical Associate Professor of Medicine

Elisabeth A. Frazier
Clinical Associate Professor of Psychiatry and Human Behavior

Arthur Frazzano
Clinical Associate Professor Emeritus of Family Medicine

Nancy J. Freeman
Clinical Associate Professor of Medicine

Michael A. Friedman
Clinical Associate Professor of Psychiatry and Human Behavior; Clinical Associate Professor of Neurology

Martin Furman
Clinical Associate Professor of Psychiatry and Human Behavior

Arthur I. Geltzer
Clinical Associate Professor Emeritus of Surgery

Thomas Germano
Clinical Associate Professor of Emergency Medicine

Sharon E. Gibson
Clinical Associate Professor of Surgery

David R. Gifford
Clinical Associate Professor of Medicine; Clinical Associate Professor of Health Services, Policy and Practice

Michael F. Gilson
Clinical Associate Professor of Medicine

Stephen E. Glinick
Clinical Associate Professor of Dermatology

Alan L. Gordon
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior

Norman M. Gordon
Clinical Associate Professor of Neurology

Lisa Gould
Clinical Associate Professor of Medicine

Neil R. Greenspan
Clinical Associate Professor of Medicine

Robert T. Griffith
Clinical Associate Professor of Pediatrics

Nicholas Grumbach
Clinical Associate Professor of Pediatrics; Clinical Associate Professor of Medicine

Nicholas Guerina
Clinical Associate Professor of Pediatrics

Gaurav Gupta
Clinical Associate Professor of Surgery

Vincent M. Harisaran
Clinical Associate Professor of Surgery

Thomas J. Haronian
Clinical Associate Professor of Emergency Medicine

Stephanie Hartselle
Clinical Associate Professor of Psychiatry and Human Behavior

Joseph I. Harwell
Clinical Associate Professor of Medicine; Clinical Associate Professor of Pediatrics

Tawfik F. Hawwa
Clinical Associate Professor of Obstetrics and Gynecology

Arnold Herman
Clinical Associate Professor Emeritus of Surgery

Robert J. Hofmann
Clinical Associate Professor of Surgery

Karyn J. Horowitz
Clinical Associate Professor of Psychiatry and Human Behavior

Barbara N. Jandasek
Clinical Associate Professor of Psychiatry and Human Behavior; Clinical Associate Professor of Pediatrics

Robert H. Janigian
Clinical Associate Professor of Surgery

Ilse M. Jenouri
Clinical Associate Professor of Emergency Medicine

Benjamin Johnson
Clinical Associate Professor of Psychiatry and Human Behavior

Robert G. M. Johnston
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior

Craig Kaufmann
Clinical Associate Professor of Psychiatry and Human Behavior

Edward Keating
Clinical Associate Professor of Medicine; Clinical Associate Professor of Medical Science

Kerri Kim
Clinical Associate Professor of Psychiatry and Human Behavior

Brian Andrew Kimble
Clinical Associate Professor of Medicine; Clinical Associate Professor of Medical Science

Boyd P. King
Clinical Associate Professor Emeritus of Medicine

Michael D. Klein
Clinical Associate Professor of Family Medicine

Robert E. Knisley
Clinical Associate Professor Emeritus of Medicine

Matthew A. Kopp
Clinical Associate Professor of Emergency Medicine

Thomas A. Krahn
Clinical Associate Professor of Medicine

George Kroumpouzos
Clinical Associate Professor of Dermatology

Jennifer F. Lambert
Clinical Associate Professor of Psychiatry and Human Behavior

Elizabeth Lange
Clinical Associate Professor of Pediatrics

Elizabeth A. Laposata
Clinical Associate Professor of Pathology and Laboratory Medicine

Joseph R. Lauro
Clinical Associate Professor of Emergency Medicine

Laura Levine
Clinical Associate Professor of Psychiatry and Human Behavior; Clinical Associate Professor of Medical Science

Rashmi Shetty Licht
Clinical Associate Professor of Surgery

Diana Elliott Lidofsky
Clinical Associate Professor of Psychiatry and Human Behavior

Richard Long
Clinical Associate Professor of Family Medicine

Salvatore J. Loporchio
Clinical Associate Professor of Surgery

Phillip R. Lucas
Clinical Associate Professor of Orthopaedics

Henry G. Magendantz
Clinical Associate Professor Emeritus of Obstetrics and Gynecology

Tariq K. Malik
Clinical Associate Professor of Medicine

Shafiq T. Mamdani
Clinical Associate Professor of Medicine

James Manis
Clinical Associate Professor Emeritus of Medicine

Louis J. Marino
Clinical Associate Professor of Psychiatry and Human Behavior

Charles E. McCoy
Clinical Associate Professor of Medicine

Edward D. Mcgookin
Clinical Associate Professor of Pediatrics; Clinical Associate Professor of Family Medicine

Kevin M. McKay
Clinical Associate Professor of Psychiatry and Human Behavior

Paul N. Mcmillan
Clinical Associate Professor Emeritus of Pathology and Laboratory Medicine

A. Sattar N. Memon
Clinical Associate Professor Emeritus of Medicine

Robert D. Meringolo
Clinical Associate Professor Emeritus of Medicine

Robin June Miller
Clinical Associate Professor of Pediatrics

Jennifer Adelson Mitty
Clinical Associate Professor of Medicine

Carmen V. Monzon
Clinical Associate Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Medicine

Samir G. Moubayed
Clinical Associate Professor Emeritus of Obstetrics and Gynecology

Brian L. Murphy
Clinical Associate Professor of Diagnostic Imaging

James R. Myers
Clinical Associate Professor of Medicine

Jack Nassau
Clinical Associate Professor of Psychiatry and Human Behavior

Guy R. Nicastri
Clinical Associate Professor of Surgery; Clinical Associate Professor of Family Medicine

Judith Nudelman
Clinical Associate Professor Emerita of Family Medicine

Richard K. Ohnmacht
Clinical Associate Professor of Pediatrics

Nicklas B. E. Oldenburg
Clinical Associate Professor of Radiation Oncology

Alvaro Olivares
Clinical Associate Professor of Psychiatry and Human Behavior

Karen Oliver
Clinical Associate Professor of Psychiatry and Human Behavior

Calvin E. Oyer
Clinical Associate Professor Emeritus of Pathology and Laboratory Medicine

Frank Paletta
Clinical Associate Professor of Surgery

Frank A. Pensa
Clinical Associate Professor of Obstetrics and Gynecology

Elliot M. Perlman
Clinical Associate Professor of Surgery

Francis J. Pescosolido
Clinical Associate Professor of Psychiatry and Human Behavior

Francine Pingitore
Clinical Associate Professor of Pediatrics; Clinical Associate Professor of Psychiatry and Human Behavior

Barry A. Plummer
Clinical Associate Professor of Psychiatry and Human Behavior

Roy M. Poses
Clinical Associate Professor of Medicine

Kenneth C. Rickler
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior and Clinical Associate Professor Emeritus of Neurology

Tina Rizack
Clinical Associate Professor of Medicine

Barbara H. Roberts
Clinical Associate Professor Emerita of Medicine

Henry J. Robidoux
Clinical Associate Professor Emeritus of Surgery

Pablo Rodriguez
Clinical Associate Professor Emeritus of Obstetrics and Gynecology

Albert M. Ross
Clinical Associate Professor of Pediatrics

Ara Sadaniantz
Clinical Associate Professor Emeritus of Medicine

Keith Scarfo
Clinical Associate Professor of Neurosurgery

Teresa L. Schraeder
Clinical Associate Professor of Family Medicine

Stuart T. Schwartz
Clinical Associate Professor of Medicine

Reza Shah-Hosseini
Clinical Associate Professor Emeritus of Obstetrics and Gynecology

Lisa B. Shea
Clinical Associate Professor of Psychiatry and Human Behavior

Thomas Sheeran
Clinical Associate Professor of Psychiatry and Human Behavior

Meera Varma Simoes
Clinical Associate Professor of Obstetrics and Gynecology

Latha Sivaprasad
Clinical Associate Professor of Medicine

Lory Snady-Mccoy
Clinical Associate Professor Emerita of Surgery

Clarence H. Soderberg
Clinical Associate Professor Emeritus of Surgery

David A. Solomon
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior

Patricia K. Spencer
Clinical Associate Professor Emerita of Diagnostic Imaging

Laura Stanton
Clinical Associate Professor of Psychiatry and Human Behavior

Pamela Steadman-Wood
Clinical Associate Professor of Psychiatry and Human Behavior

Rochelle S. Strenger
Clinical Associate Professor of Medicine

Francis M. Sullivan
Clinical Associate Professor of Emergency Medicine

Frank W. Sullivan
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior

Ghulam M. Surti
Clinical Associate Professor of Psychiatry and Human Behavior

Lynn A. Sweeney
Clinical Associate Professor of Emergency Medicine

Ross Taliano
Clinical Associate Professor of Pathology and Laboratory Medicine

Charlene A. Tate
Clinical Associate Professor of Neurology

David Robbins Tien
Clinical Associate Professor of Surgery

John F. Todaro
Clinical Associate Professor of Psychiatry and Human Behavior

Philip A. Torgan
Clinical Associate Professor Emeritus of Medicine

Scott A. Triedman
Clinical Associate Professor of Radiation Oncology

Jody Ann Underwood
Clinical Associate Professor of Psychiatry and Human Behavior; Clinical Associate Professor of Medicine

William S. Unger
Clinical Associate Professor of Psychiatry and Human Behavior

Wilson F. Utter
Clinical Associate Professor Emeritus of Pediatrics

Rendueles Villalba
Clinical Associate Professor of Psychiatry and Human Behavior

Matthew D. Vrees
Clinical Associate Professor of Surgery

Eric Ferguson Walsh
Clinical Associate Professor of Orthopaedics

Marilyn Weigner
Clinical Associate Professor of Medicine

Gary G. Wharton
Clinical Associate Professor of Obstetrics and Gynecology

William M. Whelihan
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior

Emily M. White
Clinical Associate Professor of Obstetrics and Gynecology

Janet L. Wilterdink
Clinical Associate Professor of Neurology

Harold A. Woodcome
Clinical Associate Professor Emeritus of Surgery

Margaret S. Wool
Clinical Associate Professor of Family Medicine

Tony C. Wu
Clinical Associate Professor of Medicine

Associate Professor Research

Ruhul Abid
Associate Professor of Surgery (Research)

Michael F. Armey
Associate Professor of Psychiatry and Human Behavior (Research)

Janette Baird
Associate Professor of Emergency Medicine (Research)

David H. Barker
Associate Professor of Psychiatry and Human Behavior (Research)

Kristen Benito
Associate Professor of Psychiatry and Human Behavior (Research)

Paul Bertone
Associate Professor of Medicine (Research)

Shibin Cheng
Associate Professor of Pediatrics (Research)

Li-Jin Chew
Associate Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Bum-Rak Choi
Associate Professor of Medicine (Research)

Lori Daiello
Associate Professor of Neurology (Research); Associate Professor of Health Services, Policy and Practice (Research)

Sean C. L. Deoni
Associate Professor of Diagnostic Imaging (Research); Associate Professor of Pediatrics (Research)

Susan Dickstein
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Pediatrics (Research)

Fizza S. Gillani
Associate Professor of Medicine (Research)

Christopher D. Houck
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Pediatrics (Research)

Hung-Teh Kao
Associate Professor of Psychiatry and Human Behavior (Research)

Jill A. Kreiling
Associate Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Ji Su Li
Associate Professor of Medicine (Research)

Jason Lillis
Associate Professor of Psychiatry and Human Behavior (Research)

Judy Shih-Hwa Liu
Sidney A. Fox and Dorothea Doctors Fox Associate Professor of Ophthalmology and Visual Science, Associate Professor of Molecular Biology, Cell Biology and Biochemistry (Research), Associate Professor of Neurology (Research)

Jason T. Machan
Associate Professor of Orthopaedics (Research); Associate Professor of Surgery (Research)

Lindsay M. Orchowski
Associate Professor of Psychiatry and Human Behavior (Research)

Donna R. Parker
Associate Professor of Family Medicine (Research); Associate Professor of Epidemiology (Research)

Jason Ritt
Associate Professor of Neuroscience (Research)

Jennifer A. Sanders
Associate Professor of Pediatrics (Research); Associate Professor of Pathology and Laboratory Medicine (Research)

Sunil K. Shaw
Associate Professor of Pediatrics (Research)

Nikos Tapinos
Sidney A. Fox and Dorothea Doctors Fox Associate Professor of Ophthalmology, Visual Science, and Neuroscience

Jessica L. Unick
Associate Professor of Psychiatry and Human Behavior (Research)

Anny Usheva
Associate Professor of Surgery (Research)

Alper Uzun
Associate Professor of Pediatrics (Research)

Mascha van 't Wout
Associate Professor of Psychiatry and Human Behavior (Research)

Lei Wei
Associate Professor of Orthopaedics (Research)

Hannah Wei Wu
Associate Professor of Pediatrics (Research)

Hongwei Yao
Associate Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Visiting Associate Professor

Bhrigupati Singh
Visiting Associate Professor of Psychiatry and Human Behavior

Assistant Professor

Adeel Abbasi
Assistant Professor of Medicine

Ahmed Abdelfattah
Robert J. and Nancy D. Carney University Assistant Professor of Brain Science

Liza M. Aguiar
Assistant Professor of Surgery; Assistant Professor of Pediatrics

Saud Alhusaini
Assistant Professor of Neurology

Adam R. Aluisio
Assistant Professor of Emergency Medicine

Trisha Lynn Arnold
Assistant Professor of Psychiatry and Human Behavior

Neishay Ayub
Assistant Professor of Neurology

Adina Badea
Assistant Professor of Pathology and Laboratory Medicine

Debasree Banerjee
Assistant Professor of Medicine

Thomas Barrett
Assistant Professor of Orthopaedics

Luca Bartolini
Assistant Professor of Pediatrics; Assistant Professor of Neurology; Assistant Professor of Neurosurgery

Rachel E. Beard
Assistant Professor of Surgery

Yosef Alfred Berlow
Assistant Professor of Psychiatry and Human Behavior

Lalit Beura
Assistant Professor of Molecular Microbiology and Immunology

Brad D. Blankenhorn
Assistant Professor of Orthopaedics

Leslie Ann Brick
Assistant Professor of Psychiatry and Human Behavior

E. Christine Brousseau
Assistant Professor of Obstetrics and Gynecology

Joshua C. Brown
Assistant Professor of Psychiatry and Human Behavior

Margaret Bublitz
Assistant Professor of Psychiatry and Human Behavior; Assistant Professor of Medicine

Tina Burton
Assistant Professor of Neurology

Benedito Carneiro
Assistant Professor of Medicine

Silvia Chiang
Assistant Professor of Pediatrics

Eric Cohen
Assistant Professor of Orthopaedics

Aristides Cruz
Assistant Professor of Orthopaedics

Katelyn Dannheim
Assistant Professor of Pathology and Laboratory Medicine

Michelle Rose Dawson
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Alexandra M. Deaconescu
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Andre DeSouza
Assistant Professor of Medicine

Theresa M. Desrochers
Rosenberg Family Assistant Professor of Brain Science

Monica Dhakar
Assistant Professor of Neurology

Elizabeth Dibble
Assistant Professor of Diagnostic Imaging

Jonathan Drake
Assistant Professor of Neurology

Kristin H. Dwyer
Assistant Professor of Emergency Medicine

Carsten Eickhoff
Manning Assistant Professor of Medical Science, Assistant Professor of Computer Science

Christopher Elco
Assistant Professor of Pathology and Laboratory Medicine; Assistant Professor of Dermatology

Sleiman El Jamal
Assistant Professor of Neurology

Andrew R. Evans
Assistant Professor of Orthopaedics

Karen Ferreira
Assistant Professor of Pathology and Laboratory Medicine

Hannah Elizabeth Frank
Assistant Professor of Psychiatry and Human Behavior

Jared Fridley
Assistant Professor of Neurosurgery

Andrew Fukuda
Assistant Professor of Psychiatry and Human Behavior

Stephanie Garbern
Assistant Professor of Emergency Medicine

Emily C. Gathright
Assistant Professor of Psychiatry and Human Behavior

Joseph Gil
Assistant Professor of Orthopaedics

Eric Goldstein
Assistant Professor of Neurology

Stephanie Graff
Assistant Professor of Medicine

Ricky Grisson
Assistant Professor of Pathology and Laboratory Medicine

Philip Haines
Assistant Professor of Medicine

Alison Hayward
Assistant Professor of Emergency Medicine

Kelly Holder
Assistant Professor of Medical Science

Sheldon L. Holder
Assistant Professor of Pathology and Laboratory Medicine

Raymond Hsu
Assistant Professor of Orthopaedics

Yu-Wen Alvin Huang
GLF Translational Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Tasnim Imran
Assistant Professor of Medicine

Derek Randall Jenkins
Assistant Professor of Orthopaedics

Narendra Sharma Kala
Assistant Professor of Neurology

Kelsey Anne Kangos
Assistant Professor of Psychiatry and Human Behavior

Naz Karim
Assistant Professor of Emergency Medicine

Tyler R. Kartzinel
Peggy and Henry D. Sharpe Assistant Professor of Environmental Studies, Assistant Professor of Ecology, Evolution, and Organismal Biology

Rachna Kataria
Assistant Professor of Medicine

Paul Koffer
Assistant Professor of Radiation Oncology

Zachary J. Kunicki
Assistant Professor of Psychiatry and Human Behavior

Eren Kuris
Assistant Professor of Orthopaedics

Habibe Kurt
Assistant Professor of Pathology and Laboratory Medicine

Galina Lagos
Assistant Professor of Medicine

Louis R. Lapierre
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Sanghyun Lee
Assistant Professor of Molecular Microbiology and Immunology

Adam Lewkowitz
Assistant Professor of Obstetrics and Gynecology

Justin Li
Assistant Professor of Neurosurgery

George Lisi
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Ali Mahta
Assistant Professor of Neurology

Sara Maleki
Assistant Professor of Pathology and Laboratory Medicine

Shivali Marketkar
Assistant Professor of Pathology and Laboratory Medicine

Lauren Massingham
Assistant Professor of Pediatrics

Thais P. Mather
Assistant Professor of Medical Science; Assistant Professor of Molecular Microbiology and Immunology

Aaron Maxwell
Assistant Professor of Diagnostic Imaging

Sonia Mayoral
Robert J. and Nancy D. Carney Assistant Professor of Brain Science

Rory J. Merritt
Assistant Professor of Emergency Medicine; Assistant Professor of Medical Science

Chelsea J. Miller
Assistant Professor of Radiation Oncology

Sean F. Monaghan
Assistant Professor of Surgery

Julie Monteagudo
Assistant Professor of Surgery

Katelyn Moretti
Assistant Professor of Emergency Medicine

Jessica E. Murphy
Assistant Professor of Medicine

Andrew Musits
Assistant Professor of Emergency Medicine

Matt Nassar
Assistant Professor of Neuroscience

Jacqueline L. Nesi
Assistant Professor of Psychiatry and Human Behavior

Tianyi Niu
Assistant Professor of Neurosurgery

Alyssa Lynne Norris
Assistant Professor of Psychiatry and Human Behavior

Thomas Ollila
Assistant Professor of Medicine

Joseph Pare
Assistant Professor of Emergency Medicine

Justin Parent
Assistant Professor of Psychiatry and Human Behavior

Yash Patel
Assistant Professor of Medicine

John C. Patrick
Assistant Professor of Radiation Oncology

Ari Richard Pelcovits
Assistant Professor of Medicine

Elizabeth Michelle Perelstein
Assistant Professor of Neurology

Frederike Petzschner
Assistant Professor of Psychiatry and Human Behavior

Melissa Pielech
Assistant Professor of Psychiatry and Human Behavior

Jessica S. Plavicki
Manning Assistant Professor of Pathology and Laboratory Medicine

Prarthana Prakash
Assistant Professor of Neurology

Petya Dimitrova Radoeva
Assistant Professor of Psychiatry and Human Behavior

Alexander G. Raufi
Assistant Professor of Medicine

Michael Reznik
Assistant Professor of Neurology; Assistant Professor of Neurosurgery

Stephanie Marie Ruest
Assistant Professor of Emergency Medicine; Assistant Professor of Pediatrics

Laert Rusha
Assistant Professor of Neurosurgery

Melissa L. Russo
Assistant Professor of Obstetrics and Gynecology

Marwan Saad Mohamad Saad
Assistant Professor of Medicine

Jared M. Saletin
Assistant Professor of Psychiatry and Human Behavior

Elizabeth Samuels
Assistant Professor of Epidemiology

Sasmit Sarangi
Assistant Professor of Neurology

Nishant R. Shah
Assistant Professor of Medicine; Assistant Professor of Health Services, Policy and Practice

Stephanie A. Shepard-Umaschi
Assistant Professor of Psychiatry and Human Behavior

Mohamed Sherif
Assistant Professor of Psychiatry and Human Behavior

Natasha A. Sokol
Assistant Professor of Psychiatry and Human Behavior

Daniel Spade
Assistant Professor of Pathology and Laboratory Medicine

Christoph Stretz
Assistant Professor of Neurology

Thanujaa Subramaniam
Assistant Professor of Neurology; Assistant Professor of Neurosurgery

Marybeth Sutter
Assistant Professor of Family Medicine

Konstantina Svokos
Assistant Professor of Neurosurgery

Joanne P. Szczygiel Cunha
Assistant Professor of Medicine

Ramin Tabaddor
Assistant Professor of Orthopaedics; Assistant Professor of Family Medicine

Daniella Teape
Assistant Professor of Pediatrics

Brian Theyel
Assistant Professor of Psychiatry and Human Behavior

Elizabeth Thompson
Assistant Professor of Psychiatry and Human Behavior

Louisa I. Thompson
Assistant Professor of Psychiatry and Human Behavior

Radmehr Torabi
Assistant Professor of Neurosurgery; Assistant Professor of Diagnostic Imaging

Estelle Torbey
Assistant Professor of Medicine

Hayley Treloar Padovano
Assistant Professor of Psychiatry and Human Behavior; Assistant Professor of Behavioral and Social Sciences

Yun-An Tseng
Assistant Professor of Pathology and Laboratory Medicine

Dilber Ece Uzun
Assistant Professor of Pathology and Laboratory Medicine

Ryan Van Patten
Assistant Professor of Psychiatry and Human Behavior

Audra Van Wart
Assistant Professor of Medical Science

Anne Vera Cruz
Assistant Professor of Medical Science

Chrystal Vergara-Lopez
Assistant Professor of Psychiatry and Human Behavior

LG. Ward
Assistant Professor of Psychiatry and Human Behavior

Robert Ward
Assistant Professor of Diagnostic Imaging

Ashley E. Webb
Richard and Edna Salomon Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Jordan C. White
Assistant Professor of Family Medicine; Assistant Professor of Medical Science

Rachel Wightman
Assistant Professor of Emergency Medicine; Adjunct Assistant Professor of Epidemiology

Yun Yang
Assistant Professor of Radiation Oncology

Peng Zhang
Assistant Professor of Medicine

Yang Zhou
Assistant Professor of Molecular Microbiology and Immunology

Clinical Assistant Professor

Christopher A. Abadi
Clinical Assistant Professor of Medicine

Mostafa M. Abousayed
Clinical Assistant Professor of Orthopaedics

Reshma Abraham
Clinical Assistant Professor of Medicine

Nicole Abrunzo
Clinical Assistant Professor of Pediatrics; Clinical Assistant Professor of Medicine

Emily Ackerman
Clinical Assistant Professor of Family Medicine

Benjamin Adler
Clinical Assistant Professor of Family Medicine

Katelyn Affleck
Clinical Assistant Professor of Psychiatry and Human Behavior

David C. Ahern
Clinical Assistant Professor of Psychiatry and Human Behavior

Rimsha Ahmed
Clinical Assistant Professor of Obstetrics and Gynecology

Ali Akhtar
Clinical Assistant Professor of Medicine

Rawya Al-Jabari
Clinical Assistant Professor of Psychiatry and Human Behavior

Motasem A. Al-Yacoub
Clinical Assistant Professor of Neurology

Achilles Alcaraz
Clinical Assistant Professor of Family Medicine

Holly L. Alexandre
Clinical Assistant Professor of Family Medicine

J. Winslow Alford
Clinical Assistant Professor of Family Medicine

Sadaf Ali
Clinical Assistant Professor of Psychiatry and Human Behavior

Scott Allen
Clinical Assistant Professor of Family Medicine

Elizabeth L. Altenhein
Clinical Assistant Professor of Surgery

Kim S. Amin
Clinical Assistant Professor Emeritus of Medicine

Janet Anderson
Clinical Assistant Professor of Psychiatry and Human Behavior

Rex W. Appenfeller
Clinical Assistant Professor of Medicine

Denise M. Arcand
Clinical Assistant Professor of Family Medicine

Michel A. Arcand
Clinical Assistant Professor of Orthopaedics

Stefanie P. Aronow
Clinical Assistant Professor of Pediatrics

Caley Arzamarski
Clinical Assistant Professor of Psychiatry and Human Behavior

Daniel K. Asiedu
Clinical Assistant Professor of Medicine

Sami Assad
Clinical Assistant Professor of Family Medicine

Thomas Michael Atwood
Clinical Assistant Professor of Family Medicine

Jeannine Audet
Clinical Assistant Professor of Pediatrics

Samia Ayub
Clinical Assistant Professor of Medicine

Armin Azar
Clinical Assistant Professor of Psychiatry and Human Behavior

Kevin E. Baill
Clinical Assistant Professor of Psychiatry and Human Behavior

Alexandra Baker
Clinical Assistant Professor of Pediatrics

Caroline Bancroft
Clinical Assistant Professor of Pediatrics

Rajnish Bansal
Clinical Assistant Professor of Medicine

Ross E. Barker
Clinical Assistant Professor of Neurosurgery

Keith Richard Bartolomei
Clinical Assistant Professor of Medicine

Zubair Bashir
Clinical Assistant Professor of Medicine

Orazio J. Basile
Clinical Assistant Professor Emeritus of Dermatology

Bryce Basques
Clinical Assistant Professor of Orthopaedics

Kerri L. Batra
Clinical Assistant Professor of Medicine

Maureen Baur
Clinical Assistant Professor of Medicine

Josef Bautista
Clinical Assistant Professor of Medicine

Solmaz Behtash
Clinical Assistant Professor of Family Medicine

M. David Beitle
Clinical Assistant Professor of Obstetrics and Gynecology

Michael J. Belanger
Clinical Assistant Professor of Orthopaedics

Christina Bellanti
Clinical Assistant Professor of Psychiatry and Human Behavior

Natalia Belosloudtseva
Clinical Assistant Professor of Medicine

Talya Benoff
Clinical Assistant Professor of Psychiatry and Human Behavior

Nathan Barry Beraha
Clinical Assistant Professor of Pediatrics

Geoffrey H. Berg
Clinical Assistant Professor of Medicine

Judith Berger
Clinical Assistant Professor of Psychiatry and Human Behavior

Arnaldo Berges
Clinical Assistant Professor of Psychiatry and Human Behavior

Linda J. Berman
Clinical Assistant Professor of Family Medicine

Rosa Bermudez-Emmanuelli
Clinical Assistant Professor of Obstetrics and Gynecology

Jonathan M. Bertman
Clinical Assistant Professor of Family Medicine

Michael A. Bharier
Clinical Assistant Professor of Dermatology

Shreeyukta Bhattarai
Clinical Assistant Professor of Medicine

David Bica
Clinical Assistant Professor of Orthopaedics; Clinical Assistant Professor of Family Medicine

Zoe Billinkoff
Clinical Assistant Professor of Psychiatry and Human Behavior

Karen Blackmer
Clinical Assistant Professor of Family Medicine

William Robert Blair
Clinical Assistant Professor of Family Medicine

Dawna A. Blake
Clinical Assistant Professor of Medicine

Cynthia Bliss
Clinical Assistant Professor of Pediatrics

Julie Claire Blossom-Hartley
Clinical Assistant Professor of Family Medicine

Sheenagh M. Bodkin
Clinical Assistant Professor of Medicine

Zachary Bohart
Clinical Assistant Professor of Orthopaedics

Kelsey Borner
Clinical Assistant Professor of Psychiatry and Human Behavior

Lisa D. Bowie
Clinical Assistant Professor of Family Medicine

Amy M. Boyer
Clinical Assistant Professor of Medicine

Brittney Starr Boykin
Clinical Assistant Professor of Psychiatry and Human Behavior

Lisa D. Boyle
Clinical Assistant Professor of Obstetrics and Gynecology

Michelle A. Boyle
Clinical Assistant Professor of Family Medicine

Gwynne Bragdon
Clinical Assistant Professor of Orthopaedics

Nancy H. Brand
Clinical Assistant Professor of Psychiatry and Human Behavior

Peter C. Brasch
Clinical Assistant Professor of Surgery

James A. Brcak
Clinical Assistant Professor of Psychiatry and Human Behavior

Paul Breiding
Clinical Assistant Professor of Medicine

Debra L. Brendel
Clinical Assistant Professor of Obstetrics and Gynecology

Patricia Bresky
Clinical Assistant Professor of Psychiatry and Human Behavior

Cheryl Brodsky
Clinical Assistant Professor of Obstetrics and Gynecology

Jimmette Brooks
Clinical Assistant Professor of Family Medicine

Rebecca Brown
Clinical Assistant Professor of Medicine

Karen R. Browning
Clinical Assistant Professor of Obstetrics and Gynecology

Matthew A. Brumbaugh
Clinical Assistant Professor of Medicine

Richard G. Bryan
Clinical Assistant Professor of Surgery

Jennifer Buckley
Clinical Assistant Professor of Family Medicine

Ross Budacki
Clinical Assistant Professor of Orthopaedics

Ahmet Burakgazi
Clinical Assistant Professor of Neurology

Andrew E. Burchard
Clinical Assistant Professor of Surgery

Elizabeth R. Burchard
Clinical Assistant Professor of Obstetrics and Gynecology

Caitlin Burditt
Clinical Assistant Professor of Psychiatry and Human Behavior

Alexandra C. Burns
Clinical Assistant Professor of Psychiatry and Human Behavior

Jeffrey M. Burock
Clinical Assistant Professor of Psychiatry and Human Behavior

Stephen Butler
Clinical Assistant Professor of Family Medicine

Wesley Butt
Clinical Assistant Professor of Family Medicine

Melissa A. Buttaro
Clinical Assistant Professor of Psychiatry and Human Behavior

Deborah Cahill
Clinical Assistant Professor of Anesthesiology

Joseph F. Callaghan
Clinical Assistant Professor Emeritus of Surgery

April Cambre
Clinical Assistant Professor of Psychiatry and Human Behavior

Karen T. Cammuso
Clinical Assistant Professor of Psychiatry and Human Behavior

Christopher A. Campagnari
Clinical Assistant Professor of Family Medicine

Brett Nicholas Cann
Clinical Assistant Professor of Family Medicine

Michael L. Cardenas
Clinical Assistant Professor of Radiation Oncology

Amber Marie Cardoos
Clinical Assistant Professor of Family Medicine

Jeannine Alexandra Casselman
Clinical Assistant Professor of Family Medicine

Brian C. Castelluccio
Clinical Assistant Professor of Psychiatry and Human Behavior

Colleen Cavanaugh
Clinical Assistant Professor of Obstetrics and Gynecology

Timothy Cavanaugh
Clinical Assistant Professor of Family Medicine

Dilek O. Cermik
Clinical Assistant Professor of Obstetrics and Gynecology

Tineke Chan
Clinical Assistant Professor of Surgery

Joyce R. Chang
Clinical Assistant Professor of Medicine

Manchin Chang
Clinical Assistant Professor of Medicine

Simon Chao
Clinical Assistant Professor of Orthopaedics

Tina L. Charest
Clinical Assistant Professor of Family Medicine

Yael Chatav Schonbrun
Clinical Assistant Professor of Psychiatry and Human Behavior

Elizabeth Chattillion
Clinical Assistant Professor of Psychiatry and Human Behavior

Myra Chaudary
Clinical Assistant Professor of Medicine

Iwona Chelminski
Clinical Assistant Professor of Psychiatry and Human Behavior

Andrew Chen
Clinical Assistant Professor of Orthopaedics

William Tze-Wei Chen
Clinical Assistant Professor of Medicine

Kimberley Chiappone
Clinical Assistant Professor of Psychiatry and Human Behavior

Dana Chofay
Clinical Assistant Professor of Medicine; Clinical Assistant Professor of Medical Science

Kue Chung Choi
Clinical Assistant Professor of Obstetrics and Gynecology

Sapna Chowdhry
Clinical Assistant Professor of Medicine

Fredric V. Christian
Clinical Assistant Professor of Medicine

David J. Chronley
Clinical Assistant Professor Emeritus of Pediatrics

Antony Chu
Clinical Assistant Professor of Medicine

Jeffrey D. Clement
Clinical Assistant Professor of Medicine

Ana Cojocaru
Clinical Assistant Professor of Medicine

Mauro Colavita
Clinical Assistant Professor of Obstetrics and Gynecology

Samantha Collum
Clinical Assistant Professor of Psychiatry and Human Behavior

Jeannine M. Connolly
Clinical Assistant Professor of Obstetrics and Gynecology

John M. Conte
Clinical Assistant Professor of Medicine

Molly Anne Cook
Clinical Assistant Professor of Family Medicine

Simon M. Cornelissen
Clinical Assistant Professor of Orthopaedics

Carla Correia
Clinical Assistant Professor of Psychiatry and Human Behavior

Erin Cortes
Clinical Assistant Professor of Pediatrics

Katherine E. Costa
Clinical Assistant Professor of Pediatrics

Ellen M. Costello
Clinical Assistant Professor Emerita of Psychiatry and Human Behavior

Travis Cotton
Clinical Assistant Professor of Surgery

Maria Coutinho
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Pediatrics

Casey Cragin
Clinical Assistant Professor of Psychiatry and Human Behavior

Thomas F. Crain
Clinical Assistant Professor of Medicine

Rebecca J. Crichton
Clinical Assistant Professor of Obstetrics and Gynecology

G. Valentin Cristescu
Clinical Assistant Professor of Medical Science

Laina E. Crowthers
Clinical Assistant Professor of Obstetrics and Gynecology

Marjorie L. Crozier
Clinical Assistant Professor of Psychiatry and Human Behavior

Stephen Cummings
Clinical Assistant Professor of Family Medicine

Wilfredo Curioso
Clinical Assistant Professor of Medicine

Anne Cushing-Brescia
Clinical Assistant Professor of Medicine

Catherine D'Avanzato
Clinical Assistant Professor of Psychiatry and Human Behavior

Yannick D'Silva
Clinical Assistant Professor of Medicine

Zack Dale
Clinical Assistant Professor of Medicine

Nitin S. Damle
Clinical Assistant Professor of Medicine

Rongras Damrongwatanasuk
Clinical Assistant Professor of Medicine

Eileen M. Danaher
Clinical Assistant Professor of Surgery

Anh-Tai Vinh Dang
Clinical Assistant Professor of Obstetrics and Gynecology

Katharine Darland
Clinical Assistant Professor of Medicine

Sarah Davenport
Clinical Assistant Professor of Family Medicine

Kristin C. Davidoff
Clinical Assistant Professor of Psychiatry and Human Behavior

Thamara Davis
Clinical Assistant Professor of Psychiatry and Human Behavior

Laura Deihl
Clinical Assistant Professor of Psychiatry and Human Behavior

Melinda Delaney
Clinical Assistant Professor of Pediatrics

Sara Delaporta
Clinical Assistant Professor of Family Medicine

Paul V. Del Guercio
Clinical Assistant Professor of Family Medicine

Anthony M. DeLuise
Clinical Assistant Professor of Family Medicine

Jane Mackenzie Dennison
Clinical Assistant Professor of Pediatrics

Lisa R. Denny
Clinical Assistant Professor of Family Medicine

Judith Depue
Clinical Professor Emerita of Psychiatry and Human Behavior

Hector Derreza
Clinical Assistant Professor of Medicine

Jaai Deshpande
Clinical Assistant Professor of Family Medicine

Ashok Raj Devkota
Clinical Assistant Professor of Medicine

Saud Dhillon
Clinical Assistant Professor of Medicine

Giulio G. Diamante
Clinical Assistant Professor of Surgery

Alexander Diaz de Villalvilla
Clinical Assistant Professor of Medicine

Margaret A. Dicarlo
Clinical Assistant Professor of Psychiatry and Human Behavior

Dinusha W. Dietrich
Clinical Assistant Professor of Pediatrics

Camia M. Dimock
Clinical Assistant Professor of Family Medicine

Sara DiNardo
Clinical Assistant Professor of Surgery

Jamie L. DiPietro
Clinical Assistant Professor of Psychiatry and Human Behavior

Beata E. Dizoglio
Clinical Assistant Professor of Obstetrics and Gynecology

Joseph DiZoglio
Clinical Assistant Professor of Obstetrics and Gynecology

Linda Doberstein
Clinical Assistant Professor of Pediatrics; Clinical Assistant Professor of Psychiatry and Human Behavior

Robert J. Dobrzynski
Clinical Assistant Professor of Medicine

Kimberly A. Dodd
Clinical Assistant Professor of Medicine

Lisa Ruth Domagalski
Clinical Assistant Professor of Obstetrics and Gynecology

John P. Donahue
Clinical Assistant Professor of Surgery

Deidre L. Donaldson
Clinical Assistant Professor of Family Medicine

Edmund T. Dosremedios
Clinical Assistant Professor of Orthopaedics

Megan Douglas
Clinical Assistant Professor of Medicine

Joseph F. Ducharme
Clinical Assistant Professor of Surgery

George J. Du Pont
Clinical Assistant Professor of Surgery

Robert K. Dyer
Clinical Assistant Professor of Dermatology

Samuel Eaton
Clinical Assistant Professor of Surgery

Robert Eden
Clinical Assistant Professor of Pediatrics

McKinzie Eggers
Clinical Assistant Professor of Family Medicine

Ibrahim Elgabry
Clinical Assistant Professor of Medicine

Lana G. Elhalabi
Clinical Assistant Professor of Psychiatry and Human Behavior

Ahmed Elshazly
Clinical Assistant Professor of Medicine

Martin Elson
Clinical Assistant Professor of Surgery

Christine M. Emmick
Clinical Assistant Professor of Surgery

Shannon Marie Erisman
Clinical Assistant Professor of Psychiatry and Human Behavior

Cynthia Esteban
Clinical Assistant Professor of Pediatrics

Elkin O. Estrada
Clinical Assistant Professor of Medicine

Babak Ettekal
Clinical Assistant Professor of Family Medicine

Matthew Evans
Clinical Assistant Professor of Psychiatry and Human Behavior

Tarra B. Evans
Clinical Assistant Professor of Surgery

Laura E. Fabricant
Clinical Assistant Professor of Psychiatry and Human Behavior

Khawaja Hasham Faisal
Clinical Assistant Professor of Medicine

Frank Faltus
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Jason Fanuele
Clinical Assistant Professor of Orthopaedics

Negar Faramarzi
Clinical Assistant Professor of Medicine

Joseph Farina
Clinical Assistant Professor of Medicine

Joseph Farnam
Clinical Assistant Professor of Medical Science

Elisabeth K. Farnum
Clinical Assistant Professor of Family Medicine

Jeniimarie Febres
Clinical Assistant Professor of Psychiatry and Human Behavior

Seth Feder
Clinical Assistant Professor Emeritus of Dermatology

Michael F. Felder
Clinical Assistant Professor of Family Medicine; Clinical Assistant Professor of Pediatrics

Jason D. Ferreira
Clinical Assistant Professor of Medicine

Marcolino Ferretti
Clinical Assistant Professor of Pediatrics

Sarah J. Fessler
Clinical Assistant Professor of Family Medicine

Kristin Fielding
Clinical Assistant Professor of Family Medicine

John W. Finigan
Clinical Assistant Professor of Pediatrics

Bruce E. Fischer
Clinical Assistant Professor of Medicine

Joshua E. Fischer
Clinical Assistant Professor of Medicine

Libertad Thelma Flores
Clinical Assistant Professor of Obstetrics and Gynecology

Julie Forrest
Clinical Assistant Professor of Psychiatry and Human Behavior

Aryandokht Fotros
Clinical Assistant Professor of Psychiatry and Human Behavior

Gregory A. Fox
Clinical Assistant Professor of Pediatrics

Keith Fragoza
Clinical Assistant Professor of Surgery

Laura L. Frakey
Clinical Assistant Professor of Psychiatry and Human Behavior

Caren L. Francione
Clinical Assistant Professor of Psychiatry and Human Behavior

Heather M. Frank
Clinical Assistant Professor of Psychiatry and Human Behavior

Lisa M. Freda
Clinical Assistant Professor of Psychiatry and Human Behavior

David L. Fried
Clinical Assistant Professor of Medicine

Robert John Fruggiero
Clinical Assistant Professor of Family Medicine

Karla P. Fuentes
Clinical Assistant Professor of Pediatrics

Howard Furlong
Clinical Assistant Professor of Pediatrics

Joseph R. Gaeta
Clinical Assistant Professor Emeritus of Medicine

Robyn Gagnon
Clinical Assistant Professor of Pediatrics

Alan D. Gaines
Clinical Assistant Professor of Pediatrics

Jeffrey Gaines
Clinical Assistant Professor of Medical Science

Janine Galione
Clinical Assistant Professor of Psychiatry and Human Behavior

Harris M. Galkin
Clinical Assistant Professor of Obstetrics and Gynecology

Elizabeth A. Gamble
Clinical Assistant Professor of Obstetrics and Gynecology

Niel Gandhi
Clinical Assistant Professor of Family Medicine

Michele R. Gange
Clinical Assistant Professor of Obstetrics and Gynecology

Prerna Ganjoo
Clinical Assistant Professor of Medicine

Tian Gao
Clinical Assistant Professor of Medicine

Laurie Anne Garabedian
Clinical Assistant Professor of Family Medicine

Michele A. Garazi
Clinical Assistant Professor of Medicine

Veenu Garewal
Clinical Assistant Professor of Medicine

Nikki Garren
Clinical Assistant Professor of Family Medicine

Mason C. Gasper
Clinical Assistant Professor of Neurology

Katherine L. Gassman
Clinical Assistant Professor of Family Medicine

Nathan Gelinas
Clinical Assistant Professor of Medicine

Jonathan Gershon
Clinical Assistant Professor of Psychiatry and Human Behavior

Alfredo Gil
Clinical Assistant Professor of Obstetrics and Gynecology

Malini C. Gillen
Clinical Assistant Professor of Pediatrics

Peter Joseph Gillen
Clinical Assistant Professor of Psychiatry and Human Behavior

Richard G. Gillerman
Clinical Assistant Professor of Medical Science

Caitlin Ann Gillooly
Clinical Assistant Professor of Family Medicine

Ronald M. Gilman
Clinical Assistant Professor of Medicine

Wilfredo Giordano-Perez
Clinical Assistant Professor of Family Medicine

Nicole D. Gleyzer
Clinical Assistant Professor of Family Medicine

Denise Glickman
Clinical Assistant Professor of Medicine

Lauren A. Goddard
Clinical Assistant Professor of Family Medicine

Andrea Gold
Clinical Assistant Professor of Psychiatry and Human Behavior

Arnold C. Goldberg
Clinical Assistant Professor of Family Medicine

William Arthur Golden
Clinical Assistant Professor of Medicine

Amy Goldfarb
Clinical Assistant Professor of Medicine

Debra L. Goldman
Clinical Assistant Professor of Obstetrics and Gynecology

Biju Gopalakrishnan
Clinical Assistant Professor of Neurology

Jessica Gore
Clinical Assistant Professor of Pediatrics

David M. Gottesman
Clinical Assistant Professor of Psychiatry and Human Behavior

Chelsea E. Graham
Clinical Assistant Professor of Family Medicine

Meghan Grant
Clinical Assistant Professor of Family Medicine

Bradford Clayton Gray
Clinical Assistant Professor of Surgery

David M. Greenberg
Clinical Assistant Professor of Surgery

Daniel Greenwald
Clinical Assistant Professor of Medicine

James Edward Greer
Clinical Assistant Professor of Psychiatry and Human Behavior

Iulia C. Grillo
Clinical Assistant Professor of Medicine

Alma M. Guerrero
Clinical Assistant Professor of Medicine

Medhavi Gupta
Clinical Assistant Professor of Medicine

Preeti Gupta
Clinical Assistant Professor of Medical Science

Deborah C. Gutman
Clinical Assistant Professor of Medical Science

Joshua D. Gutman
Clinical Assistant Professor Emeritus of Family Medicine

Ned H. Gutman
Clinical Assistant Professor of Medicine

Thomas E. Guttmacher
Clinical Assistant Professor of Family Medicine

Ruth Guyer
Clinical Assistant Professor of Medicine

Charles D. Hackett
Clinical Assistant Professor of Family Medicine

Neima Hadi
Clinical Assistant Professor of Medicine

Aditi V. Hajirnis
Clinical Assistant Professor of Psychiatry and Human Behavior

Mehtap Haktanir Abul
Clinical Assistant Professor of Pediatrics

Amy R. Halt
Clinical Assistant Professor of Psychiatry and Human Behavior

Geoffrey R. Hamilton
Clinical Assistant Professor of Medicine

Teresita Hamilton
Clinical Assistant Professor of Family Medicine

Sajeev Handa
Clinical Assistant Professor of Medicine; Clinical Assistant Professor of Neurology

Cynthia M. Hanna
Clinical Assistant Professor of Obstetrics and Gynecology

Fang Yin Ingrid Hao
Clinical Assistant Professor of Medicine

Emily C. Harrison
Clinical Assistant Professor of Family Medicine

Pamela A. Harrop
Clinical Assistant Professor of Medicine

Katheleen Hawes
Clinical Assistant Professor of Pediatrics

Eugene H. Healey
Clinical Assistant Professor Emeritus of Surgery

Robert C. Heffron
Clinical Assistant Professor of Family Medicine

Jennifer Hellmuth
Clinical Assistant Professor of Psychiatry and Human Behavior

Ryan P. Heney
Clinical Assistant Professor of Family Medicine

Melvin Hershkowitz
Clinical Assistant Professor Emeritus of Medicine

Suzanne R. Herzberg
Clinical Assistant Professor of Family Medicine

Nadine Hewamudalige
Clinical Assistant Professor of Family Medicine

Ellen B. Hight
Clinical Assistant Professor of Family Medicine

Lynn T. Ho
Clinical Assistant Professor of Family Medicine

Ameya Hodarkar
Clinical Assistant Professor of Medicine

Erin Hogan
Clinical Assistant Professor of Medicine

Leigh Anne Hohlstein
Clinical Assistant Professor of Psychiatry and Human Behavior

Jamie Hollenbeck
Clinical Assistant Professor of Psychiatry and Human Behavior

Peter Amberg Hollmann
Clinical Assistant Professor of Family Medicine

Jennifer J. Hosmer
Clinical Assistant Professor of Obstetrics and Gynecology

Suzanne M. Houston
Clinical Assistant Professor of Family Medicine

Jessica Huang
Clinical Assistant Professor of Family Medicine

Kirsten L. Hull
Clinical Assistant Professor of Psychiatry and Human Behavior

Heather L. Hunter
Clinical Assistant Professor of Psychiatry and Human Behavior

Jeffrey Hunter
Clinical Assistant Professor of Obstetrics and Gynecology

Brett Hurteau
Clinical Assistant Professor of Family Medicine

Alice Yung-Mei Hwang
Clinical Assistant Professor of Family Medicine

Adriana Hyams
Clinical Assistant Professor of Psychiatry and Human Behavior

Michael Iadevaia
Clinical Assistant Professor of Family Medicine

Harry Iannotti
Clinical Assistant Professor of Surgery

Gabriel Ibarra
Clinical Assistant Professor of Medicine

Farhat Ibrahim
Clinical Assistant Professor of Medicine

Sadia Iftikhar
Clinical Assistant Professor of Medicine

Omosede O. Ighile
Clinical Assistant Professor of Medicine

James Ingraham
Clinical Assistant Professor of Psychiatry and Human Behavior

Arshad Iqbal
Clinical Assistant Professor of Neurology

John Isaac
Clinical Assistant Professor of Medical Science

Barbara W. Jablow
Clinical Assistant Professor of Family Medicine

Angela Jacavone
Clinical Assistant Professor of Family Medicine

Bryan S. Jay
Clinical Assistant Professor of Diagnostic Imaging

Nathaniel J. Jellinek
Clinical Assistant Professor of Dermatology

Sharon Mullane Jezard
Clinical Assistant Professor of Family Medicine

Carolina Jimenez-Madiedo
Clinical Assistant Professor of Psychiatry and Human Behavior

Christine Johnson
Clinical Assistant Professor of Family Medicine

Eileen G. Johnson
Clinical Assistant Professor of Psychiatry and Human Behavior

Jessica N. Johnson
Clinical Assistant Professor of Family Medicine

Ormanda Johnson
Clinical Assistant Professor of Family Medicine

Donald R. Joyce
Clinical Assistant Professor of Radiation Oncology

Annmarie Jurczak
Clinical Assistant Professor of Obstetrics and Gynecology

Amr Kader
Clinical Assistant Professor of Obstetrics and Gynecology

David Kadmon
Clinical Assistant Professor of Family Medicine

Sewell I. Kahn
Clinical Assistant Professor Emeritus of Medicine

Aaron William Kaiser
Clinical Assistant Professor of Psychiatry and Human Behavior

Brett D. Kalmowitz
Clinical Assistant Professor of Medicine

Elena Kamenetsky
Clinical Assistant Professor Emerita of Neurology

Barbara T. Kao
Clinical Assistant Professor of Pediatrics

Peter Karczmar
Clinical Assistant Professor of Medicine

Kabita Karki
Clinical Assistant Professor of Medicine

Donald G. Kaufman
Clinical Assistant Professor Emeritus of Medicine

W. Scott Keigwin
Clinical Assistant Professor of Family Medicine

Amanda Aguiar Kelvey
Clinical Assistant Professor of Family Medicine

Mustapha Kemal
Clinical Assistant Professor of Neurology

Christine A. Kennedy
Clinical Assistant Professor of Family Medicine

Martin J. Kerzer
Clinical Assistant Professor of Family Medicine

Mohammad Mudassar Khan
Clinical Assistant Professor of Medicine

Pervez A. Khatib
Clinical Assistant Professor of Family Medicine

Razib Khaund
Clinical Assistant Professor of Medicine

Zia Kidwai
Clinical Assistant Professor of Medicine

Malcolm M. Kirk
Clinical Assistant Professor of Medicine

Christopher Ryan Klaus
Clinical Assistant Professor of Family Medicine

Margaret G. Klitzke
Clinical Assistant Professor Emerita of Psychiatry and Human Behavior

Eva Klonowski
Clinical Assistant Professor of Family Medicine

Kristin S. Knapp-Ines
Clinical Assistant Professor of Psychiatry and Human Behavior

Tolga N. Kokturk
Clinical Assistant Professor of Obstetrics and Gynecology

Jonathan Kole
Clinical Assistant Professor of Psychiatry and Human Behavior

Uma Kolli
Clinical Assistant Professor of Family Medicine

Liliya Koyfman
Clinical Assistant Professor of Psychiatry and Human Behavior

Altug Koymen
Clinical Assistant Professor of Obstetrics and Gynecology

Naomi R. Kramer
Clinical Assistant Professor of Medicine

Alicja U. Kreczko
Clinical Assistant Professor of Obstetrics and Gynecology

Tushar Kulkarni
Clinical Assistant Professor of Medicine

Erin E. Kunkel
Clinical Assistant Professor of Obstetrics and Gynecology

Audrey Kupchan
Clinical Assistant Professor of Medicine

Jayne A. Kurkjian-Siegel
Clinical Assistant Professor of Psychiatry and Human Behavior

Daniel K. Kwan
Clinical Assistant Professor of Surgery

Abbey Ellen Lamb
Clinical Assistant Professor of Pediatrics

Robert H. Lambe
Clinical Assistant Professor of Family Medicine

Jon K. Lambrecht
Clinical Assistant Professor of Medicine

Chad E. Lamendola
Clinical Assistant Professor of Family Medicine

Steven Lampert
Clinical Assistant Professor of Medicine

Guy Lancellotti
Clinical Assistant Professor of Medical Science

Gina La Prova
Clinical Assistant Professor of Family Medicine

Paul R. Larson
Clinical Assistant Professor of Family Medicine

Usman Lashari
Clinical Assistant Professor of Family Medicine

Nancy Lasson
Clinical Assistant Professor of Medicine

Ingrid Lauer-Arnold
Clinical Assistant Professor of Psychiatry and Human Behavior

Caitlin Lawrence
Clinical Assistant Professor of Psychiatry and Human Behavior

Benjamin Lederer
Clinical Assistant Professor of Psychiatry and Human Behavior

Ruby Lee
Clinical Assistant Professor of Psychiatry and Human Behavior

Sung-Hee Rhim Lee
Clinical Assistant Professor of Obstetrics and Gynecology

Woo Jin Lee
Clinical Assistant Professor of Orthopaedics

Michelle J. Lefebvre
Clinical Assistant Professor of Pediatrics

Kendra Levi
Clinical Assistant Professor of Family Medicine

Jonathan A. Leviss
Clinical Assistant Professor of Medicine

Jennifer S. Levy
Clinical Assistant Professor of Family Medicine

Sarah Simone Levy
Clinical Assistant Professor of Surgery

Victoria Leytin
Clinical Assistant Professor of Emergency Medicine

Tiffany Libby
Clinical Assistant Professor of Dermatology

Jonah M. Licht
Clinical Assistant Professor of Medicine

Warren E. Licht
Clinical Assistant Professor of Medicine

David P. Lichtenstein
Clinical Assistant Professor of Psychiatry and Human Behavior

Sheldon Lidofsky
Clinical Assistant Professor of Medicine

Stacey P. Lievense
Clinical Assistant Professor of Obstetrics and Gynecology

Joseph T. Lifrak
Clinical Assistant Professor of Family Medicine

Swee Yang Lim
Clinical Assistant Professor of Neurology

James Lippincott
Clinical Assistant Professor of Family Medicine

Marcia Liss
Clinical Assistant Professor of Psychiatry and Human Behavior

Jerome J. Liu
Clinical Assistant Professor of Psychiatry and Human Behavior

Roald J. Llado
Clinical Assistant Professor of Family Medicine

Douglas Moore Long
Clinical Assistant Professor of Psychiatry and Human Behavior

Gabriela Lopez
Clinical Assistant Professor of Medical Science

Yi Lu
Clinical Assistant Professor of Medicine

Melissa Ludwig
Clinical Assistant Professor of Psychiatry and Human Behavior

Vanessa Lund
Clinical Assistant Professor of Orthopaedics

Krista Lussier
Clinical Assistant Professor of Psychiatry and Human Behavior

Christopher Luttmann
Clinical Assistant Professor of Medicine

Mary E. Lyster
Clinical Assistant Professor of Family Medicine

D. Audoen Maddock
Clinical Assistant Professor of Obstetrics and Gynecology

James O. Maher
Clinical Assistant Professor of Orthopaedics

Kelby G. Maher
Clinical Assistant Professor of Medical Science

Matthew Michael Malek
Clinical Assistant Professor of Family Medicine

Steven A. Mallozzi
Clinical Assistant Professor of Medicine

Scott Joseph Mandel
Clinical Assistant Professor of Orthopaedics

Nayana Manjunath
Clinical Assistant Professor of Obstetrics and Gynecology

Elena Manning
Clinical Assistant Professor of Orthopaedics

Holly Manning
Clinical Assistant Professor of Psychiatry and Human Behavior

Jeffrey Manning
Clinical Assistant Professor of Family Medicine

Jessica C. Manyan
Clinical Assistant Professor of Family Medicine

Ginger L. Manzo
Clinical Assistant Professor of Psychiatry and Human Behavior

Pierre R. Manzo
Clinical Assistant Professor of Family Medicine

John R. Marcaccio
Clinical Assistant Professor Emeritus of Surgery

Stephanie Marchand
Clinical Assistant Professor of Pediatrics

Peter S. Margolis
Clinical Assistant Professor of Medicine

Jessica M. Marrero
Clinical Assistant Professor of Family Medicine

Carla M. Martin
Clinical Assistant Professor of Medical Science

Saul A. Martin
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Christopher M. Matkovic
Clinical Assistant Professor of Psychiatry and Human Behavior

Jennifer L. Maude
Clinical Assistant Professor of Medicine

Laura A. Mayer
Clinical Assistant Professor of Psychiatry and Human Behavior

Elizabeth Mayhall
Clinical Assistant Professor of Obstetrics and Gynecology

Shardale L. McAfee
Clinical Assistant Professor of Family Medicine

Thomas McCauley
Clinical Assistant Professor of Surgery

Katherine Donovan McCleary
Clinical Assistant Professor of Family Medicine

David W. McConville
Clinical Assistant Professor of Psychiatry and Human Behavior

Paula McEvoy
Clinical Assistant Professor of Pediatrics

Heather A. McGee
Clinical Assistant Professor of Medical Science

John McGonigle
Clinical Assistant Professor of Family Medicine

Andrew P. Mcgrath
Clinical Assistant Professor of Pediatrics

Rebecca Mclean
Clinical Assistant Professor of Psychiatry and Human Behavior

Megan D. McMahon
Clinical Assistant Professor of Obstetrics and Gynecology

Michael Mcquiggan
Clinical Assistant Professor of Psychiatry and Human Behavior

Anthony P. Mechrefe
Clinical Assistant Professor of Family Medicine

Carroll A. Medeiros
Clinical Assistant Professor of Obstetrics and Gynecology

Lucille Mehring
Clinical Assistant Professor of Medical Science

Christina M. Mele
Clinical Assistant Professor of Psychiatry and Human Behavior

Lisa M. Menard-Manlove
Clinical Assistant Professor of Family Medicine

Manuela M. Mendes
Clinical Assistant Professor of Family Medicine

Roger Mennillo
Clinical Assistant Professor of Medical Science

Mridula Menon
Clinical Assistant Professor of Medicine

Molly Zipporah Meth
Clinical Assistant Professor of Psychiatry and Human Behavior

Diane E. Minasian
Clinical Assistant Professor of Family Medicine

Jane Miniutti
Clinical Assistant Professor of Family Medicine

Ashish Misri
Clinical Assistant Professor of Medicine

Maya Mistry
Clinical Assistant Professor of Family Medicine

Cristina L. Mitchell
Clinical Assistant Professor of Family Medicine

Joyce E. Monac
Clinical Assistant Professor Emerita of Pediatrics

Keith O. Monchik
Clinical Assistant Professor of Family Medicine

Michael Monsour
Clinical Assistant Professor of Medicine

Sydney Moon
Clinical Assistant Professor of Psychiatry and Human Behavior

Cesar Mora Jaramillo
Clinical Assistant Professor of Family Medicine

Theresa Morgan
Clinical Assistant Professor of Psychiatry and Human Behavior

Richard V. Morgera
Clinical Assistant Professor of Family Medicine

William J. Morocco
Clinical Assistant Professor of Pediatrics

Gillian A. Morris
Clinical Assistant Professor of Family Medicine

Sarah Herrick Morris
Clinical Assistant Professor of Psychiatry and Human Behavior

Ryann A. Morrison
Clinical Assistant Professor of Psychiatry and Human Behavior

John Rollin Morton
Clinical Assistant Professor of Family Medicine

David Moss
Clinical Assistant Professor of Orthopaedics

Morgan Kutzman Motia
Clinical Assistant Professor of Family Medicine

Susannah L. Mozley
Clinical Assistant Professor of Psychiatry and Human Behavior

Jon A. Mukand
Clinical Assistant Professor of Orthopaedics

Donald R. Murphy
Clinical Assistant Professor of Family Medicine

Emily E. Murphy
Clinical Assistant Professor of Psychiatry and Human Behavior

John A. Murphy
Clinical Assistant Professor Emeritus of Obstetrics and Gynecology

Melissa M. Murphy
Clinical Assistant Professor of Medical Science

Anne L. Murray
Clinical Assistant Professor of Obstetrics and Gynecology

David Mysels
Clinical Assistant Professor of Psychiatry and Human Behavior

Jennifer Judd Nadeau
Clinical Assistant Professor of Psychiatry and Human Behavior

Fatima Najeeb
Clinical Assistant Professor of Medicine

Namrata Nandakumar
Clinical Assistant Professor of Surgery

Christopher Shawn Nasin
Clinical Assistant Professor of Family Medicine

Umair Nazir
Clinical Assistant Professor of Medicine

Lina R. Nemchenok
Clinical Assistant Professor of Medicine

Dawn K. Nero
Clinical Assistant Professor of Psychiatry and Human Behavior

Laura S. Nevel
Clinical Assistant Professor of Obstetrics and Gynecology

Christopher Newton
Clinical Assistant Professor of Surgery

Eric B. Newton
Clinical Assistant Professor of Medicine

Karen Y. Ng
Clinical Assistant Professor of Family Medicine

Huy N. Nguyen
Clinical Assistant Professor of Orthopaedics

Mindy Nguyen
Clinical Assistant Professor of Family Medicine

Claire Nicogossian
Clinical Assistant Professor of Psychiatry and Human Behavior

John D. Nisbet
Clinical Assistant Professor of Obstetrics and Gynecology

Arthur W. Noel
Clinical Assistant Professor of Diagnostic Imaging

Courtney Noonan
Clinical Assistant Professor of Obstetrics and Gynecology

Andrew Nowak
Clinical Assistant Professor of Medicine

Lisa Noyes-Duguay
Clinical Assistant Professor of Medicine

Kyle Nuland
Clinical Assistant Professor of Orthopaedics

Christine Nunes
Clinical Assistant Professor of Psychiatry and Human Behavior

Kathleen A. O'Heelan
Clinical Assistant Professor of Family Medicine

Olga Obraztsova
Clinical Assistant Professor of Psychiatry and Human Behavior

Melissa Alejandra Ochoa-Trochez
Clinical Assistant Professor of Family Medicine

Patricia M. Ogera
Clinical Assistant Professor of Pathology and Laboratory Medicine

Rachel Ojserkis
Clinical Assistant Professor of Psychiatry and Human Behavior

Warren Ong
Clinical Assistant Professor of Psychiatry and Human Behavior

Nwamaka Onwugbenu
Clinical Assistant Professor of Obstetrics and Gynecology

Roberto Ortiz
Clinical Assistant Professor of Medicine; Clinical Assistant Professor of Pathology and Laboratory Medicine

Pranav Oza
Clinical Assistant Professor of Medicine

Margaret Paccione-Dyszlewski
Clinical Assistant Professor of Psychiatry and Human Behavior

Cristina A. Pacheco
Clinical Assistant Professor of Family Medicine

Rafael E. Padilla
Clinical Assistant Professor of Anesthesiology

Anjali Palav
Clinical Assistant Professor of Psychiatry and Human Behavior

Daniella Palermo
Clinical Assistant Professor of Psychiatry and Human Behavior

Melissa Lorien Palma
Clinical Assistant Professor of Family Medicine

Altin Pani
Clinical Assistant Professor of Surgery

Shrema Parikh
Clinical Assistant Professor of Pediatrics

Andrea Parker
Clinical Assistant Professor of Surgery

Michelle Parker
Clinical Assistant Professor of Psychiatry and Human Behavior

Robert K. Parker
Clinical Assistant Professor of Surgery

Katherine Partridge
Clinical Assistant Professor of Psychiatry and Human Behavior

George Pasquarello
Clinical Assistant Professor of Orthopaedics

Christina Pastorello
Clinical Assistant Professor of Psychiatry and Human Behavior

Bonny Patel
Clinical Assistant Professor of Psychiatry and Human Behavior

Samir M. Patel
Clinical Assistant Professor of Psychiatry and Human Behavior

Sandhya Patel
Clinical Assistant Professor of Psychiatry and Human Behavior

Amrit Pathak
Clinical Assistant Professor of Medicine

Alfred A. Paul
Clinical Assistant Professor of Surgery

Mark Raymond Paulos
Clinical Assistant Professor of Medicine

Raymond Pavlovich
Clinical Assistant Professor of Orthopaedics

Gillian E. Pearis
Clinical Assistant Professor of Psychiatry and Human Behavior

David W. Pearson
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Edward A. Pensa
Clinical Assistant Professor of Medicine

Emily M. Pepyne
Clinical Assistant Professor of Orthopaedics

Vincent Pera
Clinical Assistant Professor of Medicine

Pranith Perera
Clinical Assistant Professor of Medicine

Marc Jonathan Perkel
Clinical Assistant Professor of Psychiatry and Human Behavior

Adrienne J. Perry
Clinical Assistant Professor of Obstetrics and Gynecology

Matthew Beard Perry
Clinical Assistant Professor of Family Medicine

Tammi-Marie K. Phillip
Clinical Assistant Professor of Psychiatry and Human Behavior

Bruce A. Phillips
Clinical Assistant Professor of Family Medicine

Justyna B. Piasecka
Clinical Assistant Professor of Psychiatry and Human Behavior

Sarah Catherine Pillemer
Clinical Assistant Professor of Psychiatry and Human Behavior

Martha A. Pizzarello
Clinical Assistant Professor of Obstetrics and Gynecology

Peter A. Pizzarello
Clinical Assistant Professor of Family Medicine

Matthew J. Plante
Clinical Assistant Professor of Family Medicine

Gabriel J. Pleasants
Clinical Assistant Professor of Family Medicine

Gabriel J. Plourde
Clinical Assistant Professor of Family Medicine

Alan D. Podis
Clinical Assistant Professor Emeritus of Surgery

Jose R. Polanco
Clinical Assistant Professor of Medicine

Christian Pope
Clinical Assistant Professor of Obstetrics and Gynecology

Michael T. Poshkus
Clinical Assistant Professor of Medicine

Ornella Judith Potter
Clinical Assistant Professor of Psychiatry and Human Behavior

Marylin M. Powers
Clinical Assistant Professor of Obstetrics and Gynecology

Glenn Stephen Prescod
Clinical Assistant Professor of Surgery; Clinical Assistant Professor of Family Medicine

Ann Back Price
Clinical Assistant Professor of Psychiatry and Human Behavior

Sherri Provencal
Clinical Assistant Professor of Psychiatry and Human Behavior

Albert J. Puerini
Clinical Assistant Professor Emeritus of Family Medicine

Margaret M. Pyle
Clinical Assistant Professor of Pediatrics

C. Brandon Qualls
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

David G. Quigley
Clinical Assistant Professor Emeritus of Orthopaedics

Kathy Radie-Keane
Clinical Assistant Professor of Radiation Oncology

Dale F. Radka
Clinical Assistant Professor of Psychiatry and Human Behavior

Keith W. L. Rafal
Clinical Assistant Professor of Family Medicine

Jason Rafferty
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Pediatrics

Anchela Rajeshwaran
Clinical Assistant Professor of Medicine

Verity Ramirez
Clinical Assistant Professor of Medicine

Rebecca W. Randall
Clinical Assistant Professor of Obstetrics and Gynecology

Paras Ranjitkar
Clinical Assistant Professor of Medicine

Rajendrasinh Rathod
Clinical Assistant Professor of Obstetrics and Gynecology

Roger D. Raymond
Clinical Assistant Professor of Medicine

James R. Rayner
Clinical Assistant Professor of Emergency Medicine

Lauren Reeves
Clinical Assistant Professor of Psychiatry and Human Behavior

Wendy P. Regan
Clinical Assistant Professor of Family Medicine

Richard A. Regnante
Clinical Assistant Professor of Medicine

Jared Reichenberg
Clinical Assistant Professor of Psychiatry and Human Behavior

Jose Rengifo
Clinical Assistant Professor of Psychiatry and Human Behavior

Staci Resnick
Clinical Assistant Professor of Medical Science

Katharine Conrad Reynolds
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Pediatrics

Anthony R. Ricci
Clinical Assistant Professor of Medicine

Donald Rice
Clinical Assistant Professor of Medicine

Mark A. Ringiewicz
Clinical Assistant Professor of Family Medicine

Robert Risica
Clinical Assistant Professor of Surgery

Dannie C. Ritchie
Clinical Assistant Professor of Family Medicine

Jorge Rivera
Clinical Assistant Professor of Surgery

Sophia Rizk
Clinical Assistant Professor of Medicine

Debra L. Roberts
Clinical Assistant Professor of Family Medicine

Alexander P. Robertson
Clinical Assistant Professor of Orthopaedics

James F. Robertson
Clinical Assistant Professor of Medicine

Courtney Robidoux
Clinical Assistant Professor of Pediatrics

Lisa Rocchio
Clinical Assistant Professor of Psychiatry and Human Behavior

Marina Rodriguez
Clinical Assistant Professor of Medicine

Andrew J. Rogers
Clinical Assistant Professor of Orthopaedics

Katrina Jane Roi
Clinical Assistant Professor of Family Medicine

Jessica Denise Rome
Clinical Assistant Professor of Medical Science

Jennifer A. Rosenbaum
Clinical Assistant Professor of Family Medicine

Mindy S. Rosenbloom
Clinical Assistant Professor of Psychiatry and Human Behavior

Michael G. Ross
Clinical Assistant Professor of Medicine

Toni M. Roth
Clinical Assistant Professor of Radiation Oncology

Emily S. Rowland
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Medical Science

Lowell J. Rubin
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Barbara M. Ruf
Clinical Assistant Professor of Psychiatry and Human Behavior

Richard J. Ruggieri
Clinical Assistant Professor of Medicine

Charles M. Ruhl
Clinical Assistant Professor of Surgery

Renee B. Rulin
Clinical Assistant Professor of Family Medicine

Lorna B. Russell
Clinical Assistant Professor of Medicine

J. Mark Ryan
Clinical Assistant Professor of Medicine

Inna Ryvkin
Clinical Assistant Professor of Family Medicine

Andrew Saal
Clinical Assistant Professor of Family Medicine

Karen Lee Saal
Clinical Assistant Professor of Medical Science

Daniel C. Sacchetti
Clinical Assistant Professor of Neurology

Henry T. Sachs
Clinical Assistant Professor of Psychiatry and Human Behavior

Gregory Sadovnikoff
Clinical Assistant Professor of Family Medicine

Jessica R. Salak
Clinical Assistant Professor of Obstetrics and Gynecology

Gisele I. Saliba
Clinical Assistant Professor of Medicine

Matthew T. Salisbury
Clinical Assistant Professor of Family Medicine

Philip Salko
Clinical Assistant Professor of Family Medicine; Clinical Assistant Professor of Orthopaedics

Jason Salter
Clinical Assistant Professor of Family Medicine

Fadi Samaan
Clinical Assistant Professor of Medicine

Sundaresan T. Sambandam
Clinical Assistant Professor of Medicine

Mary Ann Santos
Clinical Assistant Professor of Pediatrics

Kelley Sanzen
Clinical Assistant Professor of Medicine

Paul E. Sapir
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Brian Sard
Clinical Assistant Professor of Pediatrics

Brian Savoie
Clinical Assistant Professor of Surgery

Syeda M. Sayeed
Clinical Assistant Professor of Medicine

Thomas J. Scaramella
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Monica J. Schaberg
Clinical Assistant Professor Emerita of Pediatrics

Jay S. Schachne
Clinical Assistant Professor of Medicine

Sarah B. Schmidhofer
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Medical Science

Phillip J. Schmitt
Clinical Assistant Professor of Medical Science

Franklin Schneider
Clinical Assistant Professor of Medicine

David Schreiber
Clinical Assistant Professor of Medicine

Howard E. Schulman
Clinical Assistant Professor of Medicine

Jack L. Schwartzwald
Clinical Assistant Professor of Medicine

Robert H. Schwengel
Clinical Assistant Professor of Medicine

Stephen M. Scott
Clinical Assistant Professor of Medicine

Christina Scully
Clinical Assistant Professor of Psychiatry and Human Behavior

Gregg J. Selke
Clinical Assistant Professor of Psychiatry and Human Behavior

Sonia Seng
Clinical Assistant Professor of Medicine

Kurush Setna
Clinical Assistant Professor of Medicine

Robert J. Settipane
Clinical Assistant Professor of Medical Science

Catherine E. Shafts
Clinical Assistant Professor of Family Medicine

Isha Shah
Clinical Assistant Professor of Medicine

Ahmed Shahab
Clinical Assistant Professor of Medicine

Bradley A. Shapiro
Clinical Assistant Professor of Diagnostic Imaging

Jane R. Sharp
Clinical Assistant Professor of Obstetrics and Gynecology

Meghan Sharp
Clinical Assistant Professor of Psychiatry and Human Behavior

Edward P. Sheehan
Clinical Assistant Professor of Psychiatry and Human Behavior

Burton H. Shen
Clinical Assistant Professor of Medicine

Yiyun Shi
Clinical Assistant Professor of Medicine

Cullen Shipman
Clinical Assistant Professor of Family Medicine

Priscilla Shube
Clinical Assistant Professor of Family Medicine

Santina L. Siena
Clinical Assistant Professor Emerita of Obstetrics and Gynecology

Jacquelyn M. Silva
Clinical Assistant Professor of Pediatrics

Michael A. Silver
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Louis R. Simeone
Clinical Assistant Professor of Orthopaedics

Rachel Ann Simmons
Clinical Assistant Professor of Psychiatry and Human Behavior

Mitchel Sklar
Clinical Assistant Professor of Medicine

Caroline Skudlarek-Prete
Clinical Assistant Professor of Obstetrics and Gynecology

John G. Slattery
Clinical Assistant Professor of Family Medicine

Karen Smigel
Clinical Assistant Professor of Family Medicine

Matthew Joseph Smith
Clinical Assistant Professor of Orthopaedics; Clinical Assistant Professor of Family Medicine

Richard M. Smith
Clinical Assistant Professor of Psychiatry and Human Behavior

Jon S. Solis
Clinical Assistant Professor of Dermatology

John J. Solomon
Clinical Assistant Professor of Family Medicine

Sukanya Somasundar
Clinical Assistant Professor of Family Medicine

Nitesh Sood
Clinical Assistant Professor of Medical Science

Roy Souaid
Clinical Assistant Professor of Family Medicine

Marta S. Sowa
Clinical Assistant Professor of Pediatrics

Jeremy Spector
Clinical Assistant Professor of Medicine

Joel S. Spellun
Clinical Assistant Professor of Medicine

Megan B. Spencer
Clinical Assistant Professor of Psychiatry and Human Behavior

Kenneth F. Sperber
Clinical Assistant Professor of Family Medicine

Stuart Spitalnic
Clinical Assistant Professor of Emergency Medicine

Emily B. Spurrell
Clinical Assistant Professor of Psychiatry and Human Behavior

Michael Staebler
Clinical Assistant Professor of Orthopaedics

Michael L. Stanchina
Clinical Assistant Professor of Medicine

Melanie Stark
Clinical Assistant Professor of Psychiatry and Human Behavior

Brett S. Stecker
Clinical Assistant Professor of Family Medicine

Kim M. Stein
Clinical Assistant Professor of Family Medicine

Steven H. Stein
Clinical Assistant Professor of Family Medicine

Gregory J. Steinmetz
Clinical Assistant Professor of Family Medicine

Charles Leigh Stengel
Clinical Assistant Professor of Emergency Medicine

Jodi Stephenson
Clinical Assistant Professor of Psychiatry and Human Behavior

Gregory Stiener
Clinical Assistant Professor of Psychiatry and Human Behavior

Kristyn N. Storey
Clinical Assistant Professor of Psychiatry and Human Behavior

John A. Stoukides
Clinical Assistant Professor of Medicine

Michele Style
Clinical Assistant Professor of Obstetrics and Gynecology

Faizul Suhail
Clinical Assistant Professor of Medicine

Elizabeth Loring Sullivan
Clinical Assistant Professor of Obstetrics and Gynecology

James C. Sullivan
Clinical Assistant Professor of Medicine

Jamie Sullivan
Clinical Assistant Professor of Family Medicine

John James Sullivan
Clinical Assistant Professor of Medicine

Vrishali Suman
Clinical Assistant Professor of Family Medicine

Margaret A. Sun
Clinical Assistant Professor of Family Medicine

Jeanne W. Swen
Clinical Assistant Professor of Family Medicine

Jeffrey B. Syme
Clinical Assistant Professor of Family Medicine

Charu Taneja
Clinical Assistant Professor of Surgery

Gerald M. Tarnoff
Clinical Assistant Professor of Psychiatry and Human Behavior

John M. Tarro
Clinical Assistant Professor of Surgery

Shannon Lindsay Terkell
Clinical Assistant Professor of Psychiatry and Human Behavior

Anthony G. Thomas
Clinical Assistant Professor of Medicine

Vinod Thomas
Clinical Assistant Professor of Medicine

Bianca Thorpe
Clinical Assistant Professor of Family Medicine

Matthew J. Thran
Clinical Assistant Professor of Anesthesiology

Shiqiang Tian
Clinical Assistant Professor of Orthopaedics

Alexandra M. Tien
Clinical Assistant Professor of Family Medicine

Robert Tilton
Clinical Assistant Professor of Psychiatry and Human Behavior

Cristina Toba
Clinical Assistant Professor of Psychiatry and Human Behavior

David R. Tomlinson
Clinical Assistant Professor of Family Medicine

Andrew J. Tompkins
Clinical Assistant Professor of Surgery

Beth A. Toolan
Clinical Assistant Professor of Pediatrics

Tanya B. C. Tran
Clinical Assistant Professor of Psychiatry and Human Behavior

Christine L. Trask
Clinical Assistant Professor of Psychiatry and Human Behavior

Caroline A. Troise
Clinical Assistant Professor of Medicine

Jean E. Twomey
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Pediatrics

Holly Tytell
Clinical Assistant Professor of Psychiatry and Human Behavior

Jonathan D. Uri
Clinical Assistant Professor of Anesthesiology

Jason Vachon
Clinical Assistant Professor of Medical Science

Frederick Varone
Clinical Assistant Professor of Emergency Medicine

Eric Vaught
Clinical Assistant Professor of Family Medicine

Adrian Velasquez
Clinical Assistant Professor of Medicine

Shelly Verma
Clinical Assistant Professor of Family Medicine

Benjamin S. Vogel
Clinical Assistant Professor Emeritus of Obstetrics and Gynecology

W. Scott Walker
Clinical Assistant Professor of Surgery

Adam Samuel Walpert
Clinical Assistant Professor of Pediatrics

Mary Colleen Walsh
Clinical Assistant Professor of Family Medicine

Michael R. Walther
Clinical Assistant Professor of Psychiatry and Human Behavior

Thomas Warcup
Clinical Assistant Professor of Family Medicine

Marvin S. Wasser
Clinical Assistant Professor of Pediatrics

Raymond H. Welch
Clinical Assistant Professor Emeritus of Dermatology

Jiaying Wen
Clinical Assistant Professor of Medicine

Chien-Hsiang Weng
Clinical Assistant Professor of Family Medicine

Anastasia Wermert
Clinical Assistant Professor of Psychiatry and Human Behavior

Elizabeth Wheeler
Clinical Assistant Professor of Psychiatry and Human Behavior

Bridget Wieczkowski
Clinical Assistant Professor of Medical Science; Clinical Assistant Professor of Pediatrics

Katherine S. Williams
Clinical Assistant Professor of Family Medicine

Julie M. Wilson
Clinical Assistant Professor Emerita of Psychiatry and Human Behavior

Craig Woda
Clinical Assistant Professor of Medical Science

Patricia N. Wold
Clinical Assistant Professor Emerita of Psychiatry and Human Behavior

Michael D. Wolfe
Clinical Assistant Professor of Psychiatry and Human Behavior

Ivan S. Wolfson
Clinical Assistant Professor of Family Medicine

Anthony K. Wong
Clinical Assistant Professor of Orthopaedics

Arthur W. Y. Wong
Clinical Assistant Professor of Surgery

David S. Worman
Clinical Assistant Professor of Orthopaedics

Lora Wu
Clinical Assistant Professor of Psychiatry and Human Behavior

Vanessa Wu
Clinical Assistant Professor of Psychiatry and Human Behavior

Joseph Wyllie
Clinical Assistant Professor of Medicine

Athena Xifaras
Clinical Assistant Professor of Pediatrics

Mark Young
Clinical Assistant Professor of Family Medicine

Dominick Zangari
Clinical Assistant Professor of Surgery

Raymond P. Zarlengo
Clinical Assistant Professor of Pediatrics

Kimberly A. Zeller
Clinical Assistant Professor of Family Medicine

Deqiang Zhang
Clinical Assistant Professor of Medicine

Cara Zimmerman
Clinical Assistant Professor of Medicine

Assistant Professor Research

Arturo Andrade Andrade
Assistant Professor of Brain Science (Research); Assistant Professor of Neuroscience (Research)

Sarah A. Arias
Assistant Professor of Psychiatry and Human Behavior (Research)

Jennifer L. Barredo
Assistant Professor of Psychiatry and Human Behavior (Research)

Aaron V. Berard
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Melanie Lauren Bozzay
Assistant Professor of Psychiatry and Human Behavior (Research)

Tosca Braun
Assistant Professor of Psychiatry and Human Behavior (Research)

Alexander S. Brodsky
Assistant Professor of Pathology and Laboratory Medicine (Research)

Peter Bronk
Assistant Professor of Medicine (Research)

Christy Capone
Assistant Professor of Psychiatry and Human Behavior (Research)

Celeste Caviness
Assistant Professor of Psychiatry and Human Behavior (Research)

Xiaodi Chen
Assistant Professor of Pediatrics (Research)

Anna Dorota Chorzalska
Assistant Professor of Medicine (Research)

Chun-Shiang Chung
Assistant Professor of Surgery (Research)

Alicia Cohen
Assistant Professor of Family Medicine (Research); Assistant Professor of Health Services, Policy and Practice (Research)

Charlene Collibee
Assistant Professor of Psychiatry and Human Behavior (Research)

Brandon Del Pozo
Assistant Professor of Medicine (Research); Assistant Professor of Health Services, Policy and Practice (Research)

Kathryn E. Demos
Assistant Professor of Psychiatry and Human Behavior (Research)

Xiaoqun Dong
Assistant Professor of Medicine (Research)

Yovanska M. Duarte-Velez
Assistant Professor of Psychiatry and Human Behavior (Research)

Erica Eaton
Assistant Professor of Psychiatry and Human Behavior (Research)

Erin Whitney Evans
Assistant Professor of Psychiatry and Human Behavior (Research)

Alexey Fedulov
Assistant Professor of Surgery (Research)

Helen Fuchs
Assistant Professor of Medicine (Research)

Dioscaris Garcia
Assistant Professor of Orthopaedics (Research)

Monica Adriana Garcia Solache
Assistant Professor of Medicine (Research)

Sarah Garnaat
Assistant Professor of Psychiatry and Human Behavior (Research)

Carly Michelle Goldstein
Assistant Professor of Psychiatry and Human Behavior (Research)

Stephanie Paige Goldstein
Assistant Professor of Psychiatry and Human Behavior (Research)

Kathryn J. Grive
Assistant Professor of Obstetrics and Gynecology (Research)

Yingjie Guan
Assistant Professor of Medicine (Research); Assistant Professor of Pathology and Laboratory Medicine (Research)

Sheila A. Haley
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Jacqueline Hayes
Assistant Professor of Psychiatry and Human Behavior (Research)

Kristin Michelle Healey
Assistant Professor of Psychiatry and Human Behavior (Research)

Lindsay Huffhines
Assistant Professor of Psychiatry and Human Behavior (Research)

Nicole James
Assistant Professor of Obstetrics and Gynecology (Research)

Sukanta Jash
Assistant Professor of Pediatrics (Research)

Euy-Myoung Jeong
Assistant Professor of Medicine (Research)

Anatoli Kabakov
Assistant Professor of Medicine (Research)

Suchitra Kamle
Assistant Professor of Molecular Microbiology and Immunology (Research)

Galina Karashchuk
Assistant Professor of Pathology and Laboratory Medicine (Research)

Brian Kavanaugh
Assistant Professor of Psychiatry and Human Behavior (Research)

Joshua Kemp
Assistant Professor of Psychiatry and Human Behavior (Research)

Kathleen Kemp
Assistant Professor of Psychiatry and Human Behavior (Research)

Laura Korthauer
Assistant Professor of Psychiatry and Human Behavior (Research)

Morganne Ashley Kraines
Assistant Professor of Psychiatry and Human Behavior (Research)

Anastacia Y. Kudinova
Assistant Professor of Psychiatry and Human Behavior (Research)

Kirsten Langdon
Assistant Professor of Psychiatry and Human Behavior (Research)

Brian W. LeBlanc
Assistant Professor of Surgery (Research)

Changmin Lee
Assistant Professor of Molecular Microbiology and Immunology (Research)

Peter Lee
Assistant Professor of Pathology and Laboratory Medicine (Research)

Craig T. Lefort
Assistant Professor of Surgery (Research)

Olin Dehui Liang
Assistant Professor of Medicine (Research)

Bing Ma
Assistant Professor of Molecular Microbiology and Immunology (Research)

Emily McDonald
Assistant Professor of Pediatrics (Research); Assistant Professor of Pathology and Laboratory Medicine (Research)

Biswajit Mishra
Assistant Professor of Medicine (Research)

Mandar T. Naik
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

KayLoni Olson
Assistant Professor of Psychiatry and Human Behavior (Research)

Nathalie Oulhen
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Qing Ouyang
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Emily Panza
Assistant Professor of Psychiatry and Human Behavior (Research)

Rebecca K. Papas
Assistant Professor of Psychiatry and Human Behavior (Research)

Ju Park
Assistant Professor of Medicine (Research); Assistant Professor of Epidemiology (Research)

Jessica R. Peters
Assistant Professor of Psychiatry and Human Behavior (Research)

Barbara Porton
Assistant Professor of Psychiatry and Human Behavior (Research)

Frederic Pouille
Assistant Professor of Neuroscience (Research)

Jennifer M. Primack
Assistant Professor of Psychiatry and Human Behavior (Research)

Jennifer R. Ribeiro
Assistant Professor of Obstetrics and Gynecology (Research)

Giulia Righi
Assistant Professor of Psychiatry and Human Behavior (Research)

Karim Roder
Assistant Professor of Medicine (Research)

Brooke Genkin Rogers
Assistant Professor of Medicine (Research); Assistant Professor of Psychiatry and Human Behavior (Research)

Laura Sanapo
Assistant Professor of Medicine (Research)

Heather T. Schatten
Assistant Professor of Psychiatry and Human Behavior (Research)

Christoph Schorl
Assistant Professor of Biology (Research)

Eui Young So
Assistant Professor of Medicine (Research)

Estate of Angela K. Stevens
Assistant Professor of Psychiatry and Human Behavior (Research)

Xiaoli Tang
Assistant Professor of Medicine (Research)

Jun Tao
Assistant Professor of Medicine (Research)

Nicholas Tarantino
Assistant Professor of Psychiatry and Human Behavior (Research)

Sarah Ann Thomas
Assistant Professor of Psychiatry and Human Behavior (Research)

Xiaobing Tian
Assistant Professor of Pathology and Laboratory Medicine (Research)

Nilufer Nermin Turan Dural
Assistant Professor of Medicine (Research)

Lori A. Underhill
Assistant Professor of Pediatrics (Research)

Sara Vargas
Assistant Professor of Psychiatry and Human Behavior (Research)

Carlos E. Vargas-Irwin
Assistant Professor of Neuroscience (Research)

Charles A. Vaslet
Assistant Professor of Pathology and Laboratory Medicine (Research)

Edward G. Walsh
Assistant Professor of Neuroscience (Research)

Xiaotian Wang
Assistant Professor of Surgery (Research)

Sicheng Wen
Assistant Professor of Medicine (Research)

Michael S. Worden
Assistant Professor of Neuroscience (Research)

Mamiko Yajima
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Emine Bihter Yalcin
Assistant Professor of Pathology and Laboratory Medicine (Research)

Shengliang Zhang
Assistant Professor of Pathology and Laboratory Medicine (Research)

Songhua Zhang
Assistant Professor of Medicine (Research)

Lanlan Zhou
Assistant Professor of Pathology and Laboratory Medicine (Research)

Xiaoxu Zhou
Assistant Professor of Pathology and Laboratory Medicine (Research)

Distinguished Senior Lecturer

John J. Stein
Distinguished Senior Lecturer in Neuroscience

Senior Lecturer

Amy Ellen Chew
Senior Lecturer in Ecology, Evolution, and Organismal Biology

Aisling S. Dugan
Senior Lecturer in Biology; Senior Lecturer in Molecular Microbiology and Immunology

Diana M. Horrigan
Senior Lecturer in Biology; Senior Lecturer in Pathology and Laboratory Medicine

Monica Linden
Distinguished Senior Lecturer in Neuroscience

Dale A. Ritter
Senior Lecturer in Biology

Lecturer

Toni-Marie Achilli
Lecturer in Biology

William M. Holmes
Lecturer in Biology

Rebecca Y. Kartzinel
Lecturer in Biology Education; Lecturer in Ecology, Evolution, and Organismal Biology

Kristina A. Monteiro
Lecturer in Biology Education

Babette Taylor Stewart
Lecturer Emerita in Medical Science

Instructor

Terrence Vance
Instructor in Dermatology; Instructor in Epidemiology

Clinical Instructor

Aamir Ahmad
Clinical Instructor in Medical Science

Sarah Anis
Clinical Instructor in Surgery

Mohamed-Omar S. Arafeh
Clinical Instructor in Surgery

Luke Barre
Clinical Instructor in Medical Science

Mark Barsamian
Clinical Instructor in Surgery

William C. Bastan
Clinical Instructor Emeritus in Pathology and Laboratory Medicine

Katherine A. Baysinger
Clinical Instructor in Surgery

Michelle Beller
Clinical Instructor in Medical Science

Laureen Berkowitz
Clinical Instructor in Medical Science

David S. Bick
Clinical Instructor in Surgery

Richard Black
Clinical Instructor in Medical Science

Madalene Irene Boyle
Clinical Instructor in Emergency Medicine

Alex Brake
Clinical Instructor in Psychiatry and Human Behavior

John F. Butler
Clinical Instructor in Pediatrics

Jessica M. Cappellano
Clinical Instructor in Psychiatry and Human Behavior

Timothy D. Carroll
Clinical Instructor in Psychiatry and Human Behavior

Claudia A. Clarke
Clinical Instructor in Medical Science

Jane Cook
Clinical Instructor in Surgery

Maureen Crotty
Clinical Instructor in Pediatrics

Jennifer Anne Cueto
Clinical Instructor in Psychiatry and Human Behavior; Clinical Instructor in Pediatrics

Giuseppe D'Amelio
Clinical Instructor in Psychiatry and Human Behavior

Christina Marie D'Angelo
Clinical Instructor in Psychiatry and Human Behavior

Pamela Dana-Snyder
Clinical Instructor in Medical Science

Jenny David
Clinical Instructor in Obstetrics and Gynecology

Laura H. Dawson
Clinical Instructor in Pediatrics

Normand L. Decelles
Clinical Instructor in Medical Science

Suvekchha Devkota
Clinical Instructor in Medical Science

Heather Di Biasio
Clinical Instructor in Pediatrics

Charissa DiNobile
Clinical Instructor in Obstetrics and Gynecology

Matthew R. Donati
Clinical Instructor in Psychiatry and Human Behavior

Calin Drimbarean
Clinical Instructor in Medical Science

Ibrahim Eid
Clinical Instructor in Medical Science

Alaa Elnajjar
Clinical Instructor in Psychiatry and Human Behavior

Raymond Entwistle
Clinical Instructor in Pediatrics

Amy Farb
Clinical Instructor in Pediatrics

Jessica C. Farrar
Clinical Instructor in Pediatrics

Marissa A. Fast
Clinical Instructor in Pediatrics

Antonio Fernandez
Clinical Instructor in Medical Science

Ziad Fersan
Clinical Instructor in Obstetrics and Gynecology

Sarah Filone
Clinical Instructor in Psychiatry and Human Behavior

Carolyn Fleming-Fruci
Clinical Instructor in Medical Science

Barbara Miller Gaines
Clinical Instructor in Pediatrics

Jennifer M. Gassner
Clinical Instructor in Obstetrics and Gynecology

James Giddings
Clinical Instructor in Emergency Medicine

Kelvin D. Gillman
Clinical Instructor in Pediatrics

Ritu Goel
Clinical Instructor in Surgery

Michael Daniel Gomez
Clinical Instructor in Psychiatry and Human Behavior

Carin A. Got
Clinical Instructor in Anesthesiology

Scott Greenberg
Clinical Instructor in Surgery

Christine Greenia
Clinical Instructor in Psychiatry and Human Behavior

Benjamin Michael Hall
Clinical Instructor of Surgery

Heather Hall
Clinical Instructor in Psychiatry and Human Behavior

Frederick Hartman
Clinical Instructor in Surgery

Silvia M. Herrera Pinilla
Clinical Instructor in Obstetrics and Gynecology

Dylan Hershkowitz
Clinical Instructor in Psychiatry and Human Behavior

Caroline Sophie Holman
Clinical Instructor in Psychiatry and Human Behavior

Christopher Joncas
Clinical Instructor in Medical Science

Lisa Jones
Clinical Instructor in Medical Science

Carmen Khorram
Clinical Instructor in Medical Science

Jacob Adrian Lafo
Clinical Instructor in Psychiatry and Human Behavior

Thomas J. Lavin
Clinical Instructor in Psychiatry and Human Behavior

Naveh Levy
Clinical Instructor in Medical Science

Steve Bernard Louvet
Clinical Instructor in Family Medicine

Gary K. Lundstrom
Clinical Instructor in Diagnostic Imaging

Katherine P. MacCallum
Clinical Instructor of Surgery

Ian Madom
Clinical Instructor in Medical Science

Daniel Joseph Manfra
Clinical Instructor in Psychiatry and Human Behavior

Russell Morgan Marks
Clinical Instructor in Psychiatry and Human Behavior

Andrew C. Martina
Clinical Instructor in Psychiatry and Human Behavior

Melissa A. McWilliams
Clinical Instructor in Psychiatry and Human Behavior

Edward George Medeiros, Jr
Clinical Instructor in Medical Science

Leslie C. Mohlman
Clinical Instructor in Pediatrics

Dana Morris
Clinical Instructor in Psychiatry and Human Behavior

Adelaide G. Nardone
Clinical Instructor in Medical Science

Mitch Chikadibie Otu
Clinical Instructor in Psychiatry and Human Behavior

Monika Page
Clinical Instructor in Medical Science

Angelina Rose Palombo
Clinical Instructor in Medical Science

Susanne J. Patrick-MacKinnon
Clinical Instructor in Medical Science

Lori Ann Maria Perretta
Clinical Instructor in Psychiatry and Human Behavior

Benjamin Z. Phillips
Clinical Instructor in Medical Science

Loreen P. Pirnie
Clinical Instructor in Psychiatry and Human Behavior

Dieter Pohl
Clinical Instructor in Surgery

Colleen A. Powers
Clinical Instructor in Pediatrics

Victoria Elena Quinones
Clinical Instructor in Psychiatry and Human Behavior

Jennifer J. Raney
Clinical Instructor in Medical Science

Deborah Ann Rasile
Clinical Instructor in Psychiatry and Human Behavior

Nicole Rivera
Clinical Instructor in Psychiatry and Human Behavior

Dana Rosen
Clinical Instructor in Psychiatry and Human Behavior

Haitham Salem
Clinical Instructor in Psychiatry and Human Behavior

Estefania Santamaria Flores
Clinical Instructor in Obstetrics and Gynecology

Maria Santini
Clinical Instructor in Pediatrics

Kathryn Simon
Clinical Instructor in Psychiatry and Human Behavior

Ronald Smith
Clinical Instructor in Psychiatry and Human Behavior

Timothy Steinhoff
Clinical Instructor in Psychiatry and Human Behavior

Julia Tassinari
Clinical Instructor in Surgery

Tanya Thomas
Clinical Instructor in Psychiatry and Human Behavior

Keri Tilman
Clinical Instructor in Emergency Medicine

Jessica Torres
Clinical Instructor in Obstetrics and Gynecology

Yolanda Troublefield
Clinical Instructor in Medical Science

Lindsay Tse
Clinical Instructor in Medical Science

Dustin J. Tubre
Clinical Instructor in Medical Science

James Valente
Clinical Instructor in Surgery

Katherine Visser
Clinical Instructor in Psychiatry and Human Behavior

Elizabeth Stanton Wagner
Clinical Instructor in Psychiatry and Human Behavior

Micaela Anne Weaver
Clinical Instructor in Surgery; Clinical Instructor in Obstetrics and Gynecology

Colette Whitby
Clinical Instructor in Medical Science

Hugo Yamada
Clinical Instructor in Medical Science

Shiwen Yuan
Clinical Instructor in Psychiatry and Human Behavior

Alice Zhou
Clinical Instructor in Psychiatry and Human Behavior

Instructor Research

Katherine Elizabeth Darling
Instructor in Psychiatry and Human Behavior (Research)

Alger M. Fredericks
Instructor in Surgery (Research)

Haruki Higashimori
Instructor in Molecular Biology, Cell Biology and Biochemistry (Research)

Christopher Mantsounga
Instructor in Medicine (Research)

Leah Schumacher
Instructor in Psychiatry and Human Behavior (Research)

Miiko Jaakkima Sokka
Instructor in Molecular Biology, Cell Biology and Biochemistry (Research)

Nagendran Tharmalingam
Instructor in Medicine (Research)

Lijun Wang
Instructor in Orthopaedics (Research)

Investigator

Jacqueline Hynes
Investigator in Neuroscience

Nan Jiang
Investigator in Molecular Biology, Cell Biology and Biochemistry

Li Ma
Investigator in Molecular Biology, Cell Biology and Biochemistry

Chen Ming
Investigator of Neuroscience

Tanbir Najrana
Investigator in Pathology and Laboratory Medicine

Tanya C. Sanders
Investigator in Biology Education

Thomas Taetzsch
Investigator in Molecular Biology, Cell Biology and Biochemistry

Jackson Taylor
Investigator in Molecular Biology, Cell Biology and Biochemistry

Adjunct Professor

Ross Abrams
Adjunct Professor of Radiation Oncology

Fatemeh Akhlaghi
Adjunct Professor of Medicine

Mostafa Analoui
Adjunct Professor of Biology Education

Thomas F. Anders
Adjunct Professor of Psychiatry and Human Behavior

Kenneth A. Arndt
Adjunct Professor of Dermatology

David B. Baier
Adjunct Professor of Ecology, Evolution, and Organismal Biology

Charlotte M. Boney
Adjunct Professor of Pediatrics

Phil Brown
Adjunct Professor of Pathology and Laboratory Medicine

Giovanni Camussi
Adjunct Professor of Medicine

Zoe G. Cardon
Adjunct Professor of Ecology, Evolution, and Organismal Biology

Anjulika Chawla
Adjunct Professor of Pediatrics

Burke Cunha
Adjunct Professor of Medicine

Richard Dollase
Adjunct Professor of Medical Science

Samuel Dudley
Adjunct Professor of Medicine

Damian E. Dupuy
Adjunct Professor of Diagnostic Imaging

Stephen Erickson
Adjunct Professor of Psychiatry and Human Behavior

Sarah Feldstein Ewing
Adjunct Professor of Psychiatry and Human Behavior

Lynne A. Fontaine
Adjunct Professor of Pediatrics

Michael G. Goldstein
Adjunct Professor of Psychiatry and Human Behavior

Eric S. Hall
Adjunct Professor of Radiation Oncology

Anne L. Hume
Adjunct Professor of Family Medicine

Mario Jiz
Adjunct Professor of Pathology and Laboratory Medicine

Jeff G. Konin
Adjunct Professor of Family Medicine

Kerry L. Laplante
Adjunct Professor of Medicine

Margaret Lartey
Adjunct Professor of Medicine

Yow-Pin Lim
Adjunct Professor of Pathology and Laboratory Medicine

Bing Lu
Adjunct Professor of Family Medicine

Kenneth Hugh Mayer
Adjunct Professor of Medicine; Adjunct Professor of Epidemiology

Stefan McDonough
Adjunct Professor of Brain Science

Alyson J. McGregor
Adjunct Professor of Emergency Medicine

Judith S. Mercer
Adjunct Professor of Pediatrics

Justin M. Nash
Adjunct Professor of Family Medicine

John Ong'echa
Adjunct Professor of Pathology and Laboratory Medicine

Wolfgang Oster
Adjunct Professor of Pathology and Laboratory Medicine

Michael Oumano
Adjunct Professor of Radiation Oncology

Clayton Peimer
Adjunct Professor of Orthopaedics

Shmuel Reis
Adjunct Professor of Family Medicine

Caroline R. Richardson
Adjunct Professor of Family Medicine

Lynn Rothschild
Adjunct Professor of Molecular Biology, Cell Biology and Biochemistry

Amy Lynn Salisbury
Adjunct Professor of Pediatrics

Lori Scott-Sheldon
Adjunct Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Ronald Seifer
Adjunct Professor of Psychiatry and Human Behavior

Sheldon R. Simon
Adjunct Professor of Orthopaedics

Gail Skowron
Adjunct Professor of Medicine

Robert W. Sobol
Adjunct Professor of Pathology and Laboratory Medicine

Robert L. Stout
Adjunct Professor of Psychiatry and Human Behavior

Mary C. Sullivan
Adjunct Professor of Pediatrics

Jin Bo Tang
Adjunct Professor of Surgery

Tracey Taveira
Adjunct Professor of Medicine

Julie S. Taylor
Adjunct Professor of Family Medicine

Peter L. Tilkemeier
Adjunct Professor of Medicine

Thomas F. Tracy
Adjunct Professor of Surgery

Sivamainthan Vithiananthan
Adjunct Professor of Surgery

Nicholas John Wald
Adjunct Professor of Pathology and Laboratory Medicine

Risa Weisberg
Adjunct Professor of Family Medicine

Hagop Youssoufian
Adjunct Professor of Medicine

Adjunct Professor Research

Djuro Josic
Adjunct Professor of Medicine (Research)

Adjunct Associate Professor

Geoffrey Allen
Adjunct Associate Professor of Pediatrics

Christina L. Boisseau
Adjunct Associate Professor of Psychiatry and Human Behavior

Kevin J. Chang
Adjunct Associate Professor of Diagnostic Imaging

Denise Coppa
Adjunct Associate Professor of Pediatrics

R. John Davenport
Adjunct Associate Professor of Neuroscience

Jeffrey Dover
Adjunct Associate Professor of Dermatology

Patrick J. Ewanchuk
Adjunct Associate Professor of Ecology, Evolution, and Organismal Biology

Sarah A. Gaines
Adjunct Associate Professor of Emergency Medicine

Andrea Goldschmidt
Adjunct Associate Professor of Psychiatry and Human Behavior

Thomas J. Guilmette
Adjunct Associate Professor of Psychiatry and Human Behavior

Nadine T. Himelfarb
Adjunct Associate Professor of Emergency Medicine

Daniel J. Holmander
Adjunct Associate Professor of Biology Education

Brian W. Jack
Adjunct Associate Professor of Family Medicine

Liudvikas Jagminas
Adjunct Associate Professor of Medical Science

Yong S. Jong
Adjunct Associate Professor of Biology Education

Magdalena Krzystolik
Adjunct Associate Professor of Surgery

Celia P. MacDonnell
Adjunct Associate Professor of Family Medicine

Kristen A. Matteson
Adjunct Associate Professor of Obstetrics and Gynecology

Lisa H. Merck
Adjunct Associate Professor of Neurosurgery

Emily S. Miller
Adjunct Associate Professor of Obstetrics and Gynecology

Mohammed Morad
Adjunct Associate Professor of Family Medicine

Melissa Brooks Nothnagle
Adjunct Associate Professor of Family Medicine

Robert A. Partridge
Adjunct Associate Professor of Emergency Medicine

Robert Hamilton Pierce
Adjunct Associate Professor of Pathology and Laboratory Medicine

Christie J. Rizzo
Adjunct Associate Professor of Psychiatry and Human Behavior

Katina M. Robison
Adjunct Associate Professor of Obstetrics and Gynecology

Stephen J. Sheinkopf
Adjunct Associate Professor of Psychiatry and Human Behavior; Adjunct Associate Professor of Pediatrics

Shawn Riser Taylor
Adjunct Associate Professor of Family Medicine

Rajan Wadhawan
Adjunct Associate Professor of Pediatrics

Richard Wein
Adjunct Associate Professor of Surgery

Erika Werner
Adjunct Associate Professor of Obstetrics and Gynecology

Shirley Yen
Adjunct Associate Professor of Psychiatry and Human Behavior

Adjunct Associate Professor Research

Nicola M. Kouttab
Adjunct Associate Professor of Pathology and Laboratory Medicine (Research)

Attila A. Seyhan
Adjunct Associate Professor of Pathology and Laboratory Medicine (Research)

Adjunct Assistant Professor

Priya Banerjee
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Michael A. Battey
Adjunct Assistant Professor of Medical Science

Erika L. Bloom
Adjunct Assistant Professor of Medicine

Barrett W. Bready
Adjunct Assistant Professor of Biology Education

Claudia S. R. Cabrera
Adjunct Assistant Professor of Medicine

Ariel L. Camp
Adjunct Assistant Professor of Ecology, Evolution, and Organismal Biology

Paul Carmichael
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Jennifer J. Carroll
Adjunct Assistant Professor of Medicine

Brian P. Casserly
Adjunct Assistant Professor of Medicine

Ayesha Cheema-Hasan
Adjunct Assistant Professor of Pediatrics

Doreen M. Ciancaglini
Adjunct Assistant Professor of Pediatrics

Richard Clements
Adjunct Assistant Professor of Surgery

Matthew J. Collins
Adjunct Assistant Professor of Family Medicine

Selby Conrad
Adjunct Assistant Professor of Psychiatry and Human Behavior

Amy D'Agata
Adjunct Assistant Professor of Pediatrics

Catherine Degood
Adjunct Assistant Professor of Family Medicine

Yu-Ting L. Dingle
Adjunct Assistant Professor of Biology Education

Aaron Mark Drucker
Adjunct Assistant Professor of Dermatology

Naomi Bloom Gershon
Adjunct Assistant Professor of Pediatrics

Starrina Gianelloni
Adjunct Assistant Professor of Pediatrics

Katarzyna Gilek-Seibert
Adjunct Assistant Professor of Medicine

Volkan M. Gurel
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Karen M. Harnett
Adjunct Assistant Professor of Medical Science

Rachel Herz
Adjunct Assistant Professor of Psychiatry and Human Behavior

Lauri A. Hicks
Adjunct Assistant Professor of Medicine

Eric K. Huang
Adjunct Assistant Professor of Biology Education

Blanche C. Ip
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Beth A. Jerskey
Adjunct Assistant Professor of Psychiatry and Human Behavior

Bernd Laudenberg
Adjunct Assistant Professor of Family Medicine

Wen-Qing Li
Adjunct Assistant Professor of Dermatology

Laszlo Madaras
Adjunct Assistant Professor of Family Medicine

Timothy Mariano
Adjunct Assistant Professor of Psychiatry and Human Behavior

Jeffrey P. Migneault
Adjunct Assistant Professor of Family Medicine

Jeffrey S. Moffit
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Thomas F. Morgan
Adjunct Assistant Professor of Neurology

Alycia Mosley Austin
Adjunct Assistant Professor of Neuroscience

Katja Odening
Adjunct Assistant Professor of Medicine

Nimesh R. Patel
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Dipak K. Raj
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Brenda N. Roy
Adjunct Assistant Professor of Obstetrics and Gynecology

Kristin M. Scaplen
Adjunct Assistant Professor of Neuroscience

Anna Schierberl Scherr
Adjunct Assistant Professor of Psychiatry and Human Behavior

Stanley Schwartz
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Christopher G. Thanos
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Christina C. Tortolani
Adjunct Assistant Professor of Psychiatry and Human Behavior

Peter M. Yeracaris
Adjunct Assistant Professor of Family Medicine

Carolyn Te Young
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Mark Zervas
Adjunct Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Adjunct Lecturer

Melanie Dubard
Adjunct Lecturer in Psychiatry and Human Behavior

Adjunct Instructor

Briannan E. Bintz
Adjunct Instructor in Pathology and Laboratory Medicine

Chester Hedgepeth
Adjunct Instructor in Medicine

Thomas E. Noonan
Adjunct Instructor in Medicine

Ralph Philosophe
Adjunct Instructor in Obstetrics and Gynecology

Barbara Stricker
Adjunct Instructor in Obstetrics and Gynecology

Visiting Scholar

Rawan Alatigue
Visiting Scholar in Emergency Medicine

Ibrahem Aldeghaither
Visiting Scholar in Emergency Medicine

Norah Al Rasheed
Visiting Scholar in Emergency Medicine

Esam Barnawi
Visiting Scholar in Emergency Medicine

Rayane Brinck Teixeira
Visiting Scholar in Surgery

Karla Da Silva Braga
Visiting Scholar in Surgery

Khaled A. Elsaid
Visiting Scholar in Emergency Medicine

Stacia Furtado
Visiting Scholar in Pathology and Laboratory Medicine

Maite Larrucea Iruretagoyena
Visiting Scholar in Psychiatry and Human Behavior

Orsolya Beatrix Kolozsvari
Visiting Scholar in Neuroscience

Camilo Osorio
Visiting Scholar in Orthopaedics

Mengia Pedotti
Visiting Scholar in Molecular Biology, Cell Biology and Biochemistry

Inesh Sood
Visiting Scholar in Molecular Biology, Cell Biology and Biochemistry

Steffanie Tamayo
Visiting Scholar in Pathology and Laboratory Medicine

Roberta Tedesco
Visiting Scholar in Psychiatry and Human Behavior

Ritesh Thakare
Visiting Scholar in Surgery

Hang Xing
Visiting Scholar in Surgery

Jun Zhang
Visiting Scholar in Pathology and Laboratory Medicine

Visiting Scientist

Ozgun Yagmur
Visiting Scientist in Molecular Biology, Cell Biology and Biochemistry

Senior Research Associate

Jennifer F. Carr
Senior Research Associate in Molecular Biology, Cell Biology and Biochemistry

Sheryl Kopel
Senior Research Associate in Psychiatry and Human Behavior; Senior Research Associate in Pediatrics

N. Kumarasamy
Senior Research Associate in Medicine

David Paller
Senior Research Associate in Orthopaedics

Yutaka Yamamoto
Senior Research Associate in Molecular Biology, Cell Biology and Biochemistry

Research Associate

Keeley Baker
Research Associate in Neuroscience

Julie R. Bromberg
Research Associate in Emergency Medicine

Jorge Isaac Chang
Research Associate in Neuroscience

Ray Hung Chen
Research Associate in Pediatrics

Sharon Chu
Research Associate in Pathology and Laboratory Medicine

Susannah Colt
Research Associate in Pediatrics

Anthony B. Comeau
Research Associate in Neuroscience

Christopher A. Deister
Research Associate in Neuroscience

Salomi Desai
Research Associate in Orthopaedics

Elizabeth E. Eklund
Research Associate in Pathology and Laboratory Medicine

Lewis Oscar Felix Raj Lucas
Research Associate in Medicine

Maryam Ghandali
Research Associate in Pathology and Laboratory Medicine

Scott J. Gratz
Research Associate in Neuroscience

Priyadarshini S. Hirway
Research Associate in Pediatrics

Calvin Lukas Kienbacher
Research Associate in Emergency Medicine

Supaporn Kulthinee
Research Associate in Surgery

Aastha Kumari
Research Associate in Neuroscience

Helena Kwakwa
Research Associate in Medicine

Eunjeong Lee
Research Associate in Neuroscience

Xiaofei Li
Research Associate in Medicine

Yajun Liu
Research Associate in Orthopaedics

Edwin Lok
Research Associate in Medicine

Huafei Lu
Research Associate in Medicine

Adam Mani
Research Associate in Neuroscience

Margot Martinez Moreno
Research Associate in Neurosurgery

Michaela Maynard
Research Associate in Medicine

Sharon M. McKenzie
Research Associate in Medicine

Ryan Lloyd Miller
Research Associate in Neuroscience

Nahid Mohammadzadeh
Research Associate in Neurology

Seyedamin Moosavi
Research Associate in Neuroscience

Amy Morton
Research Associate in Orthopaedics

Evangelia Mylona
Research Associate in Medicine

Siena C. Napoleon
Research Associate in Medicine

Sonia Nesbitt
Research Associate in Pediatrics

Karim Oueidat
Research Associate in Diagnostic Imaging

Tuan Pham
Research Associate in Neuroscience

Andrea Pierre
Research Associate in Neuroscience

Zhen Qiao
Research Associate in Orthopaedics

Holly A. Richendrfer
Research Associate in Emergency Medicine

Mehrnoosh Samaei
Research Associate in Emergency Medicine

Elif Sengun
Research Associate in Medicine

Fadi Shehadeh
Research Associate in Medicine

Soya Sisy Sam
Research Associate in Medicine

Altar Sorkac
Research Associate in Neuroscience

Joan Stabila
Research Associate in Pediatrics

Xiaojuan Sun
Research Associate in Orthopaedics

Vida Tajiknia
Research Associate in Pathology and Laboratory Medicine

Aleksandar Todorovich
Research Associate in Diagnostic Imaging

Jay Trivedi
Research Associate in Orthopaedics

Shu-Whei Tsai
Research Associate in Pediatrics

Lijiang Wang
Research Associate in Surgery

Naohiro Yano
Research Associate in Surgery

Yuki Yokota
Research Associate in Medicine

Yiqun Zhang
Research Associate in Pathology and Laboratory Medicine

Panagiotis Ziakas
Research Associate in Medicine

Teaching Associate

Kimberly A. Adams
Teaching Associate in Medical Science

Laura L. Adams
Teaching Associate in Family Medicine

Kerry Ahern
Teaching Associate in Obstetrics and Gynecology

Barbara Alksninis
Teaching Associate in Pediatrics

Jhanev Allen
Teaching Associate in Medical Science

Ana Sofia BarberDeBrito
Teaching Associate in Obstetrics and Gynecology

Kathryn Basques
Teaching Associate in Psychiatry and Human Behavior

Patricia L. Beauchemin
Teaching Associate in Medical Science

Linda Beetlestone
Teaching Associate in Obstetrics and Gynecology

Katherine Y. Benevides
Teaching Associate in Family Medicine

Cindy Benson
Teaching Associate in Pediatrics

Mary Clare Bergen
Teaching Associate in Medicine

Kathleen E. Boyd
Teaching Associate in Medicine

Michael Brand
Teaching Associate in Medical Science

Susan Bridges
Teaching Associate in Anesthesiology

Patricia Brinegar
Teaching Associate in Family Medicine

Michelle Buffington
Teaching Associate in Obstetrics and Gynecology

William Campbell
Teaching Associate in Pediatrics

Tara A. Capuano
Teaching Associate in Obstetrics and Gynecology

Ariana Ann Carvajal
Teaching Associate in Obstetrics and Gynecology

Mary Lynn Casey
Teaching Associate in Family Medicine

Ceceley Chambers
Teaching Associate in Medical Science

Mary-Kathryn Chambers
Teaching Associate in Pediatrics

Benjamin Choiniere
Teaching Associate in Medical Science

Cynthia A. Corriveau
Teaching Associate in Family Medicine

Lauren Elizabeth Costantin
Teaching Associate in Medical Science

Stacie L. Demarais
Teaching Associate in Family Medicine

Judy DeOliveira
Teaching Associate in Medical Science

Susan DiBlasi
Teaching Associate in Obstetrics and Gynecology

Randia M. Dickerson
Teaching Associate in Medical Science

Emily DiPalma
Teaching Associate in Medical Science

Laura Doherty
Teaching Associate in Obstetrics and Gynecology

Dawn Drake
Teaching Associate in Surgery

Elizabeth Dyer
Teaching Associate in Family Medicine

Aaron Estrela
Teaching Associate in Psychiatry and Human Behavior

Samantha Ewart
Teaching Associate in Medical Science

Jacalyn Finerty
Teaching Associate in Obstetrics and Gynecology

Elana B. Finn
Teaching Associate in Family Medicine

Melissa Fitzgerald
Teaching Associate in Pediatrics

Leanne Flynn
Teaching Associate in Pediatrics

Nicole F. Flynn
Teaching Associate in Pediatrics

Lucia Fontes-Borts
Teaching Associate in Medical Science

Sherry Gagnon
Teaching Associate in Pediatrics

Lisa Gaumond
Teaching Associate in Medical Science

Megan Gernt
Teaching Associate in Surgery

Shelley J. Gertz Sigal
Teaching Associate in Medical Science

Kathleen Guarino
Teaching Associate in Pediatrics

Jacquelyn Halliday
Teaching Associate in Obstetrics and Gynecology

Briana J. Hawksley
Teaching Associate in Pediatrics

Bridget Henley
Teaching Associate in Surgery

Jane Hesser
Teaching Associate in Psychiatry and Human Behavior

Scott Hewitt
Teaching Associate in Family Medicine

James Hodson
Teaching Associate in Family Medicine

Juliette Holtzman
Teaching Associate in Medical Science

Melissa-Lauren Claudette Hooks
Teaching Associate in Medical Science

Elizabeth Howland
Teaching Associate in Obstetrics and Gynecology

Debra Hurwitz
Teaching Associate in Family Medicine

Joan L. Joseph
Teaching Associate in Medical Science

Koren V. Kanadanian
Teaching Associate in Emergency Medicine

Debra Karinski
Teaching Associate in Pediatrics

Elizabeth Kettyle
Teaching Associate in Obstetrics and Gynecology

Yeana Kim
Teaching Associate in Surgery

William Kirkpatrick
Teaching Associate in Medical Science

Sarah Kleinman
Teaching Associate in Obstetrics and Gynecology

Anne Kugler
Teaching Associate in Psychiatry and Human Behavior

Jill A. Lamson
Teaching Associate in Obstetrics and Gynecology

Andrew Lewandowski
Teaching Associate in Medical Science

Zi Ying Li
Teaching Associate in Medicine

Meghan Lovell
Teaching Associate in Obstetrics and Gynecology

Alexandra Sophie Marchioro
Teaching Associate in Medical Science

Veronica Marquis
Teaching Associate in Obstetrics and Gynecology

Sheila Martin
Teaching Associate in Surgery

Lauren L. Marvell
Teaching Associate in Surgery

Catherine Margaret May
Teaching Associate in Family Medicine

Virginia McCann
Teaching Associate in Pediatrics

Deborah A. McKenna
Teaching Associate in Psychiatry and Human Behavior

Fatima Medeiros
Teaching Associate in Family Medicine

Molly Merlino
Teaching Associate in Obstetrics and Gynecology

Lindsey Moran
Teaching Associate in Surgery

Alexandra Morang Jackson
Teaching Associate in Medical Science

Melissa Moreau
Teaching Associate in Pediatrics

Kathleen Moren
Teaching Associate in Family Medicine

Laura L. Moynihan
Teaching Associate in Medical Science

Heather W. Murphy
Teaching Associate in Obstetrics and Gynecology

Lisa M. Murphy
Teaching Associate in Emergency Medicine

Crystal M. Nadeau
Teaching Associate in Family Medicine

Linda Nanni
Teaching Associate in Obstetrics and Gynecology

Kate B. Noveau
Teaching Associate in Family Medicine

Timothy Owens
Teaching Associate in Psychiatry and Human Behavior

Anthony Parente
Teaching Associate in Medical Science

Allison M. Parkhurst
Teaching Associate in Family Medicine

Abigail Patenaude
Teaching Associate in Pediatrics

Karen Pendleton
Teaching Associate in Medicine

Lorraine E. Peterson
Teaching Associate in Medical Science

Lauren Pillsbury
Teaching Associate in Medical Science

Beth A. Pinkos
Teaching Associate in Pediatrics

Robert Plante
Teaching Associate in Psychiatry and Human Behavior

Fernanda Poyant
Teaching Associate in Medical Science

Michaela Procaccini
Teaching Associate in Orthopaedics

Louis J. Pugliese
Teaching Associate in Family Medicine

Stacy Ramsey
Teaching Associate in Obstetrics and Gynecology

Elizabeth S. Ricci
Teaching Associate in Obstetrics and Gynecology

Glenn Ricci
Teaching Associate in Medical Science

Ruby Ristuccia
Teaching Associate in Obstetrics and Gynecology

Mary Roberts
Teaching Associate in Family Medicine

Barbara B. Robinson
Teaching Associate in Pediatrics

Thomas Rodrigues
Teaching Associate in Medical Science

Shimona Romney
Teaching Associate in Medical Science

Barbara Roney
Teaching Associate in Psychiatry and Human Behavior

Jennifer Sardella
Teaching Associate in Pediatrics

Jennifer Scalia
Teaching Associate in Obstetrics and Gynecology

Jennifer L. Schwab
Teaching Associate in Pediatrics

Rebecca Lovell Scott
Teaching Associate in Medical Science

Vrithi Shah
Teaching Associate in Pediatrics

Carolyn Slack
Teaching Associate in Obstetrics and Gynecology

Christopher Sonntag
Teaching Associate in Medicine

Rebecca Sousa
Teaching Associate in Medical Science

Sarah Spinette
Teaching Associate in Pediatrics

Angela Stupinski
Teaching Associate in Family Medicine

Katie F. Tavares-Medeiros
Teaching Associate in Family Medicine

M. Diane Thompson
Teaching Associate in Family Medicine; Teaching Associate in Medical Science

Kathleen M. Tokarski
Teaching Associate in Medical Science

Christopher Tucci
Teaching Associate in Surgery

Deanna P. Upchurch
Teaching Associate in Medical Science

Laura M. Valois
Teaching Associate in Medical Science

Vanessa M. Ventura
Teaching Associate in Surgery

Sandra Victorino
Teaching Associate in Medical Science

Judith Walker
Teaching Associate in Family Medicine

Sherry Walters
Teaching Associate in Surgery

William Watkins
Teaching Associate in Orthopaedics

Sarah White
Teaching Associate in Surgery

Geoffrey L. Williams
Teaching Associate in Molecular Biology, Cell Biology and Biochemistry

Mandy Witkin
Teaching Associate in Psychiatry and Human Behavior

Jessica Wood
Teaching Associate in Family Medicine

Noell Woolley
Teaching Associate in Medical Science

Wendy Beth Young
Teaching Associate in Obstetrics and Gynecology

Associate Professor, Clinician Educator

Saurabh Agarwal
Associate Professor of Diagnostic Imaging, Clinician Educator

Tanya Ali
Associate Professor of Medicine, Clinician Educator

Jason M. Aliotta
Associate Professor of Medicine, Clinician Educator

Lauren Allister
Associate Professor of Emergency Medicine, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

Khaldoun Almhanna
Associate Professor of Medicine, Clinician Educator

Angela C. Anderson
Associate Professor of Pediatrics, Clinician Educator; Associate Professor of Emergency Medicine, Clinician Educator

Michelle Stozek Anvar
Associate Professor of Medicine, Clinician Educator

Marcella Aquino
Associate Professor of Pediatrics, Clinician Educator

Andrea E. Arena
Associate Professor of Family Medicine, Clinician Educator

Karen E. Aspry
Associate Professor of Medicine, Clinician Educator

Christopher Azzoli
Associate Professor of Medicine, Clinician Educator

Christina A. Bandera
Associate Professor of Surgery, Clinician Educator

James Barbeau
Associate Professor of Pathology and Laboratory Medicine, Clinician Educator

Jacob R. Berger
Associate Professor of Neurology, Clinician Educator

Reena A. Bhatt
Associate Professor of Surgery, Clinician Educator

William D. Binder
Associate Professor of Emergency Medicine, Clinician Educator

Alice E. Bonitati
Associate Professor of Medicine, Clinician Educator

Tanya Booker
Associate Professor of Obstetrics and Gynecology, Clinician Educator

Jeffrey M. Brody
Associate Professor of Diagnostic Imaging, Clinician Educator

William D. Brown
Associate Professor of Pediatrics, Clinician Educator; Associate Professor of Neurology, Clinician Educator

Robert T. Burke
Associate Professor Emeritus of Pediatrics, Clinician Educator

Kate E. Cahill
Associate Professor of Medicine, Clinician Educator; Associate Professor of Medical Science, Clinician Educator

Mariana Canepa
Associate Professor of Pathology and Laboratory Medicine, Clinician Educator

Geoffrey Capraro
Associate Professor of Emergency Medicine, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Alexios Carayannopoulos
Associate Professor of Neurosurgery, Clinician Educator

Cassandra L. Carberry
Associate Professor of Obstetrics and Gynecology, Clinician Educator

Gene A. Cardarelli
Associate Professor of Radiation Oncology, Clinician Educator

Gerardo P. Carino
Associate Professor of Medicine, Clinician Educator

Thomas N. Carruthers
Associate Professor of Surgery, Clinician Educator

Rabin F. Chandran
Associate Professor of Family Medicine, Clinician Educator

Prabjot Channa
Associate Professor of Surgery, Clinician Educator

Laura L. Chapman
Associate Professor of Emergency Medicine, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Kenneth K. L. Chen
Associate Professor of Medicine, Clinician Educator; Associate Professor of Obstetrics and Gynecology, Clinician Educator

Wendy Shih-wen Chen
Associate Professor of Surgery, Clinician Educator

Erica Chung
Associate Professor of Pediatrics, Clinician Educator

Sybil Cineas
Associate Professor of Medicine, Clinician Educator; Associate Professor of Medical Science, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Sevdenur Cizginer
Associate Professor of Medicine, Clinician Educator; Associate Professor of Health Services, Policy and Practice, Clinician Educator; Associate Professor of Surgery, Clinician Educator

Daniel T. Coghlin
Associate Professor of Pediatrics, Clinician Educator

Jamieson V. Cohn
Associate Professor of Emergency Medicine, Clinician Educator

Erika Constantine
Associate Professor of Emergency Medicine, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Viren A. D'Sa
Associate Professor of Pediatrics, Clinician Educator

Kristy L. Dalrymple Gaudiano
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Jennifer D. Davis
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Bradley D. DeNardo
Associate Professor of Pediatrics, Clinician Educator

Diane DerMarderosian
Associate Professor of Pediatrics, Clinician Educator; Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Abigail Donaldson
Associate Professor of Pediatrics, Clinician Educator

Kathleen R. Donise
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Gregory J. Dubel
Associate Professor of Diagnostic Imaging, Clinician Educator

Christine M. Duffy
Associate Professor of Medicine, Clinician Educator

Annmarie Dunican
Associate Professor of Surgery, Clinician Educator

David Edmonson
Associate Professor of Surgery, Clinician Educator; Associate Professor of Obstetrics and Gynecology, Clinician Educator

Heinrich Elinzano
Associate Professor of Neurology, Clinician Educator; Associate Professor of Medicine, Clinician Educator

Fadya El Rayess
Associate Professor of Family Medicine, Clinician Educator

Matthew A. Esposito
Associate Professor of Obstetrics and Gynecology, Clinician Educator

M. Khurram Faizan
Associate Professor of Pediatrics, Clinician Educator

Katherine E. Faricy-Anderson
Associate Professor of Medicine, Clinician Educator

Deirdre M. Fearon
Associate Professor of Emergency Medicine, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Jeffrey Feden
Associate Professor of Emergency Medicine, Clinician Educator

Mary M. Flynn
Associate Professor of Medicine, Clinician Educator

Sara R. Ford
Associate Professor of Pediatrics, Clinician Educator

Rachel L. Fowler
Associate Professor of Emergency Medicine, Clinician Educator

Ana Tuya Fulton
Associate Professor of Medicine, Clinician Educator; Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Melissa M. Gaitanis
Associate Professor of Medicine, Clinician Educator

Abbe L. Garcia
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Joseph Garland
Associate Professor of Medicine, Clinician Educator

Bethany J. Gentilesco
Associate Professor of Medicine, Clinician Educator; Associate Professor of Medical Science, Clinician Educator

Frantz J. Gibbs
Associate Professor of Emergency Medicine, Clinician Educator

Richard L. Gold
Associate Professor of Diagnostic Imaging, Clinician Educator

Natalia Golova
Associate Professor of Pediatrics, Clinician Educator

Theresa A. Graves
Associate Professor of Surgery, Clinician Educator

Jan C. Groblewski
Associate Professor of Surgery, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Richard Alan Haas
Associate Professor of Diagnostic Imaging, Clinician Educator; Associate Professor of Neurosurgery, Clinician Educator

Fadlallah G. Habr
Associate Professor of Medicine, Clinician Educator

Jesse Hart
Associate Professor of Pathology and Laboratory Medicine, Clinician Educator

Thaddeus W. Herliczek
Associate Professor of Diagnostic Imaging, Clinician Educator

Jennifer Herren
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Michael Herzlinger
Associate Professor of Pediatrics, Clinician Educator

Ross Hilliard
Associate Professor of Medicine, Clinician Educator

Mary M. Hillstrom
Associate Professor of Diagnostic Imaging, Clinician Educator

Linda A. Hunter
Associate Professor of Obstetrics and Gynecology, Clinician Educator

Elias Hyams
Associate Professor of Surgery, Clinician Educator

Elizabeth S. Jacobs
Associate Professor of Emergency Medicine, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Gaurav Jindal
Associate Professor of Diagnostic Imaging, Clinician Educator

Loree Kalliainen
Associate Professor of Surgery, Clinician Educator

Shuba Kamath
Associate Professor of Pediatrics, Clinician Educator

Adib Karam
Associate Professor of Diagnostic Imaging, Clinician Educator

Emily R. Katz
Associate Professor of Psychiatry and Human Behavior, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

John C. Kawaoka
Associate Professor of Dermatology, Clinician Educator

Karen L. Kerman
Associate Professor of Neurology, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Tareq Kheirbek
Associate Professor of Surgery, Clinician Educator

Humera Khurshid
Associate Professor of Medicine, Clinician Educator

Adam Klipfel
Associate Professor of Surgery, Clinician Educator

Michael P. Koster
Associate Professor of Pediatrics, Clinician Educator

Peter K. Kriz
Associate Professor of Orthopaedics, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Chandan N. Lakhiani
Associate Professor of Pediatrics, Clinician Educator

Shoshana M. Landow
Associate Professor of Dermatology, Clinician Educator

Rebecca Laptook
Associate Professor of Psychiatry and Human Behavior, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Andrew T. Levinson
Associate Professor of Medicine, Clinician Educator

David G. Lindquist
Associate Professor of Emergency Medicine, Clinician Educator

Melanie J. Lippmann
Associate Professor of Emergency Medicine, Clinician Educator

Cynthia L. Loncar
Associate Professor of Pediatrics, Clinician Educator; Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Christine M. Low
Associate Professor of Psychiatry and Human Behavior, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Elizabeth A. Lowenhaupt
Associate Professor of Psychiatry and Human Behavior, Clinician Educator; Associate Professor of Medical Science, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Stephanie N. Lueckel
Associate Professor of Surgery, Clinician Educator

Maria C. Mancebo
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Margaret M. Mannix
Associate Professor of Psychiatry and Human Behavior, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Abigail Marcaccio
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Peter J. Mazzaglia
Associate Professor of Surgery, Clinician Educator

Robin L. McKinney
Associate Professor of Pediatrics, Clinician Educator

Lynn Mcnicoll
Associate Professor of Medicine, Clinician Educator

Ryan A. McTaggart
Associate Professor of Diagnostic Imaging, Clinician Educator; Associate Professor of Neurology, Clinician Educator; Associate Professor of Neurosurgery, Clinician Educator

Anthony E. Mega
Associate Professor of Medicine, Clinician Educator; Associate Professor of Surgery, Clinician Educator

Niharika D. Mehta
Associate Professor of Medicine, Clinician Educator

Stephen T. Mernoff
Associate Professor of Neurology, Clinician Educator

Dino Messina
Associate Professor of Medicine, Clinician Educator

Kimberly V. Miller
Associate Professor of Surgery, Clinician Educator

Taro Minami
Associate Professor of Medicine, Clinician Educator; Associate Professor of Medical Science, Clinician Educator

Jennie J. Muglia
Associate Professor of Dermatology, Clinician Educator

Nadia Mujahid
Associate Professor of Medicine, Clinician Educator

Gerard Nau
Associate Professor of Medicine, Clinician Educator

John W. O'Bell
Associate Professor of Medicine, Clinician Educator

Adena Osband
Associate Professor of Surgery, Clinician Educator

Adam D. Pallant
Associate Professor of Pediatrics, Clinician Educator

Salley Pels
Associate Professor of Pediatrics, Clinician Educator

Gita S. Pensa
Associate Professor of Emergency Medicine, Clinician Educator

John A. Pezzullo
Associate Professor of Diagnostic Imaging, Clinician Educator

Michael Picchioni
Associate Professor of Medicine, Clinician Educator

Lee Alexander Polikoff
Associate Professor of Pediatrics, Clinician Educator

N. Stevenson Potter
Associate Professor of Neurology, Clinician Educator; Associate Professor of Neurosurgery, Clinician Educator

Amanda B. Pressman
Associate Professor of Medicine, Clinician Educator

Joseph S. Rabatin
Associate Professor of Medicine, Clinician Educator

Elizabeth Renaud
Associate Professor of Surgery, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Mark S. Ridlen
Associate Professor Emeritus of Diagnostic Imaging, Clinician Educator

Jeffrey S. Riese
Associate Professor of Pediatrics, Clinician Educator

Tanya Rogo
Associate Professor of Pediatrics, Clinician Educator

Leslie A. Roth
Associate Professor of Surgery, Clinician Educator

Kathleen M. Rotondo
Associate Professor of Pediatrics, Clinician Educator

G. Dean Roye
Associate Professor of Surgery, Clinician Educator

Beth A. Ryder
Associate Professor of Surgery, Clinician Educator; Associate Professor of Medical Science, Clinician Educator

Margaret Scheffler
Associate Professor of Pediatrics, Clinician Educator

Scott T. Schmidt
Associate Professor of Surgery, Clinician Educator

Ivona Sediva
Associate Professor of Pediatrics, Clinician Educator

Nishit Shah
Associate Professor of Surgery, Clinician Educator

Linda B. Shalon
Associate Professor of Pediatrics, Clinician Educator

Robert M. Shalvoy
Associate Professor of Orthopaedics, Clinician Educator

Jason M. Shapiro
Associate Professor of Pediatrics, Clinician Educator; Associate Professor of Medicine, Clinician Educator

Afreen Siddiqui
Associate Professor of Anesthesiology, Clinician Educator

Rebecca B. Silver
Associate Professor of Psychiatry and Human Behavior, Clinician Educator; Associate Professor of Pediatrics, Clinician Educator

Victoria Snegovskikh
Associate Professor of Obstetrics and Gynecology, Clinician Educator

Mindy Sobota
Associate Professor of Medicine, Clinician Educator

Peter Soukas
Associate Professor of Medicine, Clinician Educator

Philippa Gabrielle Sprinz
Associate Professor Emerita, Clinician Educator of Pediatrics

Dariusz R. Stachurski
Associate Professor of Pathology and Laboratory Medicine, Clinician Educator

Kara Stavros
Associate Professor of Neurology, Clinician Educator

Andrew H. Stephen
Associate Professor of Surgery, Clinician Educator

Philip H. Stockwell
Associate Professor of Medicine, Clinician Educator

Elizabeth M. Sutton
Associate Professor of Emergency Medicine, Clinician Educator; Associate Professor of Medical Science, Clinician Educator

David W. Swenson
Associate Professor of Diagnostic Imaging, Clinician Educator

Jie Tang
Associate Professor of Medicine, Clinician Educator

Lidia A. Vognar
Associate Professor of Medicine, Clinician Educator

Michael T. Wallach
Associate Professor Emeritus of Diagnostic Imaging, Clinician Educator and Associate Professor Emeritus of Pediatrics, Clinician Educator

Otis U. Warren
Associate Professor of Emergency Medicine, Clinician Educator

Delma-Jean Watts
Associate Professor of Pediatrics, Clinician Educator

Joseph Benjamin Weiss
Associate Professor of Medicine, Clinician Educator

Doreen L. Wiggins
Associate Professor of Surgery, Clinician Educator

Matthew D. Willis
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Oliver Wisco
Associate Professor of Dermatology, Clinician Educator

Kyle J. Wohlrab
Associate Professor of Obstetrics and Gynecology, Clinician Educator

Jennifer C. Wolff
Associate Professor of Psychiatry and Human Behavior, Clinician Educator

Karen L. Woolfall-Quinn
Associate Professor of Medicine, Clinician Educator

Hsi-Yang Wu
Associate Professor of Surgery, Clinician Educator

Ali Yalcindag
Associate Professor of Pediatrics, Clinician Educator

James W. Ziegler
Associate Professor of Pediatrics, Clinician Educator

Assistant Professor, Clinician Educator

Yasin Abul
Assistant Professor of Medicine, Clinician Educator

Yam Acharya
Assistant Professor of Medicine, Clinician Educator

Purva Agarwal
Assistant Professor of Medicine, Clinician Educator

Ameen Al-Aghil
Assistant Professor of Medicine, Clinician Educator

Ana Albuja
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Neurology, Clinician Educator

May Aldaabil
Assistant Professor of Medicine, Clinician Educator

Sibel Algon
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Emily Allen
Assistant Professor of Pediatrics, Clinician Educator

Ryan Allen
Assistant Professor of Medicine, Clinician Educator

Matthew D. Alvin
Assistant Professor of Diagnostic Imaging, Clinician Educator

Laura Margarita Amorese-O'Connell
Assistant Professor of Medicine, Clinician Educator

Jared Anderson
Assistant Professor of Emergency Medicine, Clinician Educator

Kristin L. Anderson
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Medicine, Clinician Educator

Nadeem Anis
Assistant Professor of Medicine, Clinician Educator

Jonathan Rhett Argo
Assistant Professor of Anesthesiology, Clinician Educator

Pinar Arikan
Assistant Professor of Medicine, Clinician Educator

Shyamal R. Asher
Assistant Professor of Anesthesiology, Clinician Educator

David Ashley
Assistant Professor of Family Medicine, Clinician Educator

Nicholas Asselin
Assistant Professor of Emergency Medicine, Clinician Educator

Bassam I. Aswad
Assistant Professor of Pathology and Laboratory Medicine, Clinician Educator

James Atkinson
Assistant Professor of Medicine, Clinician Educator

Eilean Attwood
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Matthew Austin
Assistant Professor of Medicine, Clinician Educator

Gabriella Juliet Avellino
Assistant Professor of Surgery, Clinician Educator

Nina K. Ayala
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Maroun Azar
Assistant Professor of Medicine, Clinician Educator

Tatiana Bakaeva
Assistant Professor of Surgery, Clinician Educator

Paul Baker
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Lauren Elaine Bakios
Assistant Professor of Surgery, Clinician Educator; Assistant Professor of Pathology and Laboratory Medicine, Clinician Educator

Peter Michael Barth
Assistant Professor of Medicine, Clinician Educator

Anneliese Beaubrun
Assistant Professor of Medicine, Clinician Educator

Andrew Charles Beck
Assistant Professor of Emergency Medicine, Clinician Educator

Allison L. Behrle Yardley
Assistant Professor of Pediatrics, Clinician Educator

Santosh Belbase
Assistant Professor of Medicine, Clinician Educator

Elizabeth Benz
Assistant Professor of Surgery, Clinician Educator

Scott E. Benzuly
Assistant Professor of Anesthesiology, Clinician Educator

Kristina Berglund
Assistant Professor of Medicine, Clinician Educator

Justin Berk
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Medicine, Clinician Educator

Meghan Beucher
Assistant Professor of Emergency Medicine, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

Christina A. Beyer
Assistant Professor of Medicine, Clinician Educator

Harikrashna Bhatt
Assistant Professor of Medicine, Clinician Educator

Roma D. Bhuta
Assistant Professor of Pediatrics, Clinician Educator

Courtney C. Bilodeau
Assistant Professor of Medicine, Clinician Educator

Ariel Birnbaum
Assistant Professor of Medicine, Clinician Educator

Alison Block
Assistant Professor of Family Medicine, Clinician Educator

Michael F. Blundin
Assistant Professor of Medicine, Clinician Educator

Timothy John Boardman
Assistant Professor of Emergency Medicine, Clinician Educator

Mark Brady
Assistant Professor of Emergency Medicine, Clinician Educator

Elizabeth Brannan
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Russell Evan Bratman
Assistant Professor of Medicine, Clinician Educator

Rimini Breakstone
Assistant Professor of Medicine, Clinician Educator

Arlen Brickman
Assistant Professor of Pathology and Laboratory Medicine, Clinician Educator

Allison W. Brindle
Assistant Professor of Pediatrics, Clinician Educator

Vanessa M. Britto
Assistant Professor of Medicine, Clinician Educator

Amy L. Brotherton
Assistant Professor of Medicine, Clinician Educator

Benjamin P. Brown
Mimi Pichey Assistant Professor of Obstetrics and Gynecology

Erica Dawn Brown
Assistant Professor of Medicine, Clinician Educator

J. Elliott Brown
Assistant Professor of Diagnostic Imaging, Clinician Educator

Tara Brown
Assistant Professor of Pediatrics, Clinician Educator

William J. Brucker
Assistant Professor of Pediatrics, Clinician Educator

Dayna Burrell
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Joseph M. Caiati
Assistant Professor of Anesthesiology, Clinician Educator

Rachael L. Caiati
Assistant Professor of Diagnostic Imaging, Clinician Educator

Nathan Calabro-Kailukaitis
Assistant Professor of Medicine, Clinician Educator

Joaquin Camara
Assistant Professor of Neurosurgery, Clinician Educator

Tyler Campbell
Assistant Professor of Diagnostic Imaging, Clinician Educator

Madeline Cancian
Assistant Professor of Surgery, Clinician Educator

Katherine F. Canfield
Assistant Professor of Pediatrics, Clinician Educator

Rebecca Watkins Cangemi
Assistant Professor of Medicine, Clinician Educator

Chelsy Caren
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Raquel Carneiro
Assistant Professor of Medicine, Clinician Educator

Bianca M. Carpentier
Assistant Professor of Diagnostic Imaging, Clinician Educator

Brady G. Case
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Sarah Case
Assistant Professor of Emergency Medicine, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

John A. Cassese
Assistant Professor of Diagnostic Imaging, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

Stephanie A. Catanese
Assistant Professor of Medicine, Clinician Educator

Abdullah Chahin
Assistant Professor of Medicine, Clinician Educator

Sandy Chai
Assistant Professor of Dermatology, Clinician Educator

Victoria C. Chang
Assistant Professor of Neurology, Clinician Educator

Wen-I. Chang
Assistant Professor of Pediatrics, Clinician Educator

Heather A. Chapman
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

George T. Charlton
Assistant Professor of Medicine, Clinician Educator

Saima Chaudhry
Assistant Professor of Neurology, Clinician Educator

Tzong-Huei Herbert Chen
Assistant Professor of Anesthesiology, Clinician Educator

Xiaolei Chen
Assistant Professor of Medicine, Clinician Educator

Vicky Cheng
Assistant Professor of Medicine, Clinician Educator

Daniel Horim Choi
Assistant Professor of Pediatrics, Clinician Educator

Pradeep Chopra
Assistant Professor of Medicine, Clinician Educator

Lydia R. Ciarallo
Assistant Professor of Emergency Medicine, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

Seth Alan Clark
Assistant Professor of Medicine, Clinician Educator; Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Alexander D. Cohen
Assistant Professor of Anesthesiology, Clinician Educator

Keith Robert Cohen
Assistant Professor of Medicine, Clinician Educator

Lauren Conner
Assistant Professor of Medicine, Clinician Educator

Michael Connolly
Assistant Professor of Surgery, Clinician Educator; Assistant Professor of Medical Science, Clinician Educator

Celeste Corcoran
Assistant Professor of Pediatrics, Clinician Educator

Patricia A. Cristofaro
Assistant Professor of Medicine, Clinician Educator

Charles Cronin
Assistant Professor of Medicine, Clinician Educator

Alisha Maureen Crowley
Assistant Professor of Medicine, Clinician Educator

David Curley
Assistant Professor of Emergency Medicine, Clinician Educator

Alicia Curtin
Assistant Professor of Family Medicine, Clinician Educator

Stephanie Jo Czech
Assistant Professor of Family Medicine, Clinician Educator; Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Deepan Dalal
Assistant Professor of Medicine, Clinician Educator; Assistant Professor of the Practice of Health Services, Policy and Practice

Kristopher Davignon
Assistant Professor of Anesthesiology, Clinician Educator

Lawrence M. Davis
Assistant Professor of Diagnostic Imaging, Clinician Educator

Julio Cesar Defillo
Assistant Professor of Medicine, Clinician Educator

Linda Rose Demello
Assistant Professor of Diagnostic Imaging, Clinician Educator

Beatriz M. DeMoranville
Assistant Professor of Medicine, Clinician Educator

Almaz S. Dessie
Assistant Professor of Emergency Medicine, Clinician Educator

Carla de Venecia
Assistant Professor of Diagnostic Imaging, Clinician Educator

Kyla Dewar
Assistant Professor of Medicine, Clinician Educator

Zobeida Diaz
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Vincent A. DiMaggio
Assistant Professor of Pediatrics, Clinician Educator

Christopher Dimarco
Assistant Professor of Dermatology, Clinician Educator

Lindsay E. Donat
Assistant Professor of Surgery, Clinician Educator

Linda L. Donegan
Assistant Professor of Diagnostic Imaging, Clinician Educator

Edward Donnelly
Assistant Professor of Neurology, Clinician Educator

Kristina M. Duarte
Assistant Professor of Family Medicine, Clinician Educator

Laura M. Edmonds
Assistant Professor of Medicine, Clinician Educator

Pamela Egan
Assistant Professor of Medicine, Clinician Educator

Renee Ross Eger
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Amy Egolf
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Samuel K. Evans
Assistant Professor of Medicine, Clinician Educator

Ronan Farrell
Assistant Professor of Medicine, Clinician Educator

Hannah Fassel
Assistant Professor of Pediatrics, Clinician Educator

Carol J. Faulkner
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Mariam Fayek
Assistant Professor of Medicine, Clinician Educator

Tamara L. Feingold-Link
Assistant Professor of Medicine, Clinician Educator

Amanda Fernandes
Assistant Professor of Medicine, Clinician Educator

Denise Fernandes
Assistant Professor of Medicine, Clinician Educator

Sean Fine
Assistant Professor of Medicine, Clinician Educator

Arkadiy Finn
Assistant Professor of Medicine, Clinician Educator

Elnaz Firoz
Assistant Professor of Dermatology, Clinician Educator

Whitney Charles Fisher
Assistant Professor of Emergency Medicine, Clinician Educator

Cecilia Crane Fix
Assistant Professor of Medicine, Clinician Educator

Devon Flaherty
Assistant Professor of Anesthesiology, Clinician Educator

Andrew Foderaro
Assistant Professor of Medicine, Clinician Educator

Christine Foley
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Colin Andrew Foley
Assistant Professor of Anesthesiology, Clinician Educator

Meghan Elizabeth Fredette
Assistant Professor of Pediatrics, Clinician Educator

Amy French
Assistant Professor of Medicine, Clinician Educator

Katharine French
Assistant Professor of Medicine, Clinician Educator

Christopher M. Furey
Assistant Professor of Family Medicine, Clinician Educator

Michael Furman
Assistant Professor of Diagnostic Imaging, Clinician Educator

Tanuja Gandhi
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Meghan Charlotte Geary
Assistant Professor of Medicine, Clinician Educator; Assistant Professor of Medical Science, Clinician Educator

Sara Geffert
Assistant Professor of Medicine, Clinician Educator

Alicia Genisca
Assistant Professor of Emergency Medicine, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

Sassine Ghanem
Assistant Professor of Medicine, Clinician Educator

Arnoldas Giedrimas
Assistant Professor of Medicine, Clinician Educator

Jeanette G. Giedrimas
Assistant Professor of Medicine, Clinician Educator

Holly C. Gil
Assistant Professor of Diagnostic Imaging, Clinician Educator

Alexis Gimovsky
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Marcoandrea Giorgi
Assistant Professor of Surgery, Clinician Educator

Sandra Jimenez Giraldo
Assistant Professor of Pediatrics, Clinician Educator

Melissa Gleckel
Assistant Professor of Family Medicine, Clinician Educator

Anupriya Gogne
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Hayley Goldbach
Assistant Professor of Dermatology, Clinician Educator

Vidya Vellore Gopinath
Assistant Professor of Medicine, Clinician Educator

Michelle Gorgone
Assistant Professor of Anesthesiology, Clinician Educator

Brett W. Goudie
Assistant Professor of Pediatrics, Clinician Educator

Dina Gozman
Assistant Professor of Emergency Medicine, Clinician Educator

Nicole L. Grenier
Assistant Professor of Dermatology, Clinician Educator

Jessica L. Guidi
Assistant Professor of Medicine, Clinician Educator

Sameer Gupta
Assistant Professor of Dermatology, Clinician Educator

Dawa Gurung
Assistant Professor of Medicine, Clinician Educator

Manoj Gurung
Assistant Professor of Medicine, Clinician Educator

Dana Guyer
Assistant Professor of Medicine, Clinician Educator

Charles Hackett
Assistant Professor of Family Medicine, Clinician Educator

Sarah E. Hagin
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Jason Halpern
Assistant Professor of Diagnostic Imaging, Clinician Educator

Maureen Hamel
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Fatma Hammad
Assistant Professor of Medicine, Clinician Educator

L. Corey Hanley
Assistant Professor of Pathology and Laboratory Medicine, Clinician Educator; Assistant Professor of Medical Science, Clinician Educator

Gita V. Harappanahally
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Neurology, Clinician Educator

Erica J. Hardy
Assistant Professor of Medicine, Clinician Educator; Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Jeanna Harvey Barnes
Assistant Professor of Diagnostic Imaging, Clinician Educator

Allison Heinly
Assistant Professor of Pediatrics, Clinician Educator

Karl Herman
Assistant Professor of Medicine, Clinician Educator

Renee Hickey
Assistant Professor of Neurology, Clinician Educator

James Barry Higgs
Assistant Professor of Medicine, Clinician Educator

Thomas Hines
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Medical Science, Clinician Educator

Laurie Hoffman
Assistant Professor of Pediatrics, Clinician Educator

Dawn B. Hogan
Assistant Professor of Family Medicine, Clinician Educator

Mary H. Hohenhaus
Assistant Professor of Medicine, Clinician Educator

Sabina Holland
Assistant Professor of Pediatrics, Clinician Educator

Sara E. Holmstrom
Assistant Professor of Emergency Medicine, Clinician Educator

Brian Honeyman
Assistant Professor of Medicine, Clinician Educator

Tao Hong
Assistant Professor of Pathology and Laboratory Medicine, Clinician Educator

Bridget Hottenstein
Assistant Professor of Medicine, Clinician Educator

Andrew Hsu
Assistant Professor of Medicine, Clinician Educator

Richard C. Huard
Assistant Professor of Pathology and Laboratory Medicine, Clinician Educator

Omar N. Hyder
Assistant Professor of Medicine, Clinician Educator

Sarah Marie Hyder
Assistant Professor of Medicine, Clinician Educator

Marsela Hyska-Campbell
Assistant Professor of Diagnostic Imaging, Clinician Educator

Jason D. Iannuccilli
Assistant Professor of Diagnostic Imaging, Clinician Educator

Ann-Marie Idusuyi
Assistant Professor of Medicine, Clinician Educator

Ambreen Ijaz
Assistant Professor of Medicine, Clinician Educator

Fransisca Indraswari
Assistant Professor of Neurology, Clinician Educator

Randall Ingham
Assistant Professor of Medicine, Clinician Educator

Ghada Issa
Assistant Professor of Diagnostic Imaging, Clinician Educator

Joseph A. Izzi
Assistant Professor of Orthopaedics, Clinician Educator

Judy L. Jang
Assistant Professor of Medicine, Clinician Educator

Thomas W. Jean
Assistant Professor of Medicine, Clinician Educator

Ambrish Jha
Assistant Professor of Medicine, Clinician Educator

Atin Jindal
Assistant Professor of Medicine, Clinician Educator

Jennie Elizabeth Johnson
Assistant Professor of Medicine, Clinician Educator

Jan Marian Karczewski
Assistant Professor of Medicine, Clinician Educator

Vania Kasper
Assistant Professor of Pediatrics, Clinician Educator

Syed Hammad Kazmi
Assistant Professor of Medicine, Clinician Educator

Alexis Kearney
Assistant Professor of Emergency Medicine, Clinician Educator

Sarah Elizabeth Keene
Assistant Professor of Emergency Medicine, Clinician Educator

Carrie Leah Kelly
Assistant Professor of Pediatrics, Clinician Educator

Eric Scott Kerns
Assistant Professor of Medicine, Clinician Educator

Mary Keszler
Assistant Professor of Pediatrics, Clinician Educator

Zain Khalid
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Hanan I. Khalil
Assistant Professor of Diagnostic Imaging, Clinician Educator

Hina Khan
Assistant Professor of Medicine, Clinician Educator

Ramu Kharel
Assistant Professor of Emergency Medicine, Clinician Educator

Hussain Khawaja
Assistant Professor of Medicine, Clinician Educator

Joshua Kim
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Dominic Kleinhenz
Assistant Professor of Orthopaedics, Clinician Educator

Matthew A. Kluge
Assistant Professor of Medicine, Clinician Educator

Julie Knutson
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Manpreet Kochhar
Assistant Professor of Pediatrics, Clinician Educator

Susan L. Koelliker
Assistant Professor of Diagnostic Imaging, Clinician Educator

Aravind Kokkirala
Assistant Professor of Medicine, Clinician Educator

Martha Kole White
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Dhimiter Kondili
Assistant Professor of Diagnostic Imaging, Clinician Educator

Nicole Korbly
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Robin A. Kremsdorf
Assistant Professor of Pediatrics, Clinician Educator

Helena Kuhn
Assistant Professor of Dermatology, Clinician Educator

Vincent Angelo LaBarbera
Assistant Professor of Neurology, Clinician Educator

Donald Labbe
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Ashley Lakin
Assistant Professor of Family Medicine, Clinician Educator

Vita Lamberson
Assistant Professor of Pediatrics, Clinician Educator

Alexis Lawrence
Assistant Professor of Emergency Medicine, Clinician Educator

Jasmin Lebastchi
Assistant Professor of Medicine, Clinician Educator

Athene Ka Wing Lee
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Jisoo Lee
Assistant Professor of Medicine, Clinician Educator

John Lee
Assistant Professor of Diagnostic Imaging, Clinician Educator

Janet Lefkowitz
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Emily A. Leveen
Assistant Professor of Medicine, Clinician Educator

William Lewis-de los Angeles
Assistant Professor of Pediatrics, Clinician Educator

Beth A. Logan
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Elizabeth Lokich
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Kristin Lombardi
Assistant Professor of Pediatrics, Clinician Educator

Michael Longo
Assistant Professor of Diagnostic Imaging, Clinician Educator

Matthew Lee Lorenz
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Medicine, Clinician Educator

Andrew Luhrs
Assistant Professor of Surgery, Clinician Educator

Natalia Lukankina
Assistant Professor of Pediatrics, Clinician Educator

Kenneth A. Lynch
Assistant Professor of Surgery, Clinician Educator

Matthew Lynch
Assistant Professor of Medicine, Clinician Educator

Rebecca MacDonell-Yilmaz
Assistant Professor of Medicine, Clinician Educator

Joanna V. MacLean
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator; Assistant Professor of Medicine, Clinician Educator

Nirav Makwana
Assistant Professor of Diagnostic Imaging, Clinician Educator

Christopher Malgieri
Assistant Professor of Anesthesiology, Clinician Educator

Alexis A. Mancini
Assistant Professor of Surgery, Clinician Educator

Seth Aaron Margolis
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Terry Albert Marryshow
Assistant Professor of Medicine, Clinician Educator

Kyle Denison Martin
Assistant Professor of Emergency Medicine, Clinician Educator

Augusto Rafael Martinez
Assistant Professor of Pediatrics, Clinician Educator

Cathy Massoud
Assistant Professor of Dermatology, Clinician Educator

Donnah Mathews
Assistant Professor of Medicine, Clinician Educator

Parijat Mathur
Assistant Professor of Medicine, Clinician Educator

Amy Marie Matson
Assistant Professor of Emergency Medicine, Clinician Educator

Mirabelle Mattar
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Kate Mayans
Assistant Professor of Medicine, Clinician Educator

Kristina McAteer
Assistant Professor of Emergency Medicine, Clinician Educator

Joseph M. McNamara
Assistant Professor of Pediatrics, Clinician Educator

Anupama Menon
Assistant Professor of Medicine, Clinician Educator

Virginia Mensah
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Laura Y. Mercurio
Assistant Professor of Emergency Medicine, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

Basma Merhi
Assistant Professor of Medicine, Clinician Educator

Stephen J. Migliori
Assistant Professor of Surgery, Clinician Educator

Brian Mikolasko
Assistant Professor of Medicine, Clinician Educator

Katherine Gallagher Miller
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Kennon S. Miller
Assistant Professor of Surgery, Clinician Educator

Kate Millington
Assistant Professor of Pediatrics, Clinician Educator

Mary Minn
Assistant Professor of Anesthesiology, Clinician Educator

Abhinav Misra
Assistant Professor of Medicine, Clinician Educator

Srilakshmi Mitta
Assistant Professor of Medicine, Clinician Educator

Gabrielle Mock
Assistant Professor of Medicine, Clinician Educator

Sameh Mohamed
Assistant Professor of Medicine, Clinician Educator

Krisztina Moldovan
Assistant Professor of Neurosurgery, Clinician Educator

Debra J. Moorhead
Assistant Professor of Family Medicine, Clinician Educator

Carla C. Moreira
Assistant Professor of Surgery, Clinician Educator

Jonathan S. Movson
Assistant Professor of Diagnostic Imaging, Clinician Educator

Mahmoud Mowafy
Assistant Professor of Medicine, Clinician Educator

Christopher Mullin
Assistant Professor of Medicine, Clinician Educator

Benjamin John Murphy
Assistant Professor of Medicine, Clinician Educator

Carolyn Frances Murphy
Assistant Professor of Medicine, Clinician Educator

Matthew Murphy
Assistant Professor of Medicine, Clinician Educator; Assistant Professor of Behavioral and Social Sciences

Arvind S. Murthy
Assistant Professor of Anesthesiology, Clinician Educator

Yayra Musabek
Assistant Professor of Medicine, Clinician Educator

Nicholas Musisca
Assistant Professor of Emergency Medicine, Clinician Educator

Drew Nagle
Assistant Professor of Medicine, Clinician Educator

Syed Naqvi
Assistant Professor of Medicine, Clinician Educator

Carolynn Joy Nassar
Assistant Professor of Medicine, Clinician Educator

David P. Neumann
Assistant Professor of Diagnostic Imaging, Clinician Educator

Iva Neupane
Assistant Professor of Medicine, Clinician Educator

Christine Nevins-Herbert
Assistant Professor of Medicine, Clinician Educator

Rebecca Newland Kilch
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Van T. Nguyen
Assistant Professor of Diagnostic Imaging, Clinician Educator

Duyu Nie
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Neurology, Clinician Educator; Assistant Professor of Neurosurgery, Clinician Educator

Rabin Niroula
Assistant Professor of Medicine, Clinician Educator

Ashlyn Morgan Norris
Assistant Professor of Medicine, Clinician Educator

Jonathan David Nyce
Assistant Professor of Pediatrics, Clinician Educator

Jennifer A. O'Brien
Assistant Professor of Medicine, Clinician Educator

Jill A. O'Brien
Assistant Professor of Medicine, Clinician Educator

Erin Elizabeth O'Connor
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

John Oghene
Assistant Professor of Medicine, Clinician Educator

Olutosin Ojugbele
Assistant Professor of Pediatrics, Clinician Educator

Nnenna Okpara
Assistant Professor of Medicine, Clinician Educator

Susan P. Oliverio
Assistant Professor of Medicine, Clinician Educator

Mercedes Pacheco
Assistant Professor of Medicine, Clinician Educator

William Page
Assistant Professor of Medicine, Clinician Educator

Jayram M. Pai
Assistant Professor of Emergency Medicine, Clinician Educator

Christopher D. Palmer
Assistant Professor of Medicine, Clinician Educator

Lynne M. Palmisciano
Assistant Professor of Emergency Medicine, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

Diane M. Parente
Assistant Professor of Medicine, Clinician Educator

Rachit Patil
Assistant Professor of Pediatrics, Clinician Educator; Assistant Professor of Neurology, Clinician Educator; Assistant Professor of Neurosurgery, Clinician Educator

Johann Patlak
Assistant Professor of Anesthesiology, Clinician Educator

Jamie Patterson
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Leland Kit Perice
Assistant Professor of Emergency Medicine, Clinician Educator

Gianna Petrone
Assistant Professor of Emergency Medicine, Clinician Educator

Daniel Philbin
Assistant Professor of Medicine, Clinician Educator

Catherine Piccoli
Assistant Professor of Diagnostic Imaging, Clinician Educator

Jessica Pineda
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator; Assistant Professor of Family Medicine, Clinician Educator

Stephan Peter Pirnie
Assistant Professor of Neurology, Clinician Educator

Brea Prindaville
Assistant Professor of Dermatology, Clinician Educator

Michael Grier Prucha
Assistant Professor of Emergency Medicine, Clinician Educator

Hassan Qadir
Assistant Professor of Medicine, Clinician Educator

Matthew I. Quesenberry
Assistant Professor of Medicine, Clinician Educator

Reema Qureshi
Assistant Professor of Medicine, Clinician Educator

Gregory Rachu
Assistant Professor of Medicine, Clinician Educator

Tharani Rajeswaran
Assistant Professor of Medicine, Clinician Educator

Shruthi Ram
Assistant Professor of Diagnostic Imaging, Clinician Educator

Vendhan Ramanujam
Assistant Professor of Anesthesiology, Clinician Educator

Vinay Rao
Assistant Professor of Medicine, Clinician Educator

Fatima Raza
Assistant Professor of Medicine, Clinician Educator

Sakeena Raza
Assistant Professor of Medicine, Clinician Educator

Matthew Reuter
Assistant Professor of Neurology, Clinician Educator

Thomas Reznik
Assistant Professor of Medicine, Clinician Educator

Iole Ribizzi-Akhtar
Assistant Professor of Medicine, Clinician Educator

Jason Richards
Assistant Professor of Neurology, Clinician Educator

Katherine Richman
Assistant Professor of Medicine, Clinician Educator

Meghan C. Riddle
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Jennifer Ritzau
Assistant Professor of Medicine, Clinician Educator

David Rivera
Assistant Professor of Surgery, Clinician Educator

Zachary Robbins
Assistant Professor of Anesthesiology, Clinician Educator

Lee Adam Robinson
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Jael Rodriguez
Assistant Professor of Medicine, Clinician Educator

Jamison E. Rogers
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Jennifer Rogers
Assistant Professor of Emergency Medicine, Clinician Educator

Ralph Rogers
Assistant Professor of Medicine, Clinician Educator

Bani Roland
Assistant Professor of Medicine, Clinician Educator

Fred A. Rotenberg
Assistant Professor of Anesthesiology, Clinician Educator

Amity Rubeor
Assistant Professor of Family Medicine, Clinician Educator

Monica Osbelia Ruiz
Assistant Professor of Pediatrics, Clinician Educator

Kyle Rumery
Assistant Professor of Medicine, Clinician Educator

Abbas Rupawala
Assistant Professor of Medicine, Clinician Educator

Jack C. Rusley
Assistant Professor of Pediatrics, Clinician Educator; Clinical Assistant Professor of Health Services, Policy and Practice

Natasha R. Rybak
Assistant Professor of Medicine, Clinician Educator

Heather Rybasack-Smith
Assistant Professor of Emergency Medicine, Clinician Educator

Celine Saade
Assistant Professor of Surgery, Clinician Educator

Noha Sadek
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Bachir Sakr
Assistant Professor of Medicine, Clinician Educator; Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Ibrahem Salloum
Assistant Professor of Medicine, Clinician Educator

Mark V. Salmon
Assistant Professor of Medicine, Clinician Educator

Cassandra Sams
Assistant Professor of Diagnostic Imaging, Clinician Educator

Martha Cristina Sanchez
Assistant Professor of Medicine, Clinician Educator

Michael Santos
Assistant Professor of Medicine, Clinician Educator

Sylvester Sarfraz
Assistant Professor of Medicine, Clinician Educator

Brenda Satterthwaite
Assistant Professor of Anesthesiology, Clinician Educator

May-Tal Sauerbrun-Cutler
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Alessandra Sax
Assistant Professor of Diagnostic Imaging, Clinician Educator

Albert A. Scappaticci
Assistant Professor of Diagnostic Imaging, Clinician Educator

Jamie Lea Schaefer
Assistant Professor of Surgery, Clinician Educator

Lauren K. Schlanger
Assistant Professor of Medicine, Clinician Educator

Justin Schleifer
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Kevin J. Scully
Assistant Professor of Pediatrics, Clinician Educator

Michael Seidler
Assistant Professor of Diagnostic Imaging, Clinician Educator

Vijairam Selvaraj
Assistant Professor of Medicine, Clinician Educator

Su-jean Seo
Assistant Professor of Dermatology, Clinician Educator

Monica Serrano Gonzalez
Assistant Professor of Pediatrics, Clinician Educator

Shideh Shafie
Assistant Professor of Emergency Medicine, Clinician Educator

Ankur Shah
Assistant Professor of Medicine, Clinician Educator

Daniel J. Shanin
Assistant Professor of Emergency Medicine, Clinician Educator

Laura Shawhughes
Assistant Professor of Medicine, Clinician Educator

Matthew Michael Sheehan
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Mae Shen
Assistant Professor of Medicine, Clinician Educator

Sarah-Kim Shields
Assistant Professor of Diagnostic Imaging, Clinician Educator

Alexei Shimanovsky
Assistant Professor of Medicine, Clinician Educator

Victor H. Shin
Assistant Professor of Medicine, Clinician Educator

Anuoluwapo Shobayo
Assistant Professor of Medicine, Clinician Educator

Nicole Siegert
Assistant Professor of Family Medicine, Clinician Educator

James E. Simmons
Assistant Professor of Medicine, Clinician Educator

Shannon Leigh Simmons
Assistant Professor of Medicine, Clinician Educator

Melissa Ann Simon
Assistant Professor of Surgery, Clinician Educator

Boris Sinayuk
Assistant Professor of Diagnostic Imaging, Clinician Educator

Krish K. Singhal
Assistant Professor of Diagnostic Imaging, Clinician Educator

Michael Sisitsky
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Jeffrey M. Slaiby
Assistant Professor of Surgery, Clinician Educator

Brett Slingsby
Assistant Professor of Pediatrics, Clinician Educator

Heather Smith
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Denis Snegovskikh
Assistant Professor of Anesthesiology, Clinician Educator

David Sobel
Assistant Professor of Surgery, Clinician Educator

Peter Soden
Assistant Professor of Surgery, Clinician Educator

Christopher Song
Assistant Professor of Medicine, Clinician Educator

Maria Soriano-Pisaturo
Assistant Professor of Medicine, Clinician Educator

Claudia Sorin
Assistant Professor of Medicine, Clinician Educator

Lindo Terry Spencer
Assistant Professor of Pediatrics, Clinician Educator

Jensy Stafford
Assistant Professor of Medicine, Clinician Educator; Assistant Professor of Family Medicine, Clinician Educator

Todd S. Stafford
Assistant Professor of Surgery, Clinician Educator

Jami Star
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Mariah Stump
Assistant Professor of Medicine, Clinician Educator

Shova Subedi
Assistant Professor of Pediatrics, Clinician Educator

Edward Suh
Assistant Professor of Anesthesiology, Clinician Educator

Rachel A. Sullivan
Assistant Professor of Surgery, Clinician Educator

Tanya Sutcliffe
Assistant Professor of Emergency Medicine, Clinician Educator; Assistant Professor of Pediatrics, Clinician Educator

Regina Sutskever
Assistant Professor of Medicine, Clinician Educator

Elizabeth Tammaro
Assistant Professor of Family Medicine, Clinician Educator

Gofran Tarabulsi
Assistant Professor of Medicine, Clinician Educator

Geetika Tewary
Assistant Professor of Medicine, Clinician Educator

Jinen Thakkar
Assistant Professor of Medicine, Clinician Educator

Meena Theva
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Molly Thomas
Assistant Professor of Anesthesiology, Clinician Educator

Michelle H. Ting
Assistant Professor of Medicine, Clinician Educator

Tonslyn Toure
Assistant Professor of Medicine, Clinician Educator

Francine Touzard Romo
Assistant Professor of Medicine, Clinician Educator

Gregory J. Towne
Assistant Professor of Anesthesiology, Clinician Educator

M. Pilar Trelles
Assistant Professor of Psychiatry and Human Behavior, Clinician Educator

Lauren Trembley
Assistant Professor of Emergency Medicine, Clinician Educator

Andrew S. Triebwasser
Assistant Professor of Anesthesiology, Clinician Educator

Martha Catherine Trimbur
Assistant Professor of Medicine, Clinician Educator

Margaret Tryforos
Assistant Professor of Family Medicine, Clinician Educator

Elizabeth Vandeventer
Assistant Professor of Diagnostic Imaging, Clinician Educator

Rahul Vanjani
Assistant Professor of Medicine, Clinician Educator

Dhruv Vasant
Assistant Professor of Medicine, Clinician Educator

Robert Y. Velasco
Assistant Professor of Medicine, Clinician Educator

Mauricio Villamar
Assistant Professor of Neurology, Clinician Educator

John S. Vitelli
Assistant Professor of Medicine, Clinician Educator

Stephen Wagner
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Carlos Wambier
Assistant Professor of Dermatology, Clinician Educator

Grace Wanjiku
Assistant Professor of Emergency Medicine, Clinician Educator

Sarah Spencer Welsh
Assistant Professor of Pediatrics, Clinician Educator

Lee V. Wesner
Assistant Professor of Anesthesiology, Clinician Educator

Aaron Paul Wheeler
Assistant Professor of Medicine, Clinician Educator

Claudia Wheeler
Assistant Professor of Neurology, Clinician Educator

Ira Whitten
Assistant Professor of Anesthesiology, Clinician Educator

Mitchell Benjamin Wice
Assistant Professor of Medicine, Clinician Educator

Sabrina M. Witherby
Assistant Professor of Medicine, Clinician Educator

Jared Wortzman
Assistant Professor of Anesthesiology, Clinician Educator

Michael Wu
Assistant Professor of Medicine, Clinician Educator

Pei-Chi Wu
Assistant Professor of Pediatrics, Clinician Educator

Shirley Wu
Assistant Professor of Emergency Medicine, Clinician Educator

Kun Yang
Assistant Professor of Medicine, Clinician Educator

Jeremy Yardley
Assistant Professor of Pediatrics, Clinician Educator

Marian Younge
Assistant Professor of Family Medicine, Clinician Educator

Hadeel Zainah
Assistant Professor of Medicine, Clinician Educator

Kate M. Zaluski
Assistant Professor of Obstetrics and Gynecology, Clinician Educator

Ran Zhang
Assistant Professor of Anesthesiology, Clinician Educator

Yu Zhao
Assistant Professor of Medicine, Clinician Educator

Shenjun Zhu
Assistant Professor of Medicine, Clinician Educator

Jennifer Barrett Zuar
Assistant Professor of Medicine, Clinician Educator

Biology

The Biology concentration invites students to study, in depth and in breadth, the science of life and living matter. Whether pursuing the Bachelor of Arts (A.B.) or Science (Sc.B.) in biology, students can expect to learn broadly in the discipline through a selection of courses in three areas: cell and molecular biology, structure and function, and organismal biology. In addition, students pursuing the Sc.B. complete a thematic track through which they gain an in-depth understanding of a particular subfield such as, Immunopathology, Ecology and Evolutionary Biology, Physiology/Biotechnology, Cell and Molecular Biology, Physical Sciences. The concentration also emphasizes practical skills and experimental design. Concentrators are required to take at least 3 courses with a laboratory or fieldwork component. Within all of these requirements, students have a high degree of flexibility and choice. Broad research opportunities are also available across several departments within the basic sciences as well.

Standard program for the A.B. Biology

The concentration program for the A.B. in Biology consists of four prerequisite courses in math, chemistry, and a statistics course as well as ten courses in biological sciences, including  at least one course in each of the following three areas: Area 1: Cell/Molecular Biology, Area 2: Structure/Function, and Area 3: Organismal Biology.

The Biology A.B. Concentration Worksheet may be a useful tool for course planning.

Prerequisites: 1
Equilibrium, Rate, and Structure
Organic Chemistry I
Single Variable Calculus, Part I (or placement. MATH 0050/MATH 0060 may be substituted for MATH 0090.)
One of the following:
Single Variable Calculus, Part II (or placement)
Single Variable Calculus, Part II (Accelerated) (or equivalent placement)
Or a statistics course, to be approved by the concentration advisor.
Ten Core Courses: 2,3,4
BIOL 0200The Foundation of Living Systems (Required course; AP credit or similar IB or A-levels accepted, placement test available.)1
Area 1 (Cell/Molecular Biology)1
Biochemistry
Genetics
Cell and Molecular Biology
Introductory Microbiology
Principles of Immunology
Biology of the Eukaryotic Cell
Developmental Biology
Conservation in the Genomics Age
21st Century Applications in Cell and Molecular Biology
Toxicology
Principles of Neurobiology
Area 2 (Structure/Function)1
Biological Design: Structural Architecture of Organisms
Invertebrate Zoology
Inquiry in Plant Biology: Analysis of Plant Growth, Reproduction and Adaptive Responses
Principles of Physiology
Biomaterials
Hormones and Behavior
Developmental Biology
Biology of Reproduction
Animal Locomotion
Toxicology
Comparative Biology of the Vertebrates
The Brain: An Introduction to Neuroscience
Area 3 (Organismal Biology)1
Diversity of Life
The Ecology and Evolution of Infectious Disease
Invertebrate Zoology
Principles of Ecology
The Evolution of Plant Diversity
Evolutionary Behavioral Ecology
Evolutionary Biology
Terrestrial Biogeochemistry and the Functioning of Ecosystems
Conservation in the Genomics Age
Animal Locomotion
Comparative Biology of the Vertebrates
Environmental Science in a Changing World
Six additional courses chosen from BIOL and/or NEUR offerings for concentrators. The Core may include up to two related sciences, with advisor approval. The Core must also include a Senior Capstone.6
SENIOR CAPSTONE: *Only apples to students who have declared in Fall 2019 or later.* To be fulfilled via ONE of the following:
1. One of the following approved courses: BIOL 1100, 1250, 1515, 1555, 1565, 1575, 1600, 1820, 1970A.
2. One semester of independent research/independent study (BIOL 1950 or BIOL 1960).
Please visit the BUE webpage for more information.
Total Credits10


Honors: Honors in biology requires a thesis and presentation based on a research project (conducted via BIOL 1950/BIOL 1960), and quality grades in the concentration. Guidelines and information on faculty research are available in the Office of Biology Undergraduate Education or found at http://www.brown.edu/academics/biology/undergraduate-education/.

Standard Program for the Sc.B. Biology

The concentration program for the Sc.B. in Biology consists of seven prerequisite courses in math, chemistry, and physics as well as thirteen to fourteen courses in biological sciences, including courses in each of the following three areas: Area 1: Cell/Molecular Biology, Area 2: Structure/Function, and Area 3: Organismal Biology, and the three-course Track.  The biological sciences requirement also requires research (BIOL 1950/BIOL 1960), which should reflect the advanced cluster.

Students pursuing a ScB in Biology have the option to substitute a course for CHEM 0360 (Organic Chemistry) in their background core. For students pursuing the Marine Biology track, an upper level course in Geological Sciences may replace CHEM 0360. For students pursuing all other tracks, BIOL 0280 (Introductory Biochemistry) may serve as the replacement course.  Please note that approval from the concentration advisor is required for these background course substitutions. If the student has already declared, then a revised concentration plan must be submitted and approved via the ASK system. If BIOL 0280 is used as a substitute for CHEM 0360, it cannot be counted as a core course or as an Area 1 course. Students planning to apply to medical or graduate school should seek additional advising (such as from the Health Careers Office) in crafting their course plan.

The Biology Sc.B. Concentration Worksheet may be a useful tool for course planning.

Prerequisites: 1
Single Variable Calculus, Part I (or placement. MATH 0050/MATH 0060 may be substituted for MATH 0090)
One of the following:
Single Variable Calculus, Part II (or placement)
Single Variable Calculus, Part II (Accelerated)
or a statistics course to be approved by the concentration advisor
Each of the following:
Equilibrium, Rate, and Structure (or IB credit)
Organic Chemistry I
Organic Chemistry II
Biochemistry
Basic Physics A (or equivalent. PHYS 0050 PHYS 0070, or ENGN 0030 may be substituted for PHYS 0030.)
Basic Physics B (or equivalent. PHYS 0060 or ENGN 0040 may be substituted for PHYS 0040.)
Core Courses: 2,3,4
BIOL 0200The Foundation of Living Systems (or placement)1
Area 1 (Cell/Molecular Biology)1
Biochemistry
Genetics
Cell and Molecular Biology
Introductory Microbiology
Principles of Immunology
Biology of the Eukaryotic Cell
Developmental Biology
Conservation in the Genomics Age
21st Century Applications in Cell and Molecular Biology
Toxicology
Principles of Neurobiology
Area 2 (Structure/Function)1
Biological Design: Structural Architecture of Organisms
Invertebrate Zoology
Inquiry in Plant Biology: Analysis of Plant Growth, Reproduction and Adaptive Responses
Principles of Physiology
Biomaterials
Hormones and Behavior
Developmental Biology
Biology of Reproduction
Animal Locomotion
Toxicology
Comparative Biology of the Vertebrates
The Brain: An Introduction to Neuroscience
Area 3 (Organismal Biology)1
Diversity of Life
The Ecology and Evolution of Infectious Disease
Invertebrate Zoology
Principles of Ecology
The Evolution of Plant Diversity
Evolutionary Behavioral Ecology
Evolutionary Biology
Terrestrial Biogeochemistry and the Functioning of Ecosystems
Conservation in the Genomics Age
Animal Locomotion
Comparative Biology of the Vertebrates
Environmental Science in a Changing World
Six additional courses chosen from BIOL and/or NEUR offerings for concentrators. The Core may include up to two related sciences, with advisor approval. The Core must also include research. 46
RESEARCH:
The two semester research requirement may be satisfied by any two of the opportunities listed below. Students may find the following resources useful in planning for research: the annual BUE-PLME Finding, Securing and Succeeding in Research workshop, the BUE Research webpage, and the Sheridan Center’s Undergraduate Research & Experiential Opportunities webpage. Following conversation and agreement with the advisor, students articulate the research plan in the ASK declaration (in the designated text box) which is submitted for review and approval by the advisor.
Choose two:
Directed Research/Independent Study
Directed Research/Independent Study
Independent Study
Independent study course in a related discipline (i.e. STEM disciplines, ENVS, PHP, etc.) if the project is relevant to the student’s learning goals and interests in the concentration.
The following COEX courses: BIOL 0190R, BIOL 0285, BIOL 0440, BIOL 0600, BIOL 0940G, BIOL 1515, BIOL 1555; NEUR 1630, CLPS 1195. New COEX courses will be considered as they are developed and offered at Brown, and as relevant to the concentration.
A summer research experience equivalent in scope and scale to work the student would pursue in a Biology independent study course. Examples include UTRAs, LINK awards, approved research programs at other institutions, etc. These experiences do not count as a course in the 10 course core requirement, but they can be used to satisfy the one semester of the research requirement. Advisors will work with students to review these experiences - drawing on a range of potential materials including a written summary of the experience, formal work plans, materials produced (i.e. presentations/papers), and in some cases a letter from the supporting advisor.
Other equivalent opportunities not listed - with approval from the concentration advisor and Dean Smith
Students are encouraged to pursue research related to their track
TRACK:
The advanced thematic track consists of three additional biological sciences courses (not including BIOL 1950/1960 research) that form a Track. Tracks include: Immuno/Pathobiology; Ecology and Evolutionary Biology; Physiology and Biotechnology; Neurobiology; Physical Sciences; Marine Biology; Cell and Molecular Biology; Biomedical Informatics. At least two track courses, and preferably all three, must be above 1000-level. Track courses should form a cohesive grouping approved by the concentration advisor.3
Biomedical Informatics - BIOL 1565 is required for this track along with 2 additional courses from the following:
Methods in Informatics and Data Science for Health
Evaluation of Health Information Systems
Artificial Intelligence in Biomedicine
Cell and Molecular Biology
Biology of the Eukaryotic Cell
Advanced Biochemistry
Biomolecular Interactions: Health, Disease and Drug Design
Developmental Biology
Biology of Reproduction
Molecular Genetics
Human Genetics and Genomics
21st Century Applications in Cell and Molecular Biology
Toxicology
Stem Cell Biology
Ecology and Evolutionary Biology
Experimental Design in Ecology
Foundations of Population Genetics
Computational Methods for Studying Demographic History with Molecular Data
Marine Biology
Community Ecology
Human Population Genomics
Conservation Biology
Biogeography
Terrestrial Biogeochemistry and the Functioning of Ecosystems
500 Million Years of Land Plants
Conservation in the Genomics Age
Human Genetics and Genomics
Animal Locomotion
Comparative Biology of the Vertebrates
Immunobiology
Host-microbiome Interactions in Health and Disease
Cancer Biology
Innate Immunity
Parasitism: Biology and Disease
Virology
Development of Vaccines to Infectious Diseases
Marine Biology
Marine Biology
GEOL (EEPS) listings 1000 level or above. Must be a coherent set of courses that are above the introductory level and approved by advisor
Neurobiology
Cell Physiology and Biophysics
Topics in Signal Transduction
Synaptic Transmission and Plasticity
Physiological Pharmacology
NEUR listings 1000 level or above
Physiology and Biotechnology
Biotechnology and Global Health
Polymer Science for Biomaterials
Cell Physiology and Biophysics
Topics in Signal Transduction
Biomaterials
Tissue Engineering
Stem Cell Engineering
Principles of Exercise Physiology
Synaptic Transmission and Plasticity
Physiological Pharmacology
Biomolecular Interactions: Health, Disease and Drug Design
Physical Sciences
Must be a coherent set of courses drawn from the Physical Sciences; courses must be above the introductory level and approved by advisor
Total Credits13

Honors: Honors in biology requires a thesis and presentation based on a research project (usually conducted via BIOL 1950/BIOL 1960), and quality grades in the concentration. Guidelines and information on faculty research are available in the Office of Biology Undergraduate Education or at http://www.brown.edu/academics/biology/undergraduate-education/.

Stipulations for Biology Programs:

  1. For double concentrations, no more than two courses may overlap (i.e., be used to meet requirements of both programs). This includes prerequisite courses.
  2. No more than two semesters of directed research may be used as concentration credits. Each does count as an individual core course towards the program, but only carry one lab credit towards the three required.
  3. A limited number of transfer or study abroad courses may be used within the program, subject to approval of advisor, and Associate Dean of Biology, Katherine Smith.

Health & Human Biology

Health and Human Biology is an interdisciplinary concentration that provides a rigorous foundation in the biological sciences with substantive course work in humanities and social sciences within a subfield of Human Health and Disease. The program includes: background courses, biology core courses, a set of theme courses, and a Senior Capstone activity. Background courses provide the essential foundations in chemistry, mathematics, methods, and basic biology. These support the Biology core, which is comprised of a flexible menu of intermediate and advanced courses. A required portion of the Biology core is Genetics, a cornerstone of human biology and its interface with other fields. The Biology core underscores the related coursework within the Health and Disease Theme. The Theme courses are social science and humanities courses that form a cohesive, thoughtful grouping. Theme groupings must be approved by the advisor. A required senior capstone course or activity builds on the program's focus.

 

Program Requirements

REQUIRED BACKGROUND:
Four (4) courses including:
MATH 0090Single Variable Calculus, Part I (or equivalent placement)1
OR
Analytic Geometry and Calculus
and Analytic Geometry and Calculus
OR
Single Variable Calculus, Part II
Single Variable Calculus, Part II (Accelerated)
CHEM 0330Equilibrium, Rate, and Structure1
BIOL 0200The Foundation of Living Systems1
Statistics course chosen with advisor's help.1
CORE PROGRAM:
In addition to the stated background in Chemistry, Math, Biology and Statistics, five (5) Biology plus four (4) coherently-grouped Theme courses, plus a Senior-Year Capstone course or project. (See description of Capstone at link below this table).
BIOLOGY:
Five (5) courses, including:5
Genetics, which can be fulfilled in the following ways:
Genetics
-OR-
Evolutionary Biology
and Cell and Molecular Biology
-OR-
Evolutionary Biology
and Introductory Microbiology
-OR-
Evolutionary Biology
and Biochemistry
Select one course in structure/function/development such as:
Biological Design: Structural Architecture of Organisms
Principles of Physiology
Developmental Biology
Animal Locomotion
Comparative Biology of the Vertebrates
The Brain: An Introduction to Neuroscience
One course in organismal/population biology such as:
The Ecology and Evolution of Infectious Disease
Invertebrate Zoology
Principles of Ecology
Evolutionary Biology
Conservation Biology
Methods in Informatics and Data Science for Health
Comparative Biology of the Vertebrates
Environmental Science in a Changing World
Or a course from the NEUR 1940 series
Two Biology or Neuroscience courses. At least one must be at the advanced level.
THEME: With the advisor's assistance, a theme is chosen and a cohesive set of courses are selected from outside of Biology and Neuroscience.4
Approved courses must be above the introductory level and at least one must be 1000-level or above
No more than TWO courses from a given department may be included in the theme portion
Students will then select from FOUR theme options: 1) Mind, Brain, Behavior, 2) Planetary Health, 3) Global Health, 4) Social Determinants of Health
SENIOR CAPSTONE ACTIVITY: Must be conducted during the senior year, fulfilled by one of the following, and related to the students learning goals in the concentration:1
1) Advisor approved senior seminar or advanced course related to the theme
2) One semester of independent research/independent study (BIOL 1950 or BIOL 1960); in the case of a senior honors thesis, both BIOL 1950 and BIOL 1960 can be used as the capstone.
3) An appropriate internship with a scholarly context can be used if coupled with a semester of independent study mentored by a Brown faculty member.
Total Credits14

CAPSTONE: See https://www.brown.edu/academics/biology/undergraduate-education/undergraduate/health-and-human-biology for more information on the Capstone Activity.

HONORS:  See more information about Honors at https://www.brown.edu/academics/biology/undergraduate-education/honors-biological-sciences.

Applied Mathematics-Biology

The Applied Math - Biology concentration recognizes that mathematics is essential to address many modern biological problems in the post genomic era. Specifically, high throughput technologies have rendered vast new biological data sets that require novel analytical skills for the most basic analyses. These technologies are spawning a new "data-driven" paradigm in the biological sciences and the fields of bioinformatics and systems biology. The foundations of these new fields are inherently mathematical, with a focus on probability, statistical inference, and systems dynamics. These mathematical methods apply very broadly in many biological fields including some like population growth, spread of disease, that predate the genomics revolution. Nevertheless, the application of these methods in areas of biology from molecular genetics to evolutionary biology has grown very rapidly in with the availability of vast amounts of genomic sequence data. Required coursework in this program aims at ensuring expertise in mathematical and statistical sciences, and their application in biology. The students will focus in particular areas of biology. The program culminates in a senior capstone experience that pairs student and faculty in creative research collaborations. 

 

Standard program for the Sc.B. degree

Required coursework in this program aims at ensuring expertise in mathematical and statistical sciences, and their application in biology. The students will focus in particular areas of biology. The program culminates in a senior capstone experience that pairs student and faculty in creative research collaborations. Applied Math – Biology concentrators are prepared for careers in medicine, public health, industry and academic research.

Required Courses:

Students are required to take all of the following courses.

MATH 0090Single Variable Calculus, Part I1
MATH 0100Single Variable Calculus, Part II1
or MATH 0170 Single Variable Calculus, Part II (Accelerated)
MATH 0180Multivariable Calculus (or equivalent placement)1
MATH 0520Linear Algebra1
or MATH 0540 Linear Algebra With Theory
CHEM 0330Equilibrium, Rate, and Structure 11
PHYS 0030Basic Physics A1
or PHYS 0050 Foundations of Mechanics
Select one of the following sequences:2
Applied Ordinary Differential Equations
and Applied Partial Differential Equations I
Methods of Applied Mathematics I
and Methods of Applied Mathematics II
APMA 1650Statistical Inference I1
or APMA 1655 Honors Statistical Inference I
APMA 1070Quantitative Models of Biological Systems1
APMA 1080Inference in Genomics and Molecular Biology1
or NEUR 2110 Statistical Neuroscience
BIOL 0200The Foundation of Living Systems (or equivalent)1
Additional Courses
One additional course in Applied Math or Biology1
We strongly recommend that Applied Mathematics-Biology concentrators take one of the programming courses on or before their first semester as a concentrator. Those who do can use it to satisfy this requirement:
Introduction to Scientific Computing
Introduction to Scientific Computing and Problem Solving
Computing Foundations: Data
Introduction to Object-Oriented Programming and Computer Science
Computer Science: An Integrated Introduction
Accelerated Introduction to Computer Science
Introduction to programming
One research-related course in Applied Math or Biology. For example:1
A senior seminar course from the APMA 193X, 194X series
A directed research/independent study course from the APMA 1970, 1971, or BIOL 1950, 1960, or NEUR 1970 series
A directed research/independent study course in a related discipline (i.e. STEM disciplines, ENVS, PHP, etc.) if the project is relevant to the student's learning goals in the concentration and with approval from the concentration advisor.
A pre-approved course that satisfies the Biology AB capstone requirement: BIOL 1100, 1250, 1515, 1555, 1565, 1575, 1600, 1820, 1970.
A relevant CURE (Course-based Undergraduate Research Experience) course: BIOL 0190R, 0190S, 0285, 0440, 0600, 0940G, 1515, 1555; NEUR 1630, CLPS 1195.
A non-research course related to the concentration along with a research experience equivalent in scope and scale to work the student would pursue in an Applied Math or Biology independent study course. Examples include UTRAs, LINK awards, research programs at other institutions, etc. This requires approval from the concentration advisor and appropriate documentation.
Other equivalent opportunities not listed - with approval from the concentration advisor.
Four classes in the biological sciences agreed upon by the student and advisor. These four courses should form a cohesive grouping in a specific area of emphasis, at least two of which should be at the 1000-level. Some example groupings are below:4
Areas of Emphasis and Suggested Courses:
Some areas of possible emphasis for focusing of elective courses are listed below. Given the large number of course offerings in the biosciences and neuroscience, students are free to explore classes in these areas that are not listed below. However, all classes must be approved by the concentration advisor. APMA 1910 cannot be used as an elective.
Biochemistry
Biochemistry
Advanced Biochemistry
Organic Chemistry I
Chemical Biology
Biotechnology and Physiology
Principles of Physiology
Cell Physiology and Biophysics
and/or appropriate bioengineering courses, such as:
Polymer Science for Biomaterials
Biomaterials
Tissue Engineering
Stem Cell Engineering
Synthetic Biological Systems
Ecology, Evolution, and Genetics
Invertebrate Zoology
and Evolutionary Biology
Principles of Ecology
and The Evolution of Plant Diversity
Genetics
Experimental Design in Ecology
Foundations of Population Genetics
Human Population Genomics
Molecular Genetics
Neuroscience
Mathematical Methods in the Brain Sciences
Neurosciences courses: See https://www.brown.edu/academics/neuroscience/undergraduate/neuroscience-concentration-requirements
Cell Physiology and Biophysics
Topics in Signal Transduction
Synaptic Transmission and Plasticity
Total Credits18

Professional Tracks

The requirements for the professional tracks include all those of each of the standard tracks, as well as the following:

Students must complete full-time professional experiences doing work that is related to their concentration programs, totaling 2-6 months, whereby each internship must be at least one month in duration in cases where students choose to do more than one internship experience.  Such work is normally done at a company, but may also be at a university under the supervision of a faculty member. Internships that take place between the end of the fall and the start of the spring semesters cannot be used to fulfill this requirement.  On completion of each professional experience, the student must write and upload to ASK a reflective essay about the experience, to be approved by the student's concentration advisor. 

On completion of each professional experience, the student must write and upload to ASK a reflective essay about the experience, to be approved by the student's concentration advisor:

  • Which courses were put to use in your summer's work?  Which topics, in particular, were important?

  • In retrospect, which courses should you have taken before embarking on your summer experience?  What are the topics from these courses that would have helped you over the summer if you had been more familiar with them?

  • Are there topics you should have been familiar with in preparation for your summer experience, but are not taught at Brown?  What are these topics?

  • What did you learn from the experience that probably could not have been picked up from course work?

  • Is the sort of work you did over the summer something you would like to continue doing once you graduate? Explain.

  • Would you recommend your summer experience to other Brown students? Explain.

Honors

Requirements and Process: Honors in the Applied Math-Biology concentration is based primarily upon an in-depth, original research project carried out under the guidance of a Brown (and usually Applied Math or BioMed) affiliated faculty advisor.  Projects must be conducted for no less than two full semesters, and students must register for two semesters of credit for the project via APMA 1970 or BIOL 1950/BIOL 1960 or similar independent study courses.  One of these courses can be used to fulfill the research-related course requirement, but the other cannot be used elsewhere in the concentration.  The project culminates in the writing of a thesis which is reviewed by the thesis advisor and a second reader.  It is essential that the student have one advisor from the biological sciences and one in Applied Mathematics.  The thesis work must be presented in the form of an oral presentation (arranged with the primary thesis advisor) or posted at the annual Undergraduate Research Day in either Applied Mathematics or Biology.  For information on registering for BIOL 1950/BIOL 1960, please see https://www.brown.edu/academics/biology/undergraduate-education/undergraduate-research

The concentrations in Applied Math (including joint concentrations) require that honors students demonstrate excellence in grades for courses in the concentration. Students must have earned grades of A or S-with-distinction in at least 70% of the courses used for concentration credit, excluding calculus and linear algebra, or be in the upper 20% of the student's cohort (as measured by the fraction of grades of A or S-with-distinction among courses used for concentration credit, excluding calculus and linear algebra). Since S with distinctions do not appear on the internal academic record or the official transcript, the department will consult directly with the Registrar’s Office to confirm a student’s grades in concentration courses. Additional guidelines and requirements for honors are published on the department website

The deadline for applying to graduate with honors in the concentration are the same as those of the biology concentrations.  However, students in the joint concentration must inform the undergraduate chair in Applied Mathematics of their intention to apply for honors by these dates.

Biochemistry & Molecular Biology

How does life work at the molecular level? This question is at the core of the concentration program Biochemistry and Molecular Biology. In earlier years of this discipline, the focus was on structure and function of proteins, nucleic acids, lipids, carbohydrates and small molecules such as vitamins. Today the logical approach and tools of biochemical science are being expanded to new areas in neuroscience, developmental biology, immunology, pharmacology and synthetic biology (the design of analogs of biological systems). Training in biochemistry begins with a foundation in mathematics, physics, chemistry and biology. Some courses offered in other departments, including engineering, geology and computer science, are also useful. A key component of this program is the year of hands-on research carried out in collaboration with a faculty member here at Brown. Faculty sponsors are drawn from both the Chemistry Department and the Division of Biology and Medicine, and include basic science and clinical faculty.

 

Standard program for the Sc.B. degree

Students must take twenty courses in biology, chemistry, mathematics, and physics, including the following core requirements, some of these may be fulfilled with AP credits.

Three courses in mathematics including two courses in MATH 0090/0100 or MATH 0170/0180 with a third class in statistics, math, or computer science 13
Two courses in physics, typically: 12
Basic Physics A
Foundations of Mechanics
Introduction to Engineering
Basic Physics B
Foundations of Electromagnetism and Modern Physics
Dynamics and Vibrations
Three courses in physical and organic chemistry:3
Equilibrium, Rate, and Structure
Organic Chemistry I
One course in biophysical chemistry:1
Biophysical and Bioinorganic Chemistry
Four courses in biochemistry:4
Biochemistry
Inquiry in Biochemistry: From Gene to Protein Function
Plus two of three upper level biochemistry courses:
Advanced Biochemistry
Chemical Biology
Biochemistry
The two semester research requirement may be satisfied by any two of the following. Students should discuss alternative arrangements or special situations directly with their concentration advisor to obtain prior approval.2
Directed Research/Independent Study
Directed Research/Independent Study
Undergraduate Research
Undergraduate Research
Select biology or chemistry COEX courses (BIOL 0190R, BIOL 0190S, BIOL 0440, BIOL 0600, BIOL 0940G, CHEM 0500)
A summer research experience with faculty in Biology or Chemistry at Brown equivalent or greater in scope and scale to work the student would pursue in a Biology or Chemistry independent study course to satisfy one semester of the research requirement.
Suggested Elective Courses:
Students are required to take five courses from the chart below or, with approval from a concentration advisor, from any science or mathematics course relevant to biochemistry, cell and molecular biology. 5
Applied Mathematics Electives:
Methods of Applied Mathematics I
Mathematical Methods in the Brain Sciences
Essential Statistics
Biology Electives:
Principles of Nutrition
Techniques in Regenerative Medicine: Cells, Scaffolds and Staining
Biotechnology in Medicine
Phage Hunters, Part I
Phage Hunters, Part II
The Foundation of Living Systems
The Ecology and Evolution of Infectious Disease
Microbes in the Environment
Inquiry in Plant Biology: Analysis of Plant Growth, Reproduction and Adaptive Responses
Genetics
Cell and Molecular Biology
Introductory Microbiology
Principles of Immunology
Principles of Physiology
Biology of the Eukaryotic Cell
Polymer Science for Biomaterials
Cell Physiology and Biophysics
Topics in Signal Transduction
Biomaterials
Stem Cell Engineering
Protein Biophysics and Structure
Synthetic Biological Systems
Physiological Pharmacology
Cancer Biology
Biomolecular Interactions: Health, Disease and Drug Design
Developmental Biology
Biology of Reproduction
Innate Immunity
Molecular Genetics
Virology
Development of Vaccines to Infectious Diseases
Drug and Gene Delivery
The Biology of Aging
Chemistry Electives:
Inorganic Chemistry
Physical Chemistry: Quantum Chemistry
Physical Chemistry: Thermodynamics and Statistical Mechanics
Computational Tools in Biochemistry and Chemical Biology
Chemical Biology
Biochemistry
Advanced Organic Chemistry
Organic Reactions
Computer Science Electives:
A First Byte of Computer Science
Introduction to Object-Oriented Programming and Computer Science
Introduction to Algorithms and Data Structures
Computer Science: An Integrated Introduction
Computer Science: An Integrated Introduction
Computational Molecular Biology
Engineering Electives:
Materials Science
Neuroscience Electives: 2
The Brain: An Introduction to Neuroscience
Biology of Hearing
Principles of Neurobiology
Neural Systems
Introduction to Neurogenetics
Neuropharmacology and Synaptic Transmission
The Diseased Brain: Mechanisms of Neurological and Psychiatric Disorders
Physics Electives:
Introduction to Relativity, Waves and Quantum Physics
Public Health Electives:
Essentials of Data Analysis
Total Credits20

Honors Requirements for Biochemistry

All ScB Biochemistry concentrators are candidates for Honors; no separate application is necessary.

The requirements for Honors in Biochemistry are:

* Students must have a majority of either As or S with distinction grades in concentration courses. 

* Two semesters of Independent Study (CHEM 0970, CHEM 0980 or BIOL 1950, BIOL 1960). Guidelines and requirements associated with Independent Study are in the Undergraduate Concentration Handbook which can be found at the department website.

* A Thesis in a form approved by the research advisor, and recommended by the research advisor.  Additional information about thesis guidelines will be provided by the Concentration Advisor in the first half of the fall semester.

Biomedical Engineering

The Sc.B. program in Biomedical Engineering is accredited by the Engineering Accreditation Commission of ABET http://www.abet.org/. It is jointly offered by the School of Engineering and the Division of Biology and Medicine as an interdisciplinary concentration designed for students interested in applying the methods and tools of engineering to the subject matter of biology and the life sciences. Alumni of the Biomedical Engineering (BME) program will achieve one or more of these program educational objectives (PEOs) within five (5) years of graduation: (1) Serve society through work or advanced study in a broad range of fields including, but not limited to, medicine, healthcare, industry, government, and academia; (2) Apply their deeply creative and versatile biomedical engineering education to solve a broad spectrum of engineering and societal challenges; and (3) Contribute as role models, mentors, or leaders in their fields. The student outcomes of this program are the ABET (1) - (7) Student Outcomes as defined by the ABET Criteria for Accrediting Engineering Programs (available online at http://www.abet.org/accreditation-criteria-policies-documents/). The Biomedical Engineering concentration shares much of the core with the other engineering programs, but is structured to include more courses in biology and chemistry, and a somewhat different emphasis in mathematics. 

The requirements regarding Mathematics, Advanced Placement, Transfer Credit, Substitutions for Required Courses, and Humanities and Social Science Courses are identical to those of the Sc.B. degree programs in Engineering. Please refer to the Engineering section of the University Bulletin for explicit guidelines.  

The Biomedical Engineering concentration shares much of the core with the other engineering programs, but is structured to include more courses in biology and chemistry, and a somewhat different emphasis in mathematics. 

 Standard program for the Sc.B. degree

1. Core Courses
ENGN 0030Introduction to Engineering1
or ENGN 0031 Honors Introduction to Engineering
ENGN 0040Dynamics and Vibrations1
ENGN 0510Electricity and Magnetism1
or ENGN 0520 Electrical Circuits and Signals
ENGN 0720Thermodynamics1
ENGN 0810Fluid Mechanics1
CHEM 0330Equilibrium, Rate, and Structure1
MATH 0190Single Variable Calculus, Part II (Physics/Engineering)1
or MATH 0170 Single Variable Calculus, Part II (Accelerated)
or MATH 0100 Single Variable Calculus, Part II
CHEM 0350Organic Chemistry I1
MATH 0200Multivariable Calculus (Physics/Engineering)1
or MATH 0180 Multivariable Calculus
or MATH 0350 Multivariable Calculus With Theory
APMA 0350Applied Ordinary Differential Equations 11
APMA 1650Statistical Inference I1
or BIOL 0495 Statistical Analysis of Biological Data
or PHP 1510 Principles of Biostatistics and Data Analysis
or APMA 1655 Honors Statistical Inference I
2. Upper Level Biomedical Engineering Curriculum
ENGN 1110Transport and Biotransport Processes1
ENGN 1210Biomechanics1
ENGN 1230Instrumentation Design1
ENGN 1490Biomaterials1
BIOL 0800Principles of Physiology1
3. Additional Biomedical Engineering Electives: Complete at least 3 courses from the following groups; other upper-level courses are subject to Concentration Advisor approval.3
Select one or two of the following:
Digital Computing Systems
Neuroengineering
Nanoengineering and Nanomedicine
Cardiovascular Engineering
Recent Advances in Biomedical Engineering
Computer Aided Visualization and Design
Biomedical Optics
Optical Microscopy: Fundamentals and Applications
Cancer Nanotechnology
Analytical Modeling for Biomechanical and Biomedical Systems
Implantable Devices
Tissue Engineering
Stem Cell Engineering
Drug and Gene Delivery
At least one or two more courses from:
Computational Molecular Biology
Algorithmic Foundations of Computational Biology
Biochemistry
Genetics
Cell and Molecular Biology
Introductory Microbiology
Principles of Immunology
Polymer Science for Biomaterials
Cell Physiology and Biophysics
Methods in Informatics and Data Science for Health
Quantitative Models of Biological Systems
Organic Chemistry II
Computational Vision
Visualizing Vision
Topics in Translational Research and Technologies
Principles of Neurobiology
Mechanisms and Meaning of Neural Dynamics
Biological Physics
21st Century Applications in Cell and Molecular Biology
4. Capstone Design 2
ENGN 1930LBiomedical Engineering Design and Innovation 11
ENGN 1931LBiomedical Engineering Design and Innovation II 11
5. General Education Requirement: At least four approved courses must be taken in the humanities and social sciences.
Total Credits21

Biophysics

Biophysics is a quantitative science at the intersection of the life and physical sciences.  It requires a significant level of competence in physics, chemistry, biology and math as reflected in the concentration requirements.  Students should work with their concentration advisor to develop a focused academic plan that complements the required research component of the concentration and allows students to develop analytical and quantitative skills.  Students are expected to take courses that will count toward the concentration for a grade (ABC/NC).

Student Goals:
Students in this concentration will:

  • Explore the relationship between biological and physical principles by successfully completing foundational courses in biology, physics, math and chemistry
  • Gain an in-depth knowledge of the interdisciplinary nature of life and physical sciences by selecting and successfully completing advanced courses in biology, physics, math, chemistry or related fields 
  • Develop skills to identify and analyze critical questions central to biophysics
  • Apply quantitative methods to problems at the interface of life and physical sciences 
  • Complete a research project with a faculty advisor that focuses on a particular theme or problem in the field of biophysics where students apply knowledge gained throughout the curriculum.

Additional detailed information about the field of Biophysics may be found at: https://www.brown.edu/academics/biology/undergraduate-education/undergraduate/biophysics

Standard program for the Sc.B. degree

Requirements

Physics
One of the following series: 2
Basic Physics A
and Basic Physics B 1
Foundations of Mechanics
and Foundations of Electromagnetism and Modern Physics
Analytical Mechanics
and Introduction to Relativity, Waves and Quantum Physics
PHYS 0470Electricity and Magnetism1
Chemisty
CHEM 0330Equilibrium, Rate, and Structure1
CHEM 0350Organic Chemistry I1
Select one other advanced Chemistry Course1
Math
MATH 0090Single Variable Calculus, Part I (or equivalent)1
MATH 0100Single Variable Calculus, Part II (or equivalent)1
MATH 0180Multivariable Calculus (or equivalent)1
Biology
BIOL 0200The Foundation of Living Systems (or equivalent)1
Select four additional biology or neuroscience courses chosen with approval of the advisor. 4
Directed Research: Students must take two semesters of research which may be satisfied by any of the opportunities listed below:2
Directed Research in Biology (BIOL 1950/BIOL 1960), Chemistry (CHEM 0970/CHEM 0980), or Physics (PHYS 1980)
COEX courses
A summer research experience in equivalent scope and scale to an independent study, but this would not count as course credit toward the concentration
Electives: Four electives in biology, physics, math/applied math, chemistry, neuroscience, engineering or computer science; at least 2 courses must be above the introductory level 24
Total Credits20

Computational Biology

Computational biology involves the analysis and discovery of biological phenomena using computational tools, and the algorithmic design and analysis of such tools. The field is widely defined and includes foundations in computer science, applied mathematics, statistics, biochemistry, molecular biology, genetics, ecology, evolution, anatomy, neuroscience, and visualization. 

Students may pursue a Bachelor of Arts or a Bachelor of Science. Students pursuing the ScB have the option of electing a concentration in Computational Biology with one of three focus areas: Computer Sciences, Biological Sciences, or Applied Mathematics & Statistics. Both programs require a senior capstone experience that pairs students and faculty in creative research collaborations.

Standard program for the A.B. degree

Prerequisites:2
Single Variable Calculus, Part II
Single Variable Calculus, Part II (Accelerated)
The Foundation of Living Systems
General Core Requirements: Biology2
Genetics
Biochemistry
Cell and Molecular Biology
General Core Requirements: Chemistry1
Equilibrium, Rate, and Structure
Organic Chemistry I
General Core Requirements: Computer Science2
Computing Foundations: Data
and Computing Foundations: Program Organization
and Computer Science: An Integrated Introduction
OR
Introduction to Object-Oriented Programming and Computer Science
and Introduction to Algorithms and Data Structures
OR
Computer Science: An Integrated Introduction
and Computer Science: An Integrated Introduction
OR
Accelerated Introduction to Computer Science
and Computer Science: An Integrated Introduction
and Introduction to Software Engineering
and Introduction to Computer Systems
and Theory of Computation
General Core Requirements: Probability & Statistics1
Statistical Inference I
OR
Advanced Introduction to Probability for Computing and Data Science
OR
Probability
Comp Bio Core Course Requirements4
Computational Molecular Biology
Inference in Genomics and Molecular Biology
AND two of the following:
Algorithmic Foundations of Computational Biology
Deep Learning
Foundations of Population Genetics
Computational Methods for Studying Demographic History with Molecular Data
Human Population Genomics
Machine Learning
Computational Probability and Statistics
Statistical Inference II
Additional course with Director approval
Total Credits12

University Writing Requirement:

As part of Brown’s writing requirement, all students must demonstrate that they have worked on their writing both in their general studies and their concentration. There are a number of ways for Computational Biology concentrators to fulfill these requirements:

Capstone Experience

Students enrolled in the computational biology concentration will complete a research project in their senior year under faculty supervision.  The themes of such projects evolve with the field and the technology, but should represent a synthesis of the various specialties of the program. The requirements are either one semester of reading and research with a CCMB Faculty member or approved advisor, or a 2000-level Computational Biology course.

Standard program for the Sc.B. degree

Prerequisites
MATH 0100Single Variable Calculus, Part II (or equivalent)1
or MATH 0170 Single Variable Calculus, Part II (Accelerated)
BIOL 0200The Foundation of Living Systems (or equivalent)1
General Core Course Requirements: Biology
BIOL 0470Genetics (prerequisite BIOL 0200 or equivalent)1
BIOL 0280Biochemistry1
or BIOL 0500 Cell and Molecular Biology
General Core Requirements: Chemisty
CHEM 0330Equilibrium, Rate, and Structure1
or CHEM 0350 Organic Chemistry I
General Core Requirements: Computer Science2-4
Computing Foundations: Data
and Computing Foundations: Program Organization
and Computer Science: An Integrated Introduction
OR
Introduction to Object-Oriented Programming and Computer Science
and Introduction to Algorithms and Data Structures
OR
Computer Science: An Integrated Introduction
and Computer Science: An Integrated Introduction
OR
Accelerated Introduction to Computer Science
and Computer Science: An Integrated Introduction
and Introduction to Software Engineering
and Introduction to Computer Systems
CSCI 0220Introduction to Discrete Structures and Probability1
General Core Requirements: Probability & Statistics
APMA 1650Statistical Inference I1
or CSCI 1450 Advanced Introduction to Probability for Computing and Data Science
or MATH 1610 Probability
General Core Requirements: Computational Biology
CSCI 1810Computational Molecular Biology1
APMA 1080Inference in Genomics and Molecular Biology1
Capstone Experience1
Directed Research/Independent Study
Individual Independent Study
Six courses in one of the following three tracks:6
Computer Science Track:
Three of the following:
Introduction to Computer Graphics
Database Management Systems
Artificial Intelligence
Machine Learning
Deep Learning
Probabilistic Methods in Computer Science
Design and Analysis of Algorithms
or other Computer Science courses approved by the concentration advisor
Three of the following:
Introduction to Computer Systems
Introduction to Software Engineering
Introduction to Software Engineering
Algorithmic Foundations of Computational Biology
Statistical Methods in Bioinformatics, I
Statistical Inference II
Foundations of Population Genetics
Computational Methods for Studying Demographic History with Molecular Data
Human Population Genomics
Computational Probability and Statistics
Biological Sciences track
At least four courses comprising a coherent theme in one of the following areas: Biochemistry, Ecology, Evolution, or Neurobiology.
AND select two courses from the following:
Statistical Inference II
Computational Probability and Statistics
Foundations of Population Genetics
Computational Methods for Studying Demographic History with Molecular Data
Human Population Genomics
Machine Learning
Deep Learning
Algorithmic Foundations of Computational Biology
Statistical Methods in Bioinformatics, I
Applied Mathematics & Statistics Track:
At least three courses from the following:
Statistical Inference II
Computational Probability and Statistics
Artificial Intelligence
Methods of Applied Mathematics II
and Methods of Applied Mathematics I
OR
Applied Partial Differential Equations I
and Applied Ordinary Differential Equations
At least three of the following:
Quantitative Models of Biological Systems
Foundations of Population Genetics
Computational Methods for Studying Demographic History with Molecular Data
Human Population Genomics
Machine Learning
Deep Learning
Algorithmic Foundations of Computational Biology
Statistical Methods in Bioinformatics, I
Total Credits18-20

Honors:

 In order to be considered a candidate for honors, students will be expected to maintain an outstanding record, with no "C's" in concentration courses and with a minimum of an "A-" average in concentration courses. In addition, students should take at least one semester, and are strongly encouraged to take 2 semesters, of reading and research with a CCMB faculty member or approved advisor. Students must submit to a public defense of their theses to be open to the CCMB community.

  • Students seeking honors are advised to choose a Thesis Advisor prior to the end of their Junior year
  • Students must complete the Registration form for Comp Bio and submit it to CCMB@BROWN.EDU

Any deviation from these rules must be approved by the director of undergraduate studies, in consultation with the student's advisor.