You're logged in as |

Biology (Undergraduate)

The Undergraduate Program in Biology offers courses at all levels of biological organization: molecular, cellular, organismal and population, and concentration programs with varying degrees of multidisciplinary or specialty foci.

There is a vast range of collaborative opportunities for independent study and research under the mentorship of the faculty. Faculty maintain research programs at the cutting edge of their fields including biochemistry, biophysics, cell biology, developmental biology, ecology, evolution, genetics, immunology, molecular biology, neurobiology, physiology, anatomy, artificial organs, microbiology, virology, experimental pathology, and pharmacology. The close administrative relationship with the Alpert Medical School offers additional opportunities for blending fundamental biology and clinical sciences in research projects at the Brown-affiliated hospitals.

The Office of Biology Undergraduate Education is a full service academic resource center, serving a key role for students of the life sciences here. The Office provides individualized advising for students at all levels; oversees student- faculty research collaborations, holds special events and maintains a variety of programs that support and enrich student life.

For further information, please visit: http://www.brown.edu/academics/biology/undergraduate-education/

Course usage information

BIOL 0030. Principles of Nutrition.

Introduces the basic principles of human nutrition, and the application of these principles to the specific needs of humans, and the role of nutrition in chronic diseases. Provides an overview of the nutrients and their use by the human body. Also examines the role of nutrients in specific functions and disease states of the body. Not for biology concentration credit. Enrollment limited to 100.

Fall BIOL0030 S01 15551 MW 8:30-9:50(01) (M. Flynn)
Course usage information

BIOL 0040. Nutrition for Fitness and Physical Activity.

Reviews the role of nutrition in physical activity and health. It is designed to provide the student with the information and skills needed to translate nutrition and physical activity recommendations into guidelines for both the athlete for maximal performance and the non-athlete to improve both health and body weight. Students will learn the use of the energy yielding nutrition in physical activity and how food choices can influence both athletic performance and long-term health through the effect on risk factors for chronic diseases. Prerequisite: BIOL 0030. Enrollment limited to 20. Instructor permission required.

Spr BIOL0040 S01 24465 T 4:00-6:30(16) (M. Flynn)
Course usage information

BIOL 0060. Introduction to Human Physiology.

An introduction to human physiology aimed primarily at undergraduates who have minimal to no Biology background or who are not concentrating in biology. Acquire a basic understanding of the physiological mechanisms that allow for the running of each major organ systems. Topics include basic cardiovascular, respiratory, urinary, digestive, endocrine, and neuromuscular function, as well as aspects of reproduction and exercise physiology. Not for biology concentration credit. Lab.

Course usage information

BIOL 0080. Biotechnology Management.

An examination of the pharmaceutical, biotechnology, and medical product industries: what they are, how they function, whence they originate, and various perspectives on why some succeed and others fail. Pathways from lab-bench to marketplace are described as are the pervasive influences of the FDA, patent office, and courts. Extensive reading; emphasis on oral presentation. Primarily intended for students planning a career in biomedical industry. Not for biology concentration credit. Students MUST register for the lecture section and the conference. Enrollment limited to 20.

Spr BIOL0080 S01 25354 T 4:00-6:30(16) (B. Bready)
Course usage information

BIOL 0100. Living Biology at Brown and Beyond.

This unique course has three goals: 1) provide students meaningful introductions to the people, places, projects and opportunities that comprise the Program in Biology at Brown, 2) foster student identities as valued members of our STEM community while helping each discover their unique interests and potential contributions, 3) arm students with a diversity of personal, professional and academic skills that will help them succeed in Biology at Brown and beyond. The course is especially tailored to those likely to pursue one of Brown’s many Biology concentrations, but it is open to all. Limited to 200 First Year students, mandatory S/NC.

Fall BIOL0100 S01 16790 M 3:00-5:30(15) (K. Smith)
Course usage information

BIOL 0140A. Topics in Science Communications: Science Journalism Practicum.

Participants will understand how to read scientific research papers to interpret their findings and communicate these to a broader lay audience; analyze and understand best practices in science writing and the challenges of covering science for mass media; interviewing; fair and balanced coverage in reporting; give and receive peer feedback. Not for concentration credit in Biological Sciences programs. Enrollment limited to 10. Instructor permission required. S/NC WRIT

Course usage information

BIOL 0140B. Communicating Science: Biological Illustration.

This Sophomore Seminar is an immersion practicum that conjoins art and science. Employing a series of techniques, students will learn the protocols of scientific rendering in an intensive hands-on approach. Field trips will include the RISD Nature Lab, the Brown Greenhouse, and John Hay collection of biomedical and botanical folios. Media will include graphite/ carbon dust; pen and ink (stipple, line); coquille board; scratch-board, colored pencil, watercolor and polymer clay. Course will be driven by project presentations focused on communicating science through illustration. Not for concentration credit in Biological Sciences programs. Instructor permission required; enrollment 10 students. 1/2 credit. SOPH

Course usage information

BIOL 0140C. Communicating Science: Animating Science.

Taught by RISD/Brown professors with the Science Ctr and Creative Mind Initiative, this course explores the pedagogy of using visual media to convey scientific concepts. The goal is to assess the quality of existing material and design new material that fill an educational need and makes science engaging and accessible. Lectures, labs, discussions, critiques and speakers. Teams collaborate on a series of short exercises leading to the creation of videos/animations explaining scientific concepts. Projects evaluated on accuracy, clarity of explanation, educational value, viewer engagement and creativity. Not for concentration credit in Biological Sciences programs. Enrollment limited to 12; instructor permission.

Fall BIOL0140C S01 15554 W 1:00-6:00 (J. Stein)
Course usage information

BIOL 0140K. Conservation Medicine.

How have fruit bats contributed to the emergence of Nipah virus in Malaysia? Is an infectious cancer going to drive the Tasmanian Devil to extinction? Will a warmer world be a sicker world? We will consider these and additional topics at the intersection of global change biology and infectious disease emergence in this course. The course should be of interest to pre-med, general biology and environmental studies concentrators seeking interdisciplinary learning classroom experience. This will satisfy "Area 3" organismal biology concentration requirement for Biology/Health-Human Biology. Expected background: BIOL 0200 or equivalent placement. Enrollment limited to 12 sophomores. Instructor permission required. SOPH

Course usage information

BIOL 0150A. Techniques and Analyses using DNA-Based Biotechnology.

Students will study and practice a range of methods used in molecular biology while examining the ways in which those tools are used in research and in the development of medical treatments. This experience, combined with the reading and discussion of selected papers from the primary literature, fosters development of a skill set critically important for the modern day biology student. Expected background: high school Biology course. Enrollment limited to 10 first year students. Instructor permission required. Half-credit course. S/NC. FYS

Fall BIOL0150A S01 16188 T 5:00-8:00PM (J. Hall)
Fall BIOL0150A L01 16189 Arranged (J. Hall)
Course usage information

BIOL 0150B. Statistical Computing for Biology.

Modern biological research is a data rich endeavor, necessitating strong quantitative and computational skills to interpret the results of experimental and observational studies. In this course we will explore the application of statistics and modeling in biological research and environmental science. Topics covered will include basic probability, experimental design, sampling, hypothesis testing and mathematical models for prediction. No prior statistics knowledge is assumed. Enrollment limited to 10 first year and sophomore students. Instructor permission, based on a portfolio review. This is a half-credit course. S/NC

Course usage information

BIOL 0150C. Methods for Extraction and Analyzing Secondary Metabolites of Medicinal Plants.

Plant secondary metabolites are currently the subject of much research interest when investigating new target compounds for potential medicine from natural products. New leads for drugs and phytomedicines from plants and plant parts have been increasing at a rapid rate especially by the pharmaceutical industry. Many plants have been selected and collected for their specific secondary compounds and healing powers by ethnobotanists in the field. The final step is the extraction, analyzing and identification of these plant specimens. Enrollment limited to 10 first year students. Instructor permission required. Half-credit course. S/NC. FYS

Spr BIOL0150C S01 25393 T 3:00-5:00 (F. Jackson)
Spr BIOL0150C L01 25394 Arranged (F. Jackson)
Course usage information

BIOL 0150D. Techniques in Regenerative Medicine: Cells, Scaffolds and Staining.

Regenerative Medicine, also known as Tissue Engineering, is the process of creating living, functional tissues to repair or replace native tissue or organ functions that have been lost due to disease or congenital defects. As such, it is a prominent scientific discipline that can either "stand alone" or complement material-based research efforts in the areas of device design, drug delivery, diagnostics and pharmaceuticals. Students will develop proficiencies in basic cell culture techniques, early stage tissue regeneration strategies and histochemical characterization of mammalian cell constructs. Enrollment limited to 10 first year students. Instructor permission required. Half-credit course. S/NC. FYS

Fall BIOL0150D S01 16044 Arranged 'To Be Arranged'
Fall BIOL0150D L01 16045 Arranged 'To Be Arranged'
Course usage information

BIOL 0160. Plants, Food, and People.

Examines the selection, breeding, cultivation and uses of food plants. Discusses the effects on agriculture of pathogens, climate change, and loss of biodiversity. Considers whether enough food can be produced for a world population of potentially 10 billion, while sustaining biodiversity and environmental quality. Course will include two papers and assistance from Writing Fellows; feedback from first paper will be available when writing second paper, WRIT. Enrollment limited to 50. WRIT

Course usage information

BIOL 0170. Biotechnology in Medicine.

Introduces undergraduates to the main technological advances currently dominating the practice of medicine. Provides an overview of the objectives, techniques, and problems related to the application of biomedical technology to the diagnosis and treatment of disease and the contemporary health care industry. Topics include: pharmaceutical development and formulation; organ replacement by prosthesis and transplantation; medical imaging; tissue engineering, therapeutic cloning, regenerative medicine; stem cells; societal, economic, and ethical issues. This course does carry Biology concentration credit.

Fall BIOL0170 S01 16046 MWF 12:00-12:50(12) 'To Be Arranged'
Course usage information

BIOL 0180. The Biology of AIDS.

AIDS represents an example of the vulnerability of humans to new infectious agents. We will review some human infectious diseases including small pox yellow fever and influenza, and then explore AIDS/HIV. First characterized in 1981, AIDS became the leading cause of death in U.S. males aged 25-44 within a decade. We will examine what factors make HIV such a potent pathogen. The course is intended for students beginning in biology. Expected: BIOL 0200, or equivalent placement. This course does carry Biology concentration credit.

Fall BIOL0180 S01 15657 MW 8:30-9:50(01) (P. Shank)
Course usage information

BIOL 0190E. Botanical Roots of Modern Medicine.

This course will explore a variety of medicinal plants found throughout the world, the diverse cultures that use them in their daily lives and the scientific underpinnings of their medicinal uses. In conjunction with readings, students will gain a hands-on approach in lab, observing, identifying and growing these plants. Enrollment limited to: 20. Students MUST register for the lecture section and the lab. FYS

Fall BIOL0190E S01 16286 MW 3:00-4:20(17) (F. Jackson)
Fall BIOL0190E L01 16287 W 4:30-5:50 (F. Jackson)
Course usage information

BIOL 0190F. Darwinian Medicine.

Explores evolutionary explanations of why we get sick, and how this can shape, or misshape, our interpretations of medicine. Draws on evolutionary genetics, population biology, molecular biology and physiology. This course will build on evolutionary biology and then focus on disease processes such as infection, aging, cancer, allergy, diabetes, and obesity. Enrollment limited to 20 first year students. FYS

Fall BIOL0190F S01 16288 TTh 1:00-2:20(10) (M. Tatar)
Course usage information

BIOL 0190P. Pride and Prejudice in the Development of Scientific Theories.

We will examine how the pace and shape of scientific progress is affected by the social/cultural context and the "personality" of the individual. We will look into how the interplay between society and the individual affects how scientific theories arise, are presented, are debated and are accepted. The course will initially focus on Charles Darwin and his theory of Natural Selection using the biography of Adrian Desmond and James Moore, "Darwin: The Life of a Tormented Evolutionist." Enrollment limited to 20 first year students. FYS DPLL

Fall BIOL0190P S01 16190 TTh 2:30-3:50(03) (S. Helfand)
Course usage information

BIOL 0190Q. Climate Change and Species Extinction.

In this course students will go beyond the headlines and delve into the science to explore the impact of climate change on species extinction. Students will explore the integration of science and technology through traditional textbooks, primary literature, open source databases, simulations, and discussions. Students will investigate the impact of climate change on species distribution, ecology, and behavior through interactive, inquiry-based, collaborative classroom investigations. Students will learn to integrate information from a variety of sources and disciplines and share their ideas through classroom discussion, written assignments, and oral presentations. Enrollment limited to 20 first year students. FYS WRIT

Course usage information

BIOL 0190R. Phage Hunters, Part I.

A research-based lab class for freshmen; both semesters are required in the sequence. Students will isolate and characterize a bacteriophage viruses found in the soil. Lab work includes isolation and purification of your own phage, DNA isolation and restriction mapping, and EM characterization of your phage. Several phages will be selected for genome sequencing over winter break, and are annotated in the spring. One hour lecture, discussion, and 3 hours lab per week. Expected: AP Biology or equivalent, and HS chemistry. Instructor permission required. Admittance based on review of applications in the first class. Limited to 20 freshmen. FYS

Fall BIOL0190R S01 15658 M 3:00-5:30(15) (S. Taylor)
Fall BIOL0190R L01 15659 Arranged (S. Taylor)
Course usage information

BIOL 0190S. Phage Hunters, Part II.

A research-based laboratory/class for freshmen; both semesters are required. Students will isolate and characterize a bacteriophage viruses found in the soil. Lab work includes isolation and purification of your own phage, DNA isolation and restriction mapping, and EM characterization of your phage. Several phages will be selected for genome sequencing over Winter Recess, and annotated in the spring. One hour of lecture/discussion, and 3 hours lab per week. Expected: AP Biology or equivalent, HS chemistry, and permission of the instructor. Students are expected to take fall and spring courses in the sequence. Enrollment limited to 20 first-year students. Instructor permission. FYS

Spr BIOL0190S S01 24548 WF 3:00-5:30(13) (S. Taylor)
Spr BIOL0190S L01 24549 M 3:00-5:30 (S. Taylor)
Course usage information

BIOL 0190T. Bioinformatics: A Practical Introduction.

The amount of biological sequence data has grown at an exponential pace and spurred the development of computational tools that allow biologists to use this information. Students will become familiar with useful bioinformatics tools used by researchers. The course will introduce concepts of information transfer in biological molecules, storage in public databases, and how to use tools to access this information and organize it meaningfully. We will explore tools for studying whole genomes, including high-throughput sequencing data to assemble genomes and mapping subsets. Students will gain hands-on experience using these tools. Expected: AP credit or equivalent placement for BIOL 0200. FYS

Course usage information

BIOL 0190U. Plant Development, Structure and Function.

This course presents an integrated account of development, structure and function in plants, especially seed plants. Enrollment limited to 20 first year students. FYS WRIT

Fall BIOL0190U S01 16191 TTh 10:30-11:50(13) (P. Heywood)
Course usage information

BIOL 0200. The Foundation of Living Systems.

A broad overview of biological systems, emphasizing patterns and processes that form the basis of life. Explores essentials of biochemistry, molecular, and cellular biology and their relationship to the larger issues of ecology, evolution, and development. Examines current research trends in biology and their influence on culture. Appropriate for all students interested in biology. Serves as a gateway course to much of the intermediate and advanced curriculum. Placement tests are offered (contact Jody_Hall@brown.edu); AP scores of 4 or 5 are equivalent to BIOL 0200, and place a student out of this course. Students will be assigned to a lab section during the second week of class.

Spr BIOL0200 S01 25413 MWF 11:00-11:50(04) (K. Miller)
Spr BIOL0200 L01 25417 Arranged (J. Hall)
Spr BIOL0200 L02 25418 Arranged (J. Hall)
Spr BIOL0200 L03 25419 Arranged (J. Hall)
Spr BIOL0200 L04 25420 Arranged (J. Hall)
Spr BIOL0200 L05 25421 Arranged (J. Hall)
Course usage information

BIOL 0210. Diversity of Life.

This course will explore biological diversity – the number of taxa, and the functions, and processes that support life – from the perspectives of ecology and evolutionary biology. It will draw on examples and case studies from the geological record, functional morphology, the evolution of organ systems in vertebrates, genomics, behavior and sexual selection in birds and invertebrates. Overarching themes will emphasize that taxonomic diversity is an emergent property of complex life on Earth, and the importance of diversity of biological functions and processes in generating and maintaining taxonomic diversity. Class Restriction: Freshmen and sophomores; others by instructor permission.

Fall BIOL0210 S01 16321 MWF 11:00-11:50(16) (J. Kellner)
Course usage information

BIOL 0280. Introductory Biochemistry.

Lectures and recitation sections explore the mechanisms involved in the principles of macromolecular structure and function, the organization and regulation of pathways for intermediary metabolism, and the transfer of information from genes to proteins. It is expected that students have taken CHEM 0350 or are taking it concurrently.

Spr BIOL0280 S01 25424 TTh 1:00-2:20(10) (L. Lapierre)
Spr BIOL0280 C01 25425 M 4:00-5:30 (L. Lapierre)
Spr BIOL0280 C02 25426 M 7:00-8:30PM (L. Lapierre)
Spr BIOL0280 C03 25427 T 4:00-5:30 (L. Lapierre)
Spr BIOL0280 C04 25428 T 7:00-8:30PM (L. Lapierre)
Spr BIOL0280 C05 25429 W 4:00-5:30 (L. Lapierre)
Spr BIOL0280 C06 25430 W 7:00-8:30PM (L. Lapierre)
Spr BIOL0280 C07 25431 Th 4:00-5:30 (L. Lapierre)
Spr BIOL0280 C08 25432 ThF 7:00-8:30PM (L. Lapierre)
Spr BIOL0280 C09 25433 F 1:00-3:00 (L. Lapierre)
Course usage information

BIOL 0285. Introductory Biochemistry Laboratory.

Working in small groups, students will examine enzymatic reactions in bacterial metabolic pathways. They will gather information from online databases, define a working model and test this model by purifying a target enzyme and characterizing its biochemical function. They will then propose a hypothesis for the enzymatic reaction mechanism and test this hypothesis by designing mutations in the enzyme active site and characterizing these mutant enzymes experimentally. Priority given to sophomore and junior students planning to enter research careers. Expected:Students have previously taken or are concurrently enrolled in BIOL 0280; preference given to students concurrently enrolled. Instructor permission required. Course credit 0.5; final grade determined for BIOL 0285.

Spr BIOL0285 S01 25434 M 1:00-5:00 (S. Taylor)
Spr BIOL0285 S02 25435 Th 2:30-6:30 (S. Taylor)
Course usage information

BIOL 0300. Endocrinology.

A basic examination of endocrinology with emphasis on hormone biosynthesis, mechanism of action, physiological roles, and endocrine pathology. Topics include: mechanism of action of steroid, amine, and peptide hormones; neuroendocrinology; reproductive endocrinology; and endocrinology of metabolism and calcium homeostasis. It is expected that students have taken BIOL 0200 (or equivalent) and CHEM 0350.

Course usage information

BIOL 0320. Vertebrate Embryology.

Introduction to the developmental anatomy of vertebrate embryos, including humans, in an evolutionary context, through lecture, discussion and microscope slide study. Gametogenesis through germ layers and their organ system derivatives. Expected: BIOL 0200, or equivalent placement, or AP Biology score of 4 or 5. Limited to 18 freshmen and 18 sophomores. Students MUST register for the lecture section and the lab.

Course usage information

BIOL 0350. The Fossil Record: Life through Time on Earth.

Course is designed for students with prior background in geology or evolutionary biology and who want to learn more about the fossil record, the origins of modern biodiversity and ecosystem structure, and interaction between organisms, and the geological and chemical cycles on the Earth. Lectures will cover major time periods during which animals and plants lived, as well as focusing on major transitions in the evolution of life on Earth. This course will fulfill requirements in both the geology/biology and evolutionary biology concentrations. Expected: BIOL 0210, GEOL 0240 or equivalent. Instructor permission, enrollment limited to 20 sophomores/juniors; register for course/lab.

Course usage information

BIOL 0380. The Ecology and Evolution of Infectious Disease.

Infectious diseases remain among the leading causes of death worldwide, and this burden is disproportionately borne by children living in low- and middle-income countries. Thus management of infectious disease remains a critical intellectual challenge in the 21st century. This course will develop and apply ecological and evolutionary theory to infectious microbes (and their hosts) via the detailed examination of a number of case studies. This will be accomplished by a combination of lectures, discussions, and readings drawn mainly from the primary literature. Assessment will be based on biweekly problem sets, two midterms and one final exam. Expected: BIOL 0200 or equivalent.

Fall BIOL0380 S01 16326 MWF 10:00-10:50(14) (D. Weinreich)
Course usage information

BIOL 0390. Vertebrate Evolution and Diversity.

An overview of vertebrate evolution that not only covers historical events, but also introduces various scientific concepts and modes of thought. Topics include past and present biodiversity, convergent evolution, biogeography, competition, continental drift, climatic change over time, the notion of evolution as progress, and a whole-animal approach to understanding evolutionary events. Enrollment limited to 50. WRIT

Course usage information

BIOL 0400. Biological Design: Structural Architecture of Organisms.

Many questions about the workings of living creatures can be answered by joining math, physics, and biology. We will identify basic physical science concepts that help biologists understand the structure and function of animals, plants, and microorganisms, and use these to study how the physical world constrains and facilitates the evolution of the extraordinary design and diversity of organisms. For first and second year students; others by permission. Recommended background: BIOL 0200, or equivalent. Enrollment limited to 40. Instructor permission required. WRIT

Fall BIOL0400 S01 16362 MWF 2:00-2:50(07) (S. Swartz)
Course usage information

BIOL 0410. Invertebrate Zoology.

A survey of invertebrate animals emphasizing evolutionary patterns and ecological relationships. Functional morphology, physiology, reproduction, development, and behavior of invertebrates will be examined. Laboratory exercises and two separate day-long field trips provide firsthand experience with the animals. Expected: BIOL 0200 or equivalent. Enrollment limited to 44. Students MUST register for the lecture section and a lab.

Fall BIOL0410 S01 16363 MWF 11:00-11:50(16) (C. Dunn)
Fall BIOL0410 L01 16364 W 1:00-3:50 (C. Dunn)
Course usage information

BIOL 0415. Microbes in the Environment.

Examines the diversity of microbial life in the environment. Surveys key services that microbes perform on land and sea, including biodegradation of contaminants in the environment and ecosystem processes related to climate change. Examines biological interactions of symbioses, quorum sensing, and antibiotic production in a ecological context. Explores the genomic mechanisms explaining phylogeny and life history strategies in microbes. Demonstrate knowledge of the diversity of microbes in the environment and benefits in an ecological/evolutionary context. Lecture based, two fieldtrips to expand appreciation for microbial ecology. BIOL 0200 or equivalent placement; CHEM 0330. Enrollment limited to 20 sophomores, juniors and seniors. WRIT

Course usage information

BIOL 0420. Principles of Ecology.

The principles, concepts, and controversies involved in the study of the distribution and abundance of plant and animal populations and their integration into natural communities. Emphasizes interactions among organisms and the hierarchical nature of ecological processes affecting individuals, populations, and communities. Expected: BIOL 0200 (or equivalent) and MATH 0090. Lectures and weekly discussion.

Spr BIOL0420 S01 25400 TTh 9:00-10:20(01) (J. Witman)
Spr BIOL0420 C01 25402 Arranged (J. Witman)
Course usage information

BIOL 0430. The Evolution of Plant Diversity.

Examines the evolutionary history of plants from a phylogenetic perspective. Introduces the science of phylogenetics - how to infer phylogenies and how to use them to understand organismal evolution. Highlights major trends in plant evolution over the past 400 million years. Lectures survey major plant lineages, with special focus on flowering plants. Weekly labs, field trips, and assignments stress basic plant anatomy and morphology, identification, and learning the local flora. Expected: BIOL 0200 (or equivalent placement). WRIT

Fall BIOL0430 S01 16366 TTh 9:00-10:20(08) (E. Edwards)
Fall BIOL0430 L01 16367 Arranged (E. Edwards)
Course usage information

BIOL 0440. Inquiry in Plant Biology: Analysis of Plant Growth, Reproduction and Adaptive Responses.

This course focuses on what plants do and how they do it. Introduces the biology of plants, their growth and development, structural features, and their cellular and organismal responses to key stimuli. Examines physiological, reproductive and developmental strategies throughout the plant life cycle and in relation to environmental challenges. During laboratory section meetings, students pursue inquiry-based group research projects addressing novel questions about mechanisms that control plant growth and development. Laboratory section is required. Prerequisites: One Brown course with laboratory section in either Biology or Chemistry. Enrollment limited to 24 students.

Spr BIOL0440 S01 25436 TTh 10:30-11:50(09) (A. DeLong)
Spr BIOL0440 L01 25437 W 2:00-4:50 (A. DeLong)
Course usage information

BIOL 0455. Coastal Ecology and Conservation.

Will enable to students to master fundamental ecological concepts and understand how this knowledge can be used to inform coastal conservation and management. Case studies from New England and elsewhere, field trips to rocky shores, salt marshes and coastal ecosystems enable students to develop scientific skills and experience the challenges of coastal conservation science. The course is aimed at freshmen and sophomores. Expected background: BIOL 0200 or equivalent placement. Enrollment limited to 10 students, and written permission required. Email (Mark_Bertness@brown.edu) to receive course application (due May 1). Admitted students register for the course in September.

Fall BIOL0455 S01 16370 TTh 1:00-2:20(10) (M. Bertness)
Course usage information

BIOL 0460. Insect Biology.

Focuses on characteristics that make insects unique and why more insect species have been described than all other organisms combined; the opportunity to investigate diversity and adaptation; their abundance, small size, and short lifespans; their importance as agents of biocontrol pollination, agricultural pests, and disease vectors. Expected: BIOL 0200 or equivalent. Enrollment limited to 20. Students MUST register for lecture AND lab. Primarily for freshmen and sophomores.

Course usage information

BIOL 0470. Genetics.

Genetic phenomena at the molecular, cellular, organismal, and population levels. Topics include transmission of genes and chromosomes, mutation, structure and regulation of the expression of the genetic material, elements of genetic engineering, and evolutionary genetics. One laboratory session and one discussion session per week. (Students should not plan to take BIOL 0470 after 1540.) Expected: BIOL 0200 (or equivalent placement). Students will be assigned to Lab sections the first week of class.

Fall BIOL0470 S01 16192 TTh 10:30-11:50(13) (M. Johnson)
Fall BIOL0470 L01 16193 Arranged (J. Hall)
Fall BIOL0470 L02 16194 Arranged (J. Hall)
Fall BIOL0470 L03 16195 Arranged (J. Hall)
Fall BIOL0470 L04 16196 Arranged (J. Hall)
Fall BIOL0470 L05 16197 Arranged (J. Hall)
Fall BIOL0470 L06 16198 Arranged (J. Hall)
Fall BIOL0470 L07 16199 Arranged (J. Hall)
Fall BIOL0470 L08 16200 Arranged (J. Hall)
Fall BIOL0470 L09 16201 Arranged (J. Hall)
Fall BIOL0470 L10 16202 Arranged (J. Hall)
Course usage information

BIOL 0480. Evolutionary Biology.

A broad introduction to the patterns and processes of evolution at diverse levels of biological organization. Topics covered include natural selection, adaptation, speciation, systematics, macroevolution, mass extinction events, and human evolution. Weekly discussion sections involve debates on original research papers. Occasional problem sets involve computer exercises with population genetics and phylogeny reconstruction. Expected: BIOL 0200 (or equivalent placement).

Fall BIOL0480 S01 16373 MWF 9:00-9:50(01) (D. Rand)
Fall BIOL0480 C01 16374 Arranged (D. Rand)
Course usage information

BIOL 0495. Statistical Analysis of Biological Data.

A first course in probability distributions and the use of statistical methods for biological data. Topics covered will include describing data, statistical inference (hypothesis tests and confidence intervals), analyzing associations, and methods for categorical data (contingency tables and odds ratios). Methods will be applied to data drawn from areas of biological inquiry. For statistics or related science credit in Biology programs. Expected background: BIOL 0200 or equivalent, math equivalent to MATH 0100. This course is for related science credit only in Biological Sciences concentration programs. Enrollment limited: 40 undergraduates-20 juniors and 20 sophomores. Registration for seniors requires permission from the instructor.

Spr BIOL0495 S01 25506 TTh 10:30-11:50(09) (S. Ramachandran)
Spr BIOL0495 C01 25507 Arranged (S. Ramachandran)
Course usage information

BIOL 0500. Cell and Molecular Biology.

This course examines the structure and function of the basic unit of an organism, the cell. An experimental approach is used to examine cellular functions, ranging from gene transcription, cell division and protein secretion, to cell motility, and signal transduction. Relevance to health and disease will be considered. Expected: BIOL 0200 (or equivalent placement).

Spr BIOL0500 S01 25438 MW 8:30-9:50(02) (A. Webb)
Course usage information

BIOL 0510. Introductory Microbiology.

Introduces role of microbes in our understanding of biology at the cellular and molecular level. Focuses on microbial significance for infectious disease, public health, genetics, biotechnology, and biogeochemical cycles. Laboratory involves basic microbiological techniques and selection and manipulation of microbes. Expected: BIOL 0200 (or equivalent placement). Students MUST register for the lecture section, conference, and the lab. Enrollment limited to 108.

Spr BIOL0510 S01 25455 MWF 1:00-1:50(06) (R. Bennett)
Spr BIOL0510 C01 25457 MWF 2:00-2:50 (R. Bennett)
Spr BIOL0510 L01 25458 T 9:00-11:50 (R. Bungiro)
Spr BIOL0510 L02 25459 T 1:00-3:50 (R. Bungiro)
Course usage information

BIOL 0530. Principles of Immunology.

Introduction to experimental and theoretical foundations of immunology. Focuses on concepts, landmark experiments and recent advances. Topics include innate and adaptive immunity; structure/function of antibody molecules and T cell receptors; regulation of immune responses through cellular interactions. Applications of concepts to medically significant issues (vaccines, transplantation, inflammation, autoimmunity, cancer, HIV/AIDS) are discussed. Interpretative analysis of experimental data is emphasized. Expected background: BIOL 0200 or equivalent placement credit.

Fall BIOL0530 S01 15660 TTh 2:30-3:50(03) (R. Bungiro)
Course usage information

BIOL 0800. Principles of Physiology.

Introduction to the function and integration of organ systems with an emphasis on human physiology. Includes basic concepts in cell and organ system physiology as well as fundamentals of modern trends in physiological science. Emphasizes the application of physical and chemical principles to organ function at both the cellular and systemic levels. Expected: BIOL 0200 or equivalent.

Fall BIOL0800 S01 16047 TTh 10:30-11:50(13) (J. Stein)
Fall BIOL0800 L01 16048 Arranged (A. Sobieraj)
Fall BIOL0800 L02 16049 Arranged (A. Sobieraj)
Fall BIOL0800 L03 16050 Arranged (A. Sobieraj)
Spr BIOL0800 S01 25355 MWF 10:00-10:50(03) (C. Hai)
Spr BIOL0800 L01 25356 Arranged (A. Sobieraj)
Spr BIOL0800 L02 25357 Arranged (A. Sobieraj)
Spr BIOL0800 L03 25358 Arranged (A. Sobieraj)
Spr BIOL0800 L04 25359 Arranged (A. Sobieraj)
Spr BIOL0800 L05 25360 Arranged (A. Sobieraj)
Course usage information

BIOL 0860. Diet and Chronic Disease.

This course addresses the relationship of food to the development and treatment of chronic diseases. Chronic diseases discussed are obesity, dyslipidemia/heart disease, diabetes mellitus, cancers and osteoporosis. Dietary recommendations for these diseases are critically assessed. Geared toward students interested in nutrition, medicine, and public health. Prerequisites: BIOL 0030 and 0800, plus permission of the instructor. Enrollment limited to 20.

Course usage information

BIOL 0920A. Controversies in Medicine.

Why and how do controversies in medicine emerge at specific moments in time? Why do scientists come to different conclusions based on the same data? Does it matter how we interpret controversies? This sophomore-level seminar critically analyzes contemporary controversies in medicine and public health. Using a case study approach, we will examine the social and political assumptions that inform important controversies. Questions related to the relationship between science, media, activism, and health inequality will be woven into the case studies. Enrollment limited to 20 sophomores. (For theme, not biology, credit in Health and Human Health and Biology only.) SOPH

Course usage information

BIOL 0940A. Viral Epidemics.

This sophomore seminar will examine epidemics (outbreaks) of viral infections from a historical perspective. We will also cover current literature and up to the minute news accounts of infectious disease related outbreaks occurring around the globe. The major focus will be on virus related diseases but any microbial outbreak in the news will be explored. The seminar will cover basic aspects of microbial pathogenesis so students can gain an appreciation of microbial host interactions. Essential writing skills will also be developed. Enrollment limited to 20 sophomore students. WRIT SOPH

Fall BIOL0940A S01 16204 Th 4:00-6:30(04) (W. Atwood)
Course usage information

BIOL 0940B. Sophomore Seminars in Biology: Life in a Shell.

This Sophomore seminar is an examination of broad themes in whole animal physiology with an emphasis on environmental adaptations. The foundation of the course will be the instructor’s recent book “Life in a Shell: A Physiologist’s View of Turtle.” A consideration of this iconic animal’s novel biological traits will lead into comparisons with our own biology and that of other animals. Topics: respiration, circulation, metabolic rate, buoyancy control, overwintering, migration, reproduction, and bone structure and function. Relevant original research papers will be used. Mandatory S/NC; enrollment of 20 students; override required. Expected: BIOL 0200 or equivalent placement credit. SOPH

Fall BIOL0940B S01 16051 T 4:00-6:30(09) (D. Jackson)
Course usage information

BIOL 0940C. Sophomore Seminar: Insect Biology.

Focuses on characteristics that make insects unique and why more insect species have been described than all other organisms combined; the opportunity to investigate diversity and adaptation; their abundance, small size, and short lifespans; their importance as agents of biocontrol pollination, agricultural pests, and disease vectors. Expected: BIOL 0200 or equivalent. Enrollment limited to 12 sophomores only. Students MUST register for lecture AND lab. SOPH

Course usage information

BIOL 0940D. Rhode Island Flora: Understanding and Documenting Local Plant Diversity.

This Sophomore Seminar focuses on species level identification of plants in Rhode Island and will cover the dominant plant species in each of the state’s main habitats including coastal wetlands and uplands, freshwater wetlands, peatlands, upland forests, and disturbed areas. Students will learn to identify plants using online interactive keys as well as more technical dichotomous keys and will also cover basic ecological processes in each habitat including the interaction of soils, geology, and hydrology. Materials related to plant morphology, plant taxonomy, plant evolution, understanding phylogenetic trees, and botanical illustration. Pre-requisites: BIOL 0200. Instructor permission required. SOPH

Fall BIOL0940D S01 16379 F 1:00-5:00 (T. Whitfeld)
Fall BIOL0940D L01 16380 Arranged (T. Whitfeld)
Course usage information

BIOL 0960. Independent Study in Science Writing.

Incorporates a nontechnical science journalism component into the BioMed curriculum. A series of four to six specific assignments are recommended, based on topics derived from another biology course taken previously by the student, whose instructor has agreed to serve as a BIOL 0960 sponsor. Assignments may include, for example, investigative or analytical reviews, or feature articles on ethical or social impacts of new discoveries. The student and instructor schedule meetings to discuss topics and due dates, review rough drafts, and evaluate completed work.

Not for concentration credit in the biological sciences programs. Permission must be obtained from the instructor prior to registering. Section numbers vary by instructor. Half credit.

Course usage information

BIOL 1040. Ultrastructure/Bioimaging.

This course examines microscopy and image analysis in the life sciences. Theoretical and practical aspects of microscopy will be discussed. Students will obtain hands-on experience with electron microscopy, light microscopy, fluorescence microscopy, and confocal microscopy. Students will learn to display images in 3D. Advanced undergraduates. Instructor permission required.

Spr BIOL1040 S01 25440 M 2:00-5:00 (G. Williams)
Course usage information

BIOL 1050. Biology of the Eukaryotic Cell.

Examines organelles and macromolecular complexes of eukaryotic cells with respect to structural and functional roles in major cellular activities. Emphasizes experimental basis for knowledge in modern cell biology using original literature, and discusses validity of current concepts. For advanced undergraduates and beginning graduate students. Complementary to BIOL 1270 and 1540. Prerequisites: BIOL 0280 or 0470 or 0500, or instructor permission. Graduate students register for BIOL 2050.

Fall BIOL1050 S01 16207 TTh 1:00-2:20(10) (K. Miller)
Fall BIOL1050 C01 16208 Th 12:00-12:50 (K. Miller)
Course usage information

BIOL 1070. Biotechnology and Global Health.

This course examines contemporary biotechnologies used to combat the predominant, worldwide problems in human health. Global health will be addressed from the scientific and engineering perspectives while integrating public health policy, health systems and economics, medical and research ethics, and technology regulation and management. This course is intended for graduate and advanced undergraduate students in biology, engineering, or related fields who have an interest in global health initiatives. Expected background: BIOL 0200 and BIOL 0800, or equivalents. Preference will be granted to graduate students in the Biotechnology and Biomedical Engineering programs. Only for related course credit in Biology, and for theme course credit in Health and Human Biology programs. Enrollment limited to 20. Instructor permission required.

Fall BIOL1070 S01 16058 TTh 2:00-3:30 (J. Schell)
Course usage information

BIOL 1090. Polymer Science for Biomaterials.

Basic principles of polymer science and its application in medicine. Topics include basic polymerization chemistry, kinetics of polymerization and depolymerization with emphasis on bioerodible polymers, characterization of polymers by physical methods, bulk and surface properties, behavior of polymers in solutions, crystallization, gelation, and liquid crystals. Hands-on experience with polymer characterization. Expected: CHEM 0350. Enrollment limited to 25.

Fall BIOL1090 S01 16059 T 1:00-3:50 (E. Mathiowitz)
Fall BIOL1090 L01 16060 Arranged (E. Mathiowitz)
Course usage information

BIOL 1100. Cell Physiology and Biophysics.

Current topics in cell physiology, with an emphasis on membrane-mediated interactions between cells and their environment. Topics may include: ion channel structure, function and regulation; intracellular regulatory molecules; mechanisms of sensory transduction; membrane receptors and second messenger systems; vesicle secretion; and cytoskeletal regulation of cell function. Lectures, discussion, and student presentations of the current literature. Expected: BIOL 0800 or NEUR 0010. Instructor permission required. Registration overrides will not be given out until after the first one or two classes. Enrollment limited to 30, and admission is based on seniority -- graduate students, seniors, then juniors. (Not for first and second-year undergraduates.)

Spr BIOL1100 S01 25363 M 3:00-5:30(13) (A. Zimmerman)
Course usage information

BIOL 1110. Topics in Signal Transduction.

Signal transduction is one of the most rapidly developing fields in biomedical sciences. Defects in signaling pathways can be responsible for diseases such as cancer, diabetes, cardiovascular disorders and psychoses. This course offers students an overview of the molecular pathways that allow cells to receive and process signals from their external environment, with an emphasis on the emerging state-of-the-art techniques used in their study. Expected background: BIOL 0200, 0280, 0470, or 0500. Enrollment limited to 20 juniors and seniors. Instructor permission required.

Fall BIOL1110 S01 16061 W 3:00-5:30(17) (E. Oancea)
Course usage information

BIOL 1120. Biomaterials.

A biomaterial is defined as a material suitable for use in medical implants that come in direct contact with patients' tissues. These include polymers, metals, and ceramics, and materials obtained from biological sources or through recombinant biotechnology. Goal: to provide comprehensive coverage of biomaterial science and technology. Emphasizes the transition from replacement to repair strategies. For advanced undergraduates and graduate students. Prerequisite: BIOL 0800 or instructor permission.

Spr BIOL1120 S01 25364 Th 4:00-6:30(17) (B. Zielinski-Habershaw)
Course usage information

BIOL 1140. Tissue Engineering.

Tissue engineering is an interdisciplinary field that incorporates progress in cellular and molecular biology, materials science, and engineering, to advance the goal of replacing or regenerating compromised tissue function. Using an integrative approach, we will examine tissue design and development, manipulation of the tissue microenvironment, and current strategies for functional reconstruction of injured tissues. Expected: CHEM 0330, plus BIOL 0500 or 0800. Enrollment limited to 20. Instructor permission required.

Fall BIOL1140 S01 17077 Th 3:00-5:50 (C. Kofron)
Course usage information

BIOL 1150. Stem Cell Engineering.

Stem cell engineering focuses on using adult, embryonic, and induced pluripotent stem cells to repair damaged or diseased tissues. This course will examine the role of stem cells in development, tissue homeostasis, and wound healing, as well as how they can be used for tissue engineering and cell-based regenerative therapies. We will also discuss the ethical, legal, and regulatory issues that accompany current and emerging stem cell engineering endeavors. The course will use an inverted lecture and classroom discussion format to effectively deliver relevant information. Emphasis is placed on oral and written communication skills applied to assignments, tests, and individual projects. As an additional part of this course, students will receive hands-on training in how to culture cells and assess samples for stemness characteristics in a group laboratory setting.

Course usage information

BIOL 1160. Principles of Exercise Physiology.

Application of the basic principles of physiology to the study of the response mechanisms of the human body during exercise. Topics include muscle and neural control, energy metabolism, cardiovascular and respiratory effects, endocrinology, principles of training, and special topics (e.g., diving, high altitude, and microgravity). Student presentations based on scientific articles are included. Expected: BIOL 0800 or written permission of the instructor.

Fall BIOL1160 S01 16063 MWF 1:00-1:50(06) (C. Hai)
Course usage information

BIOL 1180. Comparative Animal Physiology.

Comparative approach to the function and regulation of animal systems with an emphasis on vertebrates. Topics include circulation, gas exchange, neuromuscular function, excretion, acid-base and ion regulation, and temperature regulation. Considers the unity and diversity of physiological processes in animals differing both in phylogeny and environmental adaptation. Original papers are discussed. Expected: BIOL 0800 or equivalent.

Course usage information

BIOL 1190. Synaptic Transmission and Plasticity.

Synapses are the means by which the nervous system communicates. In this seminar-style course, we will explore the molecular and physiological underpinnings of synaptic transmission. We will then examine ways in which synapses can modulate their strength during development, learning, after addictive drugs, and other adaptive processes. Readings are ONLY from primary literature. Course recommended for juniors and seniors. Required: NEUR 1020. Enrollment limited to 20. Instructor's permission required, attendance at class on the first day is mandatory.

Course usage information

BIOL 1200. Protein Biophysics and Structure.

Structural Biology is the science to determine 3-dimensional structures of biomacromolecules (i.e. proteins, RNA, and DNA). These structures enable biologists to understand and explore their function. Since proteins, RNA, and DNA are the primary molecules of life, structural biology enables us to understand and influence these molecular machineries which form the basis of all biological processes. Throughout the class, the students will see examples of biologically important proteins and protein complexes that will allow them to correlate structure and biological function. Prerequisite: BIOL 0280.

Course usage information

BIOL 1210. Synthetic Biological Systems.

A multidisciplinary course that combines science and engineering providing a solid foundation in a cutting edge field of biological engineering. Synthetic biology is a mixture of biology, chemistry, engineering, genetic engineering and biophysics. It builds on recent work in systems biology which involves the modeling of biological systems, but goes further in that it involves the construction and standardization of biological parts, that fit together to form more complex systems. Expected: at least four courses beyond BIOL 0200, CHEM 0330, PHYS 0300, ENGN 0300, MATH 0900, or CSCI 0040.

Course usage information

BIOL 1220. Synthetic Biological Systems in Theory and Practice.

A multidisciplinary laboratory, lecture, and discussion based course that combines several areas of science and engineering providing a foundation in the cutting edge field of synthetic biological engineering. The field of synthetic biology is centered around trying to make biology easier to engineer. It builds on recent work in systems biology which involves the modeling of biological systems, but goes further in that it involves the construction and standardization of biological parts that fit together making complex systems. This course will combine classes, guest lectures and discussion lab visits to give students the best possible tools for understanding and applying research in synthetic biology. Expected: at least two courses in any of the key disciplines (biology, chemistry, physics, math, engineering, computer science) beyond the introductory level, and permission of the instructor.

Course usage information

BIOL 1222A. Current Topics in Functional Genomics.

A technological revolution in genomics has exponentially increased our ability to gather biological data. A host of new methods and types of analysis has arisen to accommodate this dramatic shift in data collection. The broad scope of inquiry has ushered in an era of “system-wide” approaches and brute-force strategies where rare signals can be detected and studied. In this seminar we will cover papers that embody this new approach. Students typically have taken an advanced undergraduate-level course in biology.

Fall BIOL1222A S01 16730 Arranged (N. Neretti)
Course usage information

BIOL 1250. Host-microbiome Interactions in Health and Disease.

Will focus on current understanding of how various microbiomes communicate and interact with the host and the factors that influence these interactions. We will discuss how the new technologies such as metagenomics and metabolomics have enhanced our understanding of host-microbiome interactions in health and disease. Students will have the opportunity to participate in discussions on how to apply recent discoveries to disease processes, health restoration and maintenance. The course will help students develop skills in critical thinking and in reading and evaluating original scientific literature. Expected: students with a background in basic microbiology (BIOL 0530 or its equivalent). 20 enrollment.

Spr BIOL1250 S01 25461 Th 2:30-5:30 (P. Belenky)
Course usage information

BIOL 1260. Physiological Pharmacology.

Covers the physiology of human disease (e.g., Heart failure and arrhythmia; cancer signaling pathways with a focus on breast cancer; neurological disorders such as schizophrenia and Parkinson's disease) and discusses the pharmacology of the drugs used to treat disease. A group of the most commonly prescribed drugs is discussed in terms of their fundamental modes of action and clinical importance. Expected: BIOL 0800.

Fall BIOL1260 S01 16065 TTh 10:30-11:50(13) (J. Marshall)
Course usage information

BIOL 1270. Advanced Biochemistry.

An advanced course in biochemistry, biochemical methods, and reading of the primary literature, featuring systematic coverage of the biochemistry of the central dogma, including DNA (replication, repair, recombination), RNA (regulation and mechanism of transcription, processing, turnover), and proteins structure, synthesis, modification, degradation, mechanisms of action, function). Expected: BIOL 0280, CHEM 0350, 0360. Graduate students register for BIOL 2270.

Fall BIOL1270 S01 16212 TTh 2:30-3:50(03) (A. Deaconescu)
Course usage information

BIOL 1290. Cancer Biology.

Provides a conceptual understanding of molecular events underlying development of human cancer. Focused on genetic changes leading to malignant transformation of cells. Covers cell cycle control, DNA damage, mutagenesis, cancer predisposition syndromes, oncogenic viruses, tumor immunology, metastasis, cancer chemotherapy and drug resistance. Lecture plus discussion of primary literature. Prerequisites: BIOL 0280 OR BIOL 0470 OR BIOL 0500.

Fall BIOL1290 S01 15794 MW 3:00-4:20(17) (A. Zhitkovich)
Course usage information

BIOL 1300. Biomolecular Interactions: Health, Disease and Drug Design.

Interactions between the molecules of life-proteins, RNA, DNA, membrane components-underlie all functions necessary for life. This course focuses on how nature controls these interactions, how these interactions can go awry in disease, and how we can learn the rules of these interactions to design drugs to treat disease. Students will review the physical basis of molecular interactions, learn classic and state-of-the-art high-resolution and high-throughput tools used to measure interaction, and survey the experimental and computational strategies to harness these interactions using a case study in rational drug design. Prerequisite: Introductory Biochemistry (BIOL 0280). Enrollment limited to 20; instructor permission. WRIT

Course usage information

BIOL 1310. Developmental Biology.

Covers the molecular and cellular events of development from fertilized egg to adult. Genetic basis of body form, cell fate specification and differentiation, processes controlling morphogenesis, growth, stem cells and regeneration will be examined. Differential gene regulation, intercellular signaling and their evolutionary conservation will be central to discussion of mechanisms governing developmental processes. Additional topics: developmental plasticity, impact of epigenetic and environmental factors, and basis of disease gleaned from developmental biology research. Live embryos will complement and reinforce concepts covered in class. Enrollment limited to 36. Expected: BIOL 0200 (or equivalent), and one course in genetics, cell biology or embryology.

Fall BIOL1310 S01 16214 TTh 9:00-10:20(08) (K. Wharton)
Fall BIOL1310 L01 16215 W 1:00-4:50 (K. Wharton)
Course usage information

BIOL 1330. Biology of Reproduction.

This course is an advanced, seminar-based course. Primary literature is emphasized to complement the format of extensive student seminar presentations. It is essential that students have a strong background in biology in order to gain the most from this course. The emphasis of the course is student seminar presentation and extensive discussion on the material. This is often the first opportunity for students to present/discuss science in a seminar format. Expected background: a course in Cell Biology (e.g. BIOL 0500 or 1050), and two additional Biology courses above the introductory (BIOL 0200) level. Enrollment limited to 20.

Spr BIOL1330 S01 25442 M 3:00-5:30(13) (G. Wessel)
Course usage information

BIOL 1410. Evolutionary Genetics.

This course will focus on selected topics in molecular population genetics, molecular evolution, and comparative genomics. Classic and current primary literature at the interface of evolution and genetics will be discussed in a seminar format. The laboratory involves wet-lab exercises (allozymes, PCR- RFLP, sequencing), plus computer labs using DNA analysis packages. Students will prepare a final grant proposal on specific research interests. Expected: BIOL 0470 or 0480. Students MUST register for the lecture section and the lab. Enrollment limited to 20.

Course usage information

BIOL 1420. Experimental Design in Ecology.

An overview and discussion of the basic principles used to design lab and field experiments in ecology and environmental science. Topics include: replication and statistical power, appropriate use of factorial designs, nonparametric methods, post hoc texts, natural versus manipulative experiments, experimental artifacts and impact study design. Discussions based on primary literature and a new text. Expected: BIOL 0420.

Course usage information

BIOL 1425. Phylogenetic Biology.

This course is the study of the evolutionary relationships between organisms, and the use of evolutionary relationships to understand other aspects of organism biology. This course will provide a detailed picture of the statistical, mathematical, and computational tools for building phylogenies and using them to study evolution. Enrollment is by instructor permission. Students will present scientific papers in class and complete a final project consisting of their own phylogenetic analysis. Expected Background: Evolutionary Biology and quantitative methods (such as statistics, computation, or math). Open to juniors, seniors, and graduate students. Enrollment limited to 16.

Spr BIOL1425 S01 25404 TTh 1:00-2:20(10) (C. Dunn)
Spr BIOL1425 C01 25405 Arranged (C. Dunn)
Course usage information

BIOL 1430. Population Genetics.

Population genetics considers the genetic basis of evolution: temporal changes in the genetic composition of populations in response to processes such as mutation, natural selection and random sampling effects. Starting from first principles, this course will develop a theoretical understanding of these dynamics. We will also explore the application of these tools to genomic-scale data in order to quantify the influence of various evolutionary processes at work in natural populations. Assessments will be based on problem sets, two midterm exams and one final exam. Prerequisites: MATH 0100 and one of BIOL 0470 or 0480, or permission.

Fall BIOL1430 S01 16381 MWF 11:00-11:50(16) (D. Weinreich)
Course usage information

BIOL 1440. Marine Biology.

An examination of current topics in the ecology of marine organisms and communities. Current literature and ideas are analyzed in a seminar format (5hr/week). A class research project provides hands-on experience with designing and interpreting experimental field work. Prerequisites: BIOL 0410 and 0420. Instructor's permission required.

Course usage information

BIOL 1450. Community Ecology.

This course will explore foundational concepts in community ecology, and will draw on examples and case studies from marine and terrestrial ecosystems, including species-rich tropical rain forests and coral reefs, the marine intertidal and benthic environments, and species-poor forests and grasslands of the temperate zone. Overarching themes will emphasize theoretical frameworks to understand the evolutionary origins and maintenance of this biological diversity. This will be accomplished using traditional lectures, weekly student-led discussions, readings of the primary literature, and class activities. Expected background: BIOL 200 or equivalent placement; and BIOL 0420; OR instructor permission.

Spr BIOL1450 S01 25406 MWF 10:00-10:50(03) (J. Kellner)
Course usage information

BIOL 1465. Human Population Genomics.

An introduction to human genomics and the evolutionary forces that shape observed genetic variation across humans today. Topics will include the relationship among humans and other primates, human population genetics and genomics, and examples of the concomitant evolution of both cultural traits and domesticated organisms. Assignments include a class presentation and reviewing papers on a selected topic. Expected background: BIOL 0470 or 0480, and BIOL 0495, PHP 2500, or equivalent. Enrollment limited to 25. Instructor permission required. WRIT

Course usage information

BIOL 1470. Conservation Biology.

Conservation Biology is the scientific study of the phenomena that affect the maintenance, loss, and restoration of biological diversity. Topics covered include: 1) the impacts of global warming, species invasions, and habitat destruction on biodiversity, 2) strategies developed to combat these threats, and 3) a consideration of key economic and ethical tradeoffs. Special attention will be paid to current debate and controversy within this rapidly emerging field of study. Readings will include the primary literature. A term-paper will be required. Prerequisite: BIOL 0420 or instructor permission. Enrollment limited to 30.

Fall BIOL1470 S01 16382 TTh 9:00-10:20(08) (D. Sax)
Course usage information

BIOL 1475. Biogeography.

Will provide an overview of the field of biogeography--the study of geography of living organisms. Class meetings will be split between lectures and discussions. Each discussion will expose students to foundational papers, which set the context for the field's development, and more recent papers, which show where the field is headed. Each student will conduct a short (but time consuming) original research project on some topic in biogeography. Prerequisites: BIOL 0420 and 0480. Expected: one taxonomy-based course (e.g., BIOL 0410, 0430, or 0460). Enrollment limited to 15 juniors, seniors, and graduate students. Instructor permission required.

Course usage information

BIOL 1480. Terrestrial Biogeochemistry and the Functioning of Ecosystems.

Three fundamental multidisciplinary questions will be addressed: How do ecosystems work? What limits the growth of life on Earth? How are humans altering the framework in which all life exists? Earth is basically a closed chemical system, and the reactions that support life are fueled by sunlight. But added to this chemistry and physics is the tremendous influence of life. Life created an oxygen atmosphere; the evolution of biological nitrogen fixation exponentially increased how many organisms could exist, and the soils that support human food production developed only by biologically-mediated processes. Throughout Earth's 4.5 billion-year history changes in Earth's basic biogeochemical processes have been fairly slow. Under our inattentive stewardship, we have almost instantaneously altered all of the major element cycles. We will focus heavily on what these changes mean for life on Earth. Instructor permission required.

Course usage information

BIOL 1485. Climate Change Health and Ecology.

Explore the linkages between climate change and health. Students will come to appreciate the topic through the foundations of the primary disciplines relevant to the field including global health, environmental change, disease ecology, and others. Climate-health linkages will be learned through weekly case studies addressed collectively through student-centered lectures, discussion of the primary literature, groups activities and guest lectures from campus faculty on topics ranging from climate migration to infectious disease range shifts. Expected background: BIOL 0475, or BIOL 1470, or PHP 1070, or PHP 1920, or equivalent experience with instructor's permission. Enrollment limited to 12 juniors and seniors.

Course usage information

BIOL 1495. 500 Million Years of Land Plants.

Explores the evolution of terrestrial plants and the ecosystems they structure. Introduces the fossil record of plants and basic patterns of plant diversification on land. Highlights major trends in the evolution of plant morphology, anatomy, and ecology. Lectures survey the diversity and community structure of different geological time periods. Weekly discussion sections, field trips, and assignments examine major evolutionary trends, particularly with regard to climatic changes over time. Expected: BIOL 0400, BIOL 0430, (or equivalent placement). Enrollment limited to 15 students; instructor permission; register for section and conference.

Fall BIOL1495 S01 17055 MWF 9:00-9:50(01) (A. Leslie)
Course usage information

BIOL 1500. Plant Physiological Ecology.

An in-depth look at plant ecological strategy, focusing on the anatomical and physiological adaptations of plants to particular environments. Additional topics include plant-animal interactions, historical biogeography, and community assembly processes. A comparative, phylogenetic approach is emphasized. Lectures present a broad overview of topics, and discussions focus on current outstanding problems. Lab exercises provide hands-on experience in designing experiments, measuring plant performance, and scientific writing. Required laboratory hours to be arranged by the instructor. Expected: BIOL 0430 or BIOL 0440. Enrollment limited to 15.

Spr BIOL1500 S01 25509 MW 10:30-11:50 (E. Edwards)
Spr BIOL1500 L01 25510 Arranged (E. Edwards)
Course usage information

BIOL 1520. Innate Immunity.

Innate immunity is the initial response to microbes that prevents infection of the host. It acts within minutes to hours, allowing the development of the adaptive response in vertebrates. It is the sole mechanism of defense in invertebrates such as insects. The components and mechanisms dictating this response are explored. Prerequisite: BIOL 0530. Enrollment limited to 30. Graduate students must obtain instructor permission.

Fall BIOL1520 S01 15661 MW 8:30-9:50(01) (L. Brossay)
Course usage information

BIOL 1540. Molecular Genetics.

Even in this era when whole genome DNA sequencing has become routine, there are still thousands of eukaryotic genes with unknown functions. Genetic screens for mutations that alter pathways of interest remain the premier approach to understanding gene function in the context of the organism. In Molecular Genetics students will learn the key concepts involved in designing and interpreting genetic screens using the powerful tools available in model animal, plant, and fungal organisms. Students will also learn how to understand and analyze results presented in the primary scientific literature. Furthermore, students will gain an appreciation of how the field of genetics has changed through discoveries and technological advances made over the past 50 years. Graduate students should register for BIOL 2540.

Spr BIOL1540 S01 25444 TTh 2:30-3:50(11) (J. Bender)
Course usage information

BIOL 1545. Human Genetics and Genomics.

This course will exemplify the power of genetically informed approaches to understanding human biology. It is intended for advanced undergraduate students and graduate students; prerequisites include BIOL0470 or equivalent. The course is based in lectures, reading material (textbook and primary literature), and in-class discussions. Course topics include: medical genetics and genomics; methods to study human genotypes and related phenotypes; industry-related topics; and ethical and societal implications of genome science. It will benefit students with career interests in basic science, medicine, biotechnology, or science policy. Enrollment is limited to 20 students; selection will be based on seniority, prerequisites, and registration order.

Spr BIOL1545 S01 25446 TTh 9:00-10:20(01) (E. Morrow)
Course usage information

BIOL 1550. Biology of Emerging Microbial Diseases.

Emerging diseases influence the health of human populations in less developed countries and are expected to have similar effects worldwide. Rising incidence of "new" diseases underscores the need for knowledge of infection mechanisms and their outcomes. Focuses on biochemical, genetic, cellular and immunological events of emerging pathogens and host responses. Expected: BIOL 0470 or BIOL 0530.

Spr BIOL1550 S01 25462 MWF 1:00-1:50(06) (C. de Graffenried)
Course usage information

BIOL 1555. Methods in Biomedical Informatics.

Will provide a methodological survey of approaches used in biomedical informatics. Particular emphasis given to formalisms and algorithms used within the context of biomedical research and health care, including those used in biomolecular sequence analysis, electronic health records, clinical decision support, and public health surveillance. Practical programming skills will also be taught within these contexts. The final project of the course is to demonstrate an understanding of biomedical informatics approaches through development of a solution within biomedical research or healthcare context. Prerequisite: introductory statistics course. Enrollment: 30 students. For biological science concentrators, graduate students, others with permission.

Spr BIOL1555 S01 25687 W 3:00-5:30(14) (N. Sarkar)
Course usage information

BIOL 1560. Virology.

Emphasizes the understanding of molecular mechanisms of viral pathogenesis. Begins with a general introduction to the field of virology and then focuses on the molecular biology of specific viruses that are associated with human disease. Lectures based on current literature. Prerequisite: BIOL 0280, 0470, or 0530, or instructor permission.

Course usage information

BIOL 1565. Survey of Biomedical Informatics.

Survey course provides overview of field of biomedical informatics. Topics include computer science, healthcare, biology, social science. This course is designed to be complementary to BIOL 1555. Emphasis given to understanding the organization of biomedical information, effective management of information using computer technology, impact of such technology on biomedical research, education, patient care. Major aim explores the process of developing and applying computational and information science techniques for assessing current information practices, determining information needs of health care providers and patients, developing interventions or supporting clinical practice using informatics, and evaluating the impact of informatics solutions from a biomedical perspective.

Fall BIOL1565 S01 17079 TTh 10:30-11:50(13) (N. Sarkar)
Course usage information

BIOL 1600. Development of Vaccines to Infectious Diseases.

Provides background steps involved in vaccine development, from conceptualization to production to deployment. Considers infectious diseases and associated vaccines in context of community health. Appropriate for students wanting to gain an understanding of vaccine science. Provides a foundation for advanced courses in immunology and infectious disease, biomedical research, or medical/graduate studies. Activities include a weekly section meeting for discussion of relevant primary literature, and a final project of the student’s choice in the form of an in–class presentation, a research paper or an approved alternative format. Expected: BIOL 0200 or equivalent placement; BIOL 0530, and at least one additional biology course.

Spr BIOL1600 S01 24624 MW 3:00-4:20(14) (R. Bungiro)
Course usage information

BIOL 1800. Animal Locomotion.

How and why do animals run, jump, swim and fly? Physiology, anatomy, ecology, and evolutionary history all influence, and are influenced by, the way animals move around. We will integrate analyses from many levels of biological organization - from molecular motors, through bone-muscle systems, to biogeography - with methods and approaches from mechanics, fluid dynamics, and robotics. Expected: BIOL 0800 and PHYS 0030. Instructor permission required. WRIT

Course usage information

BIOL 1820. Environmental Health and Disease.

Fundamental concepts relating to the adverse effects of chemical agents on human health. Topics include dose-response relationships, absorption, distribution, metabolism, excretion, mechanisms of toxicity, and the effects of selected environmental toxicants on organ systems. Many of these concepts will be reinforced through the use of a case-study approach where a pertinent environmental issue is incorporated into the ongoing lectures. Expected: BIOL 0500 and BIOL 0800, plus either ENVS 0490 or BIOL 0420.
Advanced students have priority.

Spr BIOL1820 S01 24623 MW 8:30-9:50(02) (J. Plavicki)
Course usage information

BIOL 1850. Environmental and Genetic Toxicology.

Human disease is produced by complex interactions between inherited genetic predisposition and environmental exposures. These interactions will be explored at the molecular, cellular, and systemic levels. Prototype diseases will include hereditary disorders of hemoglobin, hypercholesterolemia, birth defects, and cancer. Expected: Cell Biology.

Course usage information

BIOL 1870. Techniques and Clinical Applications in Pathobiology.

A methodology course featuring laboratory and lecture instruction in established and leading-edge technologies. Examples: flow cytometry (multi-parameter analysis, cell sorting); molecular biology (PCR, real time PCR, in situ hybridization, microarrays, DNA sequencing, bioinformatics); digital imaging (image acquisition, processing and analysis); confocal microscopy ; histology and immunohistochemistry(confocal, immuno-histochemistry).

Course usage information

BIOL 1880. Comparative Biology of the Vertebrates.

The biology, structure, and evolutionary history of the vertebrates considered phylogenetically, emphasizing evolution of the major body systems. Stresses an evolutionary approach to the correlation of structure and function with environment and mode of life. Labs include dissection of several different vertebrates and comparative osteological material. Emphasis of course is on critical thinking rather than memorization of material. Recommended: BIOL 0320 or 0800. First year students must obtain instructor permission to register. Enrollment limited to 32. Students MUST register for the lecture section and the lab.

Course usage information

BIOL 1890. Human Histology.

This course will provide an in-depth treatment of the “stuff we are made of” and the wonderful logic of its organization. This course focuses fi rst on the biology of the four basic tissues (epithelium, connective tissue, muscle and nerve) and second, how they contribute to the functional anatomy of all organs and systems. For Pfizer students only.

Course usage information

BIOL 1920B. Health Inequality in Historical Perspective.

Seminar takes a historical perspective to explore causes of health inequality in the US. Draws on studies from the 19th century-present. Examines socio–political and economic context of health/disease, focusing on how race, class, and gender shape the experience of health, disease causality, and public health responses. Includes health consequences of immigration, incarceration, race-based medicine, the Chicago heatwave, and Katrina. BIOL 0200 and work in Africana Studies and/or science-technology courses suggested. Not for biology concentration credit. It is suitable for a theme course for HHB. Enrollment restricted to 20, third- and fourth-year students.

Course usage information

BIOL 1920C. Social Contexts of Disease.

What shapes our understandings of disease, and what makes a disease real? How might we explain the demise of formerly prevalent diseases and the arrival of others? How do politics, technologies, and institutions affect conceptions of disease and structure their treatment? Will examine the impact of social context on patients' experiences of disease, including clinical, scientific, and public health approaches. Will consider disease in relation to social relationships, power of the state to regulate disease, and cultural care of the body. Enrollment limited to 20 students; instructor permission required; serves as Capstone in Health and Human Biology. Not for concentration credit. WRIT

Course usage information

BIOL 1920D. Race, Difference and Biomedical Research: Historical Considerations.

This advanced seminar places the current debate over race, health, and genetics in historical context. An overarching goal is to understand how the social world informs the scientific questions we ask, design of research studies, and interpretation of findings. How have the theories and practices of biomedical science and technology produced knowledge of “race” and racial difference historically? How does race relate to gender and class? What are the implications of this debate for understanding health inequality? Previous coursework in Africana Studies, biomedical science, history of science, and/or science and technology studies preferred. Enrollment limited to 20; instructor permission. WRIT

Spr BIOL1920D S01 24622 W 3:00-5:30(14) (L. Braun)
Course usage information

BIOL 1941A. Plants in a Changing Planet.

Plants are the foundation of Earth's ecosystems and essential to human survival and civilization. This seminar will examine the physiological, ecological, and evolutionary responses of plants to rapid environmental change, and the consequences for agriculture and the structure and function of natural systems. Expected background: at least one of the following courses - BIOL 0420, 0430, 0440, 0480, or ENVS 0490.

Course usage information

BIOL 1950. Directed Research/Independent Study.

Directed research/independent study in biological sciences: basic science, social studies of biomedical science, and clinically-oriented projects, mentored by individual faculty members in the Division of Biology and Medicine. Sites include campus and hospital based facilities. Projects can serve as the basis for Honors theses, or to fulfill research requirements in a Bio-Med concentration program. Students planning to use 1950/1960 to fulfill a concentration requirement must receive approval from the concentration advisor. No more than two (2) semesters of BIOL 1950/1960 may be used toward a concentration program in the biological sciences. Faculty from outside the Division may supervise projects for bio-med program concentrators, but should do so using their Department's own Independent Study course number.

Course usage information

BIOL 1960. Directed Research/Independent Study.

Directed research/independent study in biological sciences: basic science, social studies of biomedical science, and clinically-oriented projects, mentored by individual faculty members in the Division of Biology and Medicine. Sites include campus and hospital based facilities. Projects can serve as the basis for Honors theses, or to fulfill research requirements in a Bio-Med concentration program. Students planning to use 1950/1960 to fulfill a concentration requirement must receive approval from the concentration advisor. No more than two (2) semesters of BIOL 1950/1960 may be used toward a concentration program in the biological sciences. Faculty from outside the Division may supervise projects for bio-med program concentrators, but should do so using their Department's own Independent Study course number.

Course usage information

BIOL 1970A. Stem Cell Biology.

Senior seminar course will provide an interactive forum by which up to twenty seniors (and qualified juniors with permission) will explore the biology of stem cells from their humble beginnings in the embryo to their potential use in regenerative medicine. The potency and regulation of embryonic and adult stem cell populations derived from diverse organisms will be contrasted with laboratory-derived human stem-like cells for biomedical applications. Critical reading of classical and modern literature in the field of stem cell biology will form the basis of student-led presentations, papers and ethical forums. Expected: biochemistry, genetics and/or cell biology. Instructor permission; 20 students.

Fall BIOL1970A S01 16218 M 2:00-4:30 (R. Freiman)
Course usage information

BIOL XLIST. Courses of Interest to Biology Concentrators.

Associate Dean of Biology and Chair

Edward Hawrot

Director of Undergraduate Affairs

Katherine F. Smith

Professor

Roy K. Aaron
Professor of Orthopaedics; Professor of Molecular Pharmacology, Physiology and Biotechnology

Eli Y. Adashi
Professor of Medical Science

Edward Akelman
Professor of Orthopaedics

Jorge E. Albina
Professor of Surgery

Ruben Alvero
Professor of Obstetrics and Gynecology

Joseph F. Amaral
Professor Emeritus of Surgery

James A. Arrighi
Professor of Medicine; Professor of Diagnostic Imaging; Professor of Medical Science

Walter J. Atwood
Professor of Medical Science

Samuel I. Beale
Professor Emeritus of Biology

Jose Behar
Professor Emeritus of Medicine

Judith L. Bender
Professor of Biology

Richard J. Bennett
Professor of Biology

David M. Berson
Sidney A. Fox and Dorothea Doctors Fox Professor of Ophthalmology and Visual Science

Mark D. Bertness
Robert P. Brown Professor of Biology

Richard W. Besdine
David S. Greer, M.D., Professor of Geriatric Medicine

Christine Anne Biron
Esther Elizabeth Brintzenhoff Professor of Medical Science

Andrew S. Blum
Professor of Neurology

Kim Boekelheide
Professor of Medical Science

Jeffrey Borkan
Professor of Family Medicine; Professor of Medical Science

Christopher T. Born
Intrepid Heroes Professor of Orthopedic Surgery

Wayne D. Bowen
Upjohn Professor of Pharmacology

Elizabeth Brainerd
Professor of Biology; Professor of Medical Science

Lundy Braun
Professor of Medical Science; Professor of Africana Studies

Andrew S. Brem
Professor Emeritus of Pediatrics

Laurent Brossay
Charles A. and Helen B. Stuart Professor of Medical Science

Larry K. Brown
Professor of Psychiatry and Human Behavior

John E. Buster
Professor of Obstetrics and Gynecology

Blake Cady
Professor Emeritus of Surgery

Anthony Angelo Caldamone
Professor of Surgery (Urology); Professor of Pediatrics

Angela Caliendo
Professor of Medicine

Andrew G. Campbell
Professor of Medical Science

Michael Carey
Professor of Psychiatry and Human Behavior; Professor of Behavioral and Social Sciences

Charles C. J. Carpenter
Professor of Medicine

Stephen R. Carr
Professor of Obstetrics and Gynecology

Mary A. Carskadon
Professor of Psychiatry and Human Behavior

Angelina C. A. Carvalho
Professor Emerita of Medicine

William J. Cashore
Professor Emeritus of Pediatrics

Kimberle C. Chapin
Professor of Pathology and Laboratory Medicine; Professor of Medicine

Qian Chen
Michael G. Ehrlich, MD Professor of Orthopedic Research

William Cioffi
J. Murray Beardsley Professor of Surgery

Andrew Cohen
Professor of Medicine

Annette W. Coleman
Stephen T Olney Professor Emerita of Natural History

John R. Coleman
Professor Emeritus of Biology

Barry William Connors
L. Herbert Ballou University Professor of Neuroscience

Donald R. Coustan
Professor of Obstetrics and Gynecology

Joseph Crisco
Henry F. Lippitt Professor of Orthopedics

John J. Cronan
Charles and Elfriede Collis-Frances Weeden Gibson Professor of Diagnostic Imaging in the Warren Alpert Medical School

James P. Crowley
Professor Emeritus of Medicine

Susan Cu-Uvin
Professor of Obstetrics and Gynecology; Professor of Health Services, Policy and Practice; Professor of Medicine

Michele G. Cyr
Professor of Medicine; Professor of Medical Science

Albert E. Dahlberg
Professor Emeritus of Medical Science

Robert Paul Davis
Professor Emeritus of Medical Science

Suzanne M. De La Monte
Professor of Pathology and Laboratory Medicine; Professor of Neurosurgery

Ronald A. DeLellis
Professor Emeritus of Pathology and Laboratory Medicine

Penelope Hill Dennehy
Professor of Pediatrics

Phyllis A. Dennery
Sylvia Kay Hassenfeld Professor of Pediatrics

Paul A. Disilvestro
Professor of Obstetrics and Gynecology

John P. Donoghue
Henry Merritt Wriston Professor of Neuroscience

Gary S. Dorfman
Professor Emeritus of Diagnostic Imaging

Samuel Dudley
Ruth and Paul Levinger Professor of Cardiology

Damian E. Dupuy
Professor of Diagnostic Imaging

Lance D. Dworkin
Professor of Medicine

J. Donald Easton
Professor Emeritus of Neurology

Charles B. Eaton
Professor of Family Medicine; Professor of Epidemiology

Michael G. Ehrlich
Vincent Zecchino, MD Professor of Orthopaedic Surgery

Jack A. Elias
Frank L. Day Professor of Biology

Mel H. Epstein
Professor Emeritus of Neurosurgery

Nathan B. Epstein
Professor Emeritus of Psychiatry and Human Behavior

Paul Fadale
Professor of Orthopaedics

Justin R. Fallon
Professor of Medical Science; Professor of Psychiatry and Human Behavior

Anne Fausto-Sterling
Nancy Duke Lewis Professor Emerita of Biology

Patricia Jeanne Flanagan
Professor of Pediatrics

Timothy P. Flanigan
Professor of Medicine; Professor of Health Services, Policy and Practice

Braden C. Fleming
Lucy Lippitt Professor of Orthopedics

Edwin N. Forman
Professor Emeritus of Pediatrics

Jennifer F. Friedman
Professor of Epidemiology; Professor of Pediatrics

Joseph H. Friedman
Professor of Neurology

Gregory K. Fritz
Professor of Psychiatry and Human Behavior

Karen L. Furie
Samuel I. Kennison, M.D., and Bertha S. Kennison Professor of Clinical Neuroscience

Generoso G. Gascon
Professor Emeritus of Neurology and Professor Emeritus of Pediatrics

Stephen M. Gatesy
Professor of Biology; Professor of Medical Science

Susan A. Gerbi
George D. Eggleston Professor of Biochemistry

Arvin S. Glicksman
Professor Emeritus of Medical Science

Ziya L. Gokaslan
Professor of Neurosurgery

Richard J. Goldberg
Professor of Psychiatry and Human Behavior

George E. Goslow Jr
Professor Emeritus of Biology

Cornelius O. Granai
Professor of Obstetrics and Gynecology

Stefan Gravenstein
Professor of Medicine

Benjamin D. Greenberg
Professor of Psychiatry and Human Behavior

Paul B. Greenberg
Professor of Surgery (Ophthalmology)

Samuel H. Greenblatt
Professor Emeritus of Neurosurgery

A. Gerson Greenburg
Professor Emeritus of Surgery

Philip A. Gruppuso
Professor of Pediatrics; Professor of Medical Science; Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Jason B. Hack
Professor of Emergency Medicine

Chi-Ming Hai
Professor of Medical Science

David T. Harrington
Professor of Surgery

Anne C. Hart
Professor of Neuroscience

Edward Hawrot
Alva O. Way University Professor of Medical Science

Stephen L. Helfand
Professor of Biology

Peter Heywood
Professor of Biology

Frederic G. Hoppin Jr
Professor Emeritus of Physiology and Professor Emeritus of Medicine

Jeffrey I. Hunt
Professor of Psychiatry and Human Behavior

Vincent R. Hunt
Professor Emeritus of Family Medicine

Benjamin T. Jackson
Professor Emeritus of Surgery

Donald C. Jackson
Professor Emeritus of Medical Science

Ivor M. D. Jackson
Professor Emeritus of Medicine

Gregory D. Jay
Professor of Emergency Medicine; Professor of Medicine; Professor of Engineering (Research)

Conrad E. Johanson
Professor Emeritus of Neurosurgery

Lenworth N. Johnson
Professor of Surgery (Ophthalmology)

Agnes B. Kane
Professor of Medical Science

Julie Kauer
Professor of Medical Science; Professor of Neuroscience

William E. Kaye
Professor Emeritus of Surgery and Professor Emeritus of Medicine

Martin B. Keller
Professor Emeritus of Psychiatry and Human Behavior

Martin Keszler
Professor of Pediatrics

Nabil Y. Khoury
Professor Emeritus of Obstetrics and Gynecology

Louise Sadler Kiessling
Professor Emerita of Pediatrics and Professor Emerita of Family Medicine

Timothy Kinsella
Professor of Radiation Oncology

Petra Klinge
Professor of Neurosurgery

Paul M. Knopf
Charles A. and Helen B. Stuart Professor Emeritus of Medical Science

Robert Kohn
Professor of Psychiatry and Human Behavior

Gideon Koren
Professor of Medicine

Jonathan David Kurtis
Professor of Pathology and Laboratory Medicine

Edward V. Lally
Professor of Medicine

Arthur Landy
University Professor Emeritus of Medical Science

Abbot Laptook
Professor of Pediatrics

W. Dwayne Lawrence
Professor of Pathology and Laboratory Medicine

Louis A. Leone
Professor Emeritus of Medicine

Barry M. Lester
Professor of Psychiatry and Human Behavior; Professor of Pediatrics

Mitchell M. Levy
Professor of Medicine

William J. Lewander
Professor of Emergency Medicine; Professor of Pediatrics

James G. Linakis
Professor of Emergency Medicine; Professor of Pediatrics

Diane Lipscombe
Professor of Neuroscience

Paul Liu
Professor of Surgery (Plastic Surgery)

Sydney Louis
Professor Emeritus of Neurology

Francois I. Luks
Professor of Surgery; Professor of Obstetrics and Gynecology; Professor of Pediatrics

Martha B. Mainiero
Professor of Diagnostic Imaging

Paul F. Malloy
Professor of Psychiatry and Human Behavior

David E. Mandelbaum
Professor of Neurology; Professor of Pediatrics

Anthony Lynn Mansell
Professor Emeritus of Pediatrics

Donald J. Marsh
Frank L. Day Professor Emeritus of Biology

John Marshall
Professor of Medical Science

Edith Mathiowitz
Professor of Medical Science; Professor of Engineering

William W. Mayo-Smith
Professor of Diagnostic Imaging

Elinore McCance-Katz
Professor of Behavioral and Social Sciences; Professor of Psychiatry and Human Behavior

F. Dennis McCool
Professor of Medicine

Charles J. McDonald
Professor Emeritus of Medical Science

James T. McIlwain
Sidney A. Fox and Dorothea Doctors Fox Professor Emeritus of Opthalmology and Visual Science

Antone A. Medeiros
Professor Emeritus of Medicine

Michael J. Mello
Professor of Emergency Medicine; Professor of Health Services, Policy and Practice; Professor of Medical Science

Ulrike Mende
Professor of Medicine

Leonard A. Mermel
Professor of Medicine

Geralyn M. L. Messerlian
Professor of Pathology and Laboratory Medicine; Professor of Obstetrics and Gynecology

Ivan W. Miller
Professor of Psychiatry and Human Behavior

Kenneth Raymond Miller
Professor of Biology

Richard P. Millman
Professor of Medicine; Professor of Pediatrics

Christopher I. Moore
Professor of Neuroscience

Jeffrey R. Morgan
Professor of Medical Science; Professor of Engineering

David J. Morris
Professor Emeritus of Pathology and Laboratory Medicine

Paul E. Morrissey
Professor of Surgery

Douglass H. Morse
Hermon Bumpus Professor Emeritus of Biology

Steven F. Moss
Professor of Medicine

Albert S. Most
Professor of Medicine

Kimberly L. Mowry
Robin Chemers Neustein Professor of Biomedicine

John B. Murphy
Professor of Medicine; Professor of Family Medicine

Timothy P. Murphy
Professor of Diagnostic Imaging

Deborah L. Myers
Professor of Obstetrics and Gynecology

Wade Myers
Professor of Psychiatry and Human Behavior

Eleftherios E. Mylonakis
Dean's Professor of Medical Science

Justin M. Nash
Professor of Family Medicine; Professor of Psychiatry and Human Behavior

Georg Noren
Professor Emeritus of Neurosurgery

William Oh
Sylvia Kay Hassenfeld Professor Emeritus of Pediatrics

Steven M. Opal
Professor of Medicine

Brett Douglas Owens
Professor of Orthopaedics

James F. Padbury
William and Mary Oh - William and Elsa Zopfi Professor of Pediatrics for Perinatal Research

Mark A. Palumbo
Professor of Orthopaedics

Michael A. Paradiso
Sidney A. Fox and Dorothea Doctors Fox Professor of Ophthalmology and Visual Science

Alfred F. Parisi
Professor Emeritus of Medicine

Teri B. Pearlstein
Professor of Psychiatry and Human Behavior; Professor of Medicine

Georges Peter
Professor Emeritus of Pediatrics

Katharine A. Phillips
Professor of Psychiatry and Human Behavior

Maureen Phipps
Chace-Joukowsky Professor of Obstetrics and Gynecology

M. Halit Pinar
Professor of Pathology and Laboratory Medicine

Athena Poppas
Professor of Medicine

Raymond O. Powrie
Professor of Medicine; Professor of Obstetrics and Gynecology

Lawrence H. Price
Professor of Psychiatry and Human Behavior

M. Ruhul Quddus
Professor of Pathology and Laboratory Medicine

Peter J. Quesenberry
Paul Calabresi, M.D. Professor of Oncology

Abrar Qureshi
Professor of Dermatology; Professor of Epidemiology

David M. Rand
Stephen T. Olney Professor of Natural History

Steven A. Rasmussen
Mary E. Zucker Professor of Psychiatry and Human Behavior

Robert A. Reenan
Professor of Biology

Murray Resnick
Professor of Pathology and Laboratory Medicine

Louis B. Rice
Joukowsky Family Professor of Medicine

Josiah D. Rich
Professor of Medicine; Professor of Epidemiology

Thomas Roberts
Professor of Ecology and Evolutionary Biology

Leslie Robinson-Bostom
Professor of Dermatology

Randy Rockney
Professor of Pediatrics; Professor of Family Medicine; Professor of Medical Science

Frank G. Rothman
Professor Emeritus of Biology

Boris Rotman
Professor Emeritus of Medical Science

Sharon I. Rounds
Professor of Medicine; Professor of Pathology and Laboratory Medicine

Dwight J. Rouse
Professor of Obstetrics and Gynecology; Professor of Epidemiology

Stephen P. Salloway
Professor of Neurology; Professor of Psychiatry and Human Behavior

Jerome Sanes
Professor of Neuroscience

Fred Jay Schiffman
Sigal Family Professor of Humanistic Medicine

Robert Schwartz
Professor Emeritus of Pediatrics

John M. Sedivy
Hermon C. Bumpus Professor of Biology

Ronald Seifer
Professor of Psychiatry and Human Behavior

Frank W. Sellke
Karl E. Karlson, MD and Gloria A. Karlson Professor of Cardiothoracic Surgery

Peter R. Shank
Professor Emeritus of Medical Science

M. Tracie Shea
Professor of Psychiatry and Human Behavior

David Sheinberg
Professor of Neuroscience

Mark Sigman
Krishnamurthi Family Professor of Urology

Brian Silver
Professor of Neurology

Fredric Joel Silverblatt
Professor Emeritus of Medicine

James A. Simmons
Professor of Biology

Don B. Singer
Professor Emeritus of Pathology and Laboratory Medicine

Peter S. Smith
Professor Emeritus of Pediatrics

Stephen R. Smith
Professor Emeritus of Family Medicine

Anthony Spirito
Professor of Psychiatry and Human Behavior

Barry S. Stein
Professor Emeritus of Surgery

Edward S. Sternick
Professor Emeritus of Radiation Oncology

Sharon M. Swartz
Professor of Biology; Professor of Engineering

Joseph D. Sweeney
Professor of Pathology and Laboratory Medicine; Professor of Medicine

Patrick J. Sweeney
Professor Emeritus of Obstetrics and Gynecology

Robert M. Swift
Professor of Psychiatry and Human Behavior

Marc Tatar
Professor of Biology

Richard M. Terek
Professor of Orthopaedics

Walter R. Thayer Jr
Professor Emeritus of Medical Science

Peter G. Trafton
Professor Emeritus of Orthopaedics

Glenn A. Tung
Professor of Diagnostic Imaging

Allan R. Tunkel
Professor of Medicine; Professor of Medical Science

Audrey R. Tyrka
Professor of Psychiatry and Human Behavior

Herman H. Vandenburgh
Professor Emeritus of Pathology and Laboratory Medicine and Professor Emeritus of Molecular Pharmacology, Physiology and Biotechnology

Michael P. Vezeridis
Professor of Surgery

Betty R. Vohr
Professor of Pediatrics

Jonathan K. Waage
Professor Emeritus of Biology

Jack R. Wands
Jeffrey and Kimberley Greenberg - Artemis and Martha Joukowsky Professor of Gastroenterology

Arnold-Peter Weiss
Professor of Orthopaedics

Katharine D. Wenstrom
Professor of Obstetrics and Gynecology

Gary M. Wessel
Professor of Biology

Robert J. Westlake
Professor Emeritus of Psychiatry and Human Behavior

Kristi A. Wharton
Professor of Biology

Edward Wing
Professor of Medical Science

Rena R. Wing
Professor of Psychiatry and Human Behavior

Jon D. Witman
Professor of Biology

Ronald A. Yankee
Professor Emeritus of Medicine

Anatoly Zhitkovich
Professor of Medical Science

Stanley Zimmering
Professor Emeritus of Biology

Anita L. Zimmerman
Professor of Medical Science

Mark Zimmerman
Professor of Psychiatry and Human Behavior

Brian Zink
Frances Weeden Gibson-Edward A. Inannuccilli, MD, Professor of Emergency Medicine

Caron Zlotnick
Professor of Psychiatry and Human Behavior; Professor of Medicine; Professor of Obstetrics and Gynecology

Professor Research

Kailash C. Agarwal
Professor Emeritus of Medical Science (Research)

Alfred Ayala
Professor of Surgery (Research)(Trauma)

Piero Biancani
Professor Emeritus of Medicine (Research)

Beth C. Bock
Professor of Psychiatry and Human Behavior (Research); Professor of Behavioral and Social Sciences (Research)

Ming-Yu Wang Chu
Professor Emerita of Medicine (Research)

Shih-Hsi Chu
Professor Emeritus of Molecular Pharmacology, Physiology and Biotechnology (Research)

Suzanne Colby
Professor of Psychiatry and Human Behavior (Research); Professor of Behavioral and Social Sciences (Research)

Leslie B. Gordon
Professor of Pediatrics (Research)

Elizabeth O. Harrington
Professor of Medicine (Research)

Douglas C. Hixson
Professor Emeritus of Pathology and Laboratory Medicine (Research) and Professor Emeritus of Medicine (Research)

Valerie S. Knopik
Professor of Psychiatry and Human Behavior (Research); Professor of Behavioral and Social Sciences (Research)

Chun Geun Lee
Professor of Molecular Microbiology and Immunology (Research)

Richard H. Longabaugh
Professor Emeritus of Psychiatry and Human Behavior (Research)

Elizabeth L. McQuaid
Professor of Psychiatry and Human Behavior (Research); Professor of Pediatrics (Research)

Eduardo A. Nillni
Professor Emeritus of Molecular Biology, Cell Biology and Biochemistry (Research) and Professor Emeritus of Medicine (Research)

Glenn E. Palomaki
Professor of Pathology and Laboratory Medicine (Research)

Bai-Chuan Pan
Professor Emeritus of Medicine (Research)

Jonathan S. Reichner
Professor of Surgery (Research)

Robert J. Smith
Professor of Medicine (Research); Professor of Health Services, Policy and Practice (Research)

Peter Snyder
Professor of Neurology (Research)

Nancy L. Thompson
Professor Emerita of Pathology and Laboratory Medicine (Research) and Professor Emerita of Medicine (Research)

Jennifer Tidey
Professor of Psychiatry and Human Behavior (Research); Professor of Behavioral and Social Sciences (Research)

Shuping Tong
Professor of Medicine (Research)

Shougang Zhuang
Professor of Medicine (Research)

Professor Clinical

Gowri Anandarajah
Professor of Family Medicine (Clinical); Professor of Medical Science (Clinical)

Diane J. Angelini
Professor Emerita of Obstetrics and Gynecology (Clinical)

Timothy Babineau
Professor of Surgery (Clinical)

Mara G. Coyle
Professor of Pediatrics (Clinical)

Curtis Doberstein
Professor of Neurosurgery (Clinical)

Vickie R. Driver
Professor of Orthopaedics (Clinical)

Janet Grace
Professor of Psychiatry and Human Behavior (Clinical)

Colin Harrington
Professor of Psychiatry and Human Behavior (Clinical); Professor of Medical Science (Clinical); Professor of Medicine (Clinical)

Pamela C. High
Professor of Pediatrics (Clinical)

Margaret M. Howard
Professor of Psychiatry and Human Behavior (Clinical); Professor of Medicine (Clinical)

Arlet G. Kurkchubasche
Professor of Surgery (Clinical); Professor of Pediatrics (Clinical)

Syed A. Latif
Professor of Pathology and Laboratory Medicine (Clinical)

Paul B. Lieberman
Professor of Psychiatry and Human Behavior (Clinical)

Debra J. Lobato
Professor of Psychiatry and Human Behavior (Clinical); Professor of Pediatrics (Clinical)

Kathleen M. McCarten
Professor Emerita of Pediatrics (Clinical) and Professor Emerita of Diagnostic Imaging (Clinical)

Marjorie A. Murphy
Professor of Surgery (Clinical)(Ophthalmology)

Linda Nici
Professor of Medicine (Clinical)

Richard B. Noto
Professor of Diagnostic Imaging (Clinical)

Bonnie O'Connor
Professor Emerita of Pediatrics (Clinical)

Barbara Schepps
Professor Emerita of Diagnostic Imaging (Clinical)

Linda K. Snelling
Professor of Pediatrics (Clinical); Professor of Surgery (Clinical)

Rosalind M. Vaz
Professor Emerita of Pediatrics (Clinical)

Conrad W. Wesselhoeft Jr
Professor Emeritus of Surgery (Clinical)

Clinical Professor

Scott David Berns
Clinical Professor of Pediatrics; Clinical Professor of Health Services, Policy and Practice

Rosemarie Bigsby
Clinical Professor of Pediatrics; Clinical Professor of Psychiatry and Human Behavior

Andrew S. Blazar
Clinical Professor Emeritus of Obstetrics and Gynecology

Walter A. Brown
Clinical Professor of Psychiatry and Human Behavior

Gary Bubly
Clinical Professor of Emergency Medicine; Clinical Professor of Medicine

Frederick W. Burgess
Clinical Professor of Surgery (Anesthesiology)

Joseph Chazan
Clinical Professor Emeritus of Medicine

William M. Corrao
Clinical Professor of Medicine

Robert Crausman
Clinical Professor of Medicine

Lynn C. Epstein
Clinical Professor Emerita of Psychiatry and Human Behavior and Clinical Professor Emerita of Behavioral and Social Sciences

Martin E. Felder
Clinical Professor Emeritus of Surgery

B. Allen Flaxman
Clinical Professor Emeritus of Dermatology

Roberta E. Goldman
Clinical Professor of Family Medicine

Frederick G. Guggenheim
Clinical Professor of Psychiatry and Human Behavior

Kathleen C. Hittner
Clinical Professor Emerita of Surgery (Anesthesiology)

Edward A. Iannuccilli
Clinical Professor Emeritus of Medicine

Charles B. Kahn
Clinical Professor Emeritus of Medicine

Stephen R. Kaplan
Clinical Professor Emeritus of Medicine

Joel M. Kaufman
Clinical Professor of Neurology

Arthur B. Kern
Clinical Professor Emeritus of Dermatology

Peter D. Kramer
Clinical Professor Emeritus of Psychiatry and Human Behavior

Carol Landau
Clinical Professor of Psychiatry and Human Behavior; Clinical Professor of Medicine

Mary D. Lekas
Clinical Professor Emerita of Surgery

Hon Fong Louie Mark
Clinical Professor Emerita of Pediatrics

Andrew Maslow
Clinical Professor of Surgery (Anesthesiology)

Jack M. Monchik
Clinical Professor of Surgery (Endocrine Surgery)

Andrew T. Nathanson
Clinical Professor of Emergency Medicine

Elizabeth M. Nestor
Clinical Professor of Emergency Medicine

Lyman A. Page
Clinical Professor Emeritus of Pediatrics

Robert B. Patterson
Clinical Professor of Surgery

Victor E. Pricolo
Clinical Professor of Medical Science

Lawrence Proano
Clinical Professor of Emergency Medicine

Herbert Rakatansky
Clinical Professor Emeritus of Medicine

Patricia R. Recupero
Clinical Professor of Psychiatry and Human Behavior; Clinical Professor of Medical Science

Michelle L. Rickerby
Clinical Professor of Psychiatry and Human Behavior

Philip R. Rizzuto
Clinical Professor of Surgery (Ophthalmology)

Stephen N. Rous
Clinical Professor Emeritus of Surgery

Francis H. Scola
Clinical Professor Emeritus of Diagnostic Imaging

Samir A. Shah
Clinical Professor of Medicine

Arun Singh
Clinical Professor Emeritus of Surgery

Boris Skurkovich
Clinical Professor of Pediatrics

Jacques G. Susset
Clinical Professor Emeritus of Surgery (Urology)

King W. To
Clinical Professor of Surgery (Ophthalmology)

Barbara Tylenda
Clinical Professor of Psychiatry and Human Behavior

Marguerite B. Vigliani
Clinical Professor of Obstetrics and Gynecology

Barry W. Wall
Clinical Professor of Psychiatry and Human Behavior

Marjorie E. Weishaar
Clinical Professor of Psychiatry and Human Behavior

Joel K. Weltman
Clinical Professor Emeritus of Medicine

Russell E. White
Clinical Professor of Surgery

John P. Wincze
Clinical Professor of Psychiatry and Human Behavior

Professor MBL

Mark Alliegro
Professor of Molecular Biology, Cell Biology and Biochemistry (MBL)

Zoe G. Cardon
Professor of Ecology and Evolutionary Biology (MBL)

Linda Deegan
Professor of Ecology and Evolutionary Biology (MBL)

Jonathan David Gitlin
Professor of Molecular Biology, Cell Biology and Biochemistry (MBL)

Roger T. Hanlon
Professor of Ecology and Evolutionary Biology (MBL)

Associate Professor

J. Gary Abuelo
Associate Professor of Medicine

Sun Ho Ahn
Associate Professor of Diagnostic Imaging

Carlos Aizenman
Associate Professor of Neuroscience

Rebecca H. Allen
Associate Professor of Medical Science; Associate Professor of Obstetrics and Gynecology

Brian K. Alverson
Associate Professor of Pediatrics; Associate Professor of Medical Science

Siraj Amanullah
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

David C. Anthony
Associate Professor of Family Medicine

Mary B. Arnold
Associate Professor Emerita of Pediatrics

Michael K. Atalay
Associate Professor of Diagnostic Imaging; Associate Professor of Medicine

Barbara E. Barker
Associate Professor Emerita of Pathology and Laboratory Medicine

Gilad Barnea
Associate Professor of Neuroscience

Jay M. Baruch
Associate Professor of Emergency Medicine

George P. Bayliss
Associate Professor of Medicine

Curt G. Beckwith
Associate Professor of Medicine

Michael D. Beland
Associate Professor of Diagnostic Imaging

G. Jesse Bender
Associate Professor of Pediatrics

Joseph M. Bliss
Associate Professor of Pediatrics

Andrew G. Bostom
Associate Professor of Medicine

Jerrold L. Boxerman
Associate Professor of Diagnostic Imaging

Linda L. Brown
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

Weibiao Cao
Associate Professor of Pathology and Laboratory Medicine; Associate Professor of Medicine

E. Jane Carter
Associate Professor of Medicine

Kevin J. Chang
Associate Professor of Diagnostic Imaging

Kevin P. Charpentier
Associate Professor of Surgery

Elizabeth S. Chen
Associate Professor of Medical Science; Associate Professor of Health Services, Policy and Practice

Eunyoung Cho
Associate Professor of Dermatology; Associate Professor of Epidemiology

Thomas H. Chun
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

Jennifer G. Clarke
Associate Professor of Obstetrics and Gynecology

Brian Clyne
Associate Professor of Emergency Medicine; Associate Professor of Medical Science

Erika D'Agata
Associate Professor of Medicine

Eric M. Darling
Associate Professor of Medical Science; Associate Professor of Engineering; Associate Professor of Orthopaedics

Alison DeLong
Associate Professor of Biology

Vera A. De Palo
Associate Professor of Medicine

Joseph Anthony Diaz
Associate Professor of Medicine; Associate Professor of Medical Science

Daniel P. Dickstein
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Diagnostic Imaging; Associate Professor of Pediatrics

Susan Dickstein
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Pediatrics

Thomas Dipetrillo
Associate Professor of Radiation Oncology

John E. Donahue
Associate Professor of Pathology and Laboratory Medicine; Associate Professor of Neurology

David M. Dosa
Associate Professor of Health Services, Policy and Practice; Associate Professor of Medicine

Susan Duffy
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

Casey Dunn
Associate Professor of Biology

Craig P. Eberson
Associate Professor of Orthopaedics

Erika J. Edwards
Associate Professor of Ecology and Evolutionary Biology

Thomas K. Egglin
Associate Professor of Diagnostic Imaging

Allan Erickson
Associate Professor Emeritus of Medicine

William G. Fairbrother
Associate Professor of Biology

Lloyd R. Feit
Associate Professor of Pediatrics

Richard N. Freiman
Associate Professor of Medical Science

Rebekah L. Gardner
Associate Professor of Medicine

Aris C. Garro
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

Paul George
Associate Professor of Family Medicine; Associate Professor of Medical Science

Reginald Y. Gohh
Associate Professor of Medicine

Dragan Golijanin
Associate Professor of Surgery (Urology)

Geetha Gopalakrishnan
Associate Professor of Medicine

David J. Grand
Associate Professor of Diagnostic Imaging

Emily P. Green
Associate Professor of Medical Science

Rogers C. Griffith
Associate Professor of Pathology and Laboratory Medicine

Katrine Hansen
Associate Professor of Pathology and Laboratory Medicine

Roman A. Hayda
Associate Professor of Orthopaedics

Diane Hoffman-Kim
Associate Professor of Medical Science; Associate Professor of Engineering

Andrew W. Holowinsky
Associate Professor Emeritus of Molecular Biology, Cell Biology and Biochemistry

Cynthia L. Jackson
Associate Professor of Pathology and Laboratory Medicine

Mahesh V. Jayaraman
Associate Professor of Diagnostic Imaging; Associate Professor of Neurology; Associate Professor of Neurosurgery

Elissa Jelalian
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Pediatrics

Gerwald Jogl
Associate Professor of Biology

Mark A. Johnson
Associate Professor of Biology

Rich Jones
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Neurology

Rami Kantor
Associate Professor of Medicine

Julie Katarincic
Associate Professor of Orthopaedics

Leo Kobayashi
Associate Professor of Emergency Medicine

W. Curt Lafrance
Associate Professor of Psychiatry and Human Behavior; Associate Professor of Neurology

Michelle A. Lally
Associate Professor of Medicine; Associate Professor of Health Services, Policy and Practice

Robert E. Lambiase
Associate Professor of Diagnostic Imaging

Lucia Larson
Associate Professor of Medicine

Elizabeth Lazarus
Associate Professor of Diagnostic Imaging

Otto Liebmann
Associate Professor of Emergency Medicine

John R. Lonks
Associate Professor of Medicine; Associate Professor of Medical Science

Ana P. Lourenco
Associate Professor of Diagnostic Imaging

Kristen A. Matteson
Associate Professor of Obstetrics and Gynecology

James R. McCartney
Associate Professor Emeritus of Psychiatry and Human Behavior

Kelly Ann McGarry
Associate Professor of Medicine

John E. McGeary
Associate Professor of Psychiatry and Human Behavior

Alyson J. McGregor
Associate Professor of Emergency Medicine

George R. McKendall
Associate Professor of Medicine

Roland C. Merchant
Associate Professor of Emergency Medicine; Associate Professor of Epidemiology

Ralph P. Miech
Associate Professor Emeritus of Medical Science

Margaret Miller
Associate Professor of Medicine

Thomas J. Miner
Associate Professor of Surgery

Eric M. Morrow
Associate Professor of Biology; Associate Professor of Psychiatry and Human Behavior

Christopher S. Muratore
Associate Professor of Surgery; Associate Professor of Pediatrics

Aman Nanda
Associate Professor of Medicine

Anthony M. Napoli
Associate Professor of Emergency Medicine

Marguerite A. Neill
Associate Professor of Medicine

Thomas Ng
Associate Professor of Surgery (Cardiothoracic Surgery)

Elena Oancea
Associate Professor of Medical Science

Frank L. Overly
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

Gyan Pareek
Associate Professor of Surgery (Urology)

Robert L. Patrick
Associate Professor of Medical Science

Chanika Phornphutkul
Associate Professor of Pediatrics

Paul A. Pirraglia
Associate Professor of Medicine

Stephen Porder
Associate Professor of Ecology and Evolutionary Biology and Environment and Society

Kittichai Promrat
Associate Professor of Medicine

Sohini Ramachandran
Associate Professor of Ecology and Evolutionary Biology

Megan L. Ranney
Associate Professor of Emergency Medicine; Associate Professor of Health Services, Policy and Practice

Charles R. Rardin
Associate Professor of Obstetrics and Gynecology

Neha P. Raukar
Associate Professor of Emergency Medicine

Anthony M. Reginato
Associate Professor of Medicine; Associate Professor of Dermatology

Joseph F. Renzulli
Associate Professor of Surgery (Urology)

Abdalla Rifai
Associate Professor Emeritus of Pathology and Laboratory Medicine

Jeffrey M. Rogg
Associate Professor of Diagnostic Imaging

James L. Rudolph
Associate Professor of Medicine; Associate Professor of Health Services, Policy and Practice

Raymond Richard Russell
Associate Professor of Medicine

Arthur R. Salomon
Associate Professor of Biology; Associate Professor of Chemistry

Juan Sanchez-Esteban
Associate Professor of Pediatrics

Neil Sarkar
Associate Professor of Medical Science; Associate Professor of Health Services, Policy and Practice

Daniel L. Savitt
Associate Professor of Emergency Medicine

Dov F. Sax
Associate Professor of Ecology and Evolutionary Biology and Environmental Studies

Mark Schleinitz
Associate Professor of Medicine

Jigme M. Sethi
Associate Professor of Medicine

Barry L. Sharaf
Associate Professor of Medicine

Katherine M. Sharkey
Associate Professor of Medicine; Associate Professor of Psychiatry and Human Behavior

Douglas G. Shemin
Associate Professor of Medicine

Gregory M. Soares
Associate Professor of Diagnostic Imaging

Patrick K. Sullivan
Associate Professor of Surgery

Eleanor M. Summerhill
Associate Professor of Medicine

Dominick Tammaro
Associate Professor of Medicine

Diana O. Treaba
Associate Professor of Pathology and Laboratory Medicine

Geoffrey Tremont
Associate Professor of Psychiatry and Human Behavior

Robert J. Tubbs
Associate Professor of Emergency Medicine

Li Juan Wang
Associate Professor of Pathology and Laboratory Medicine

Daniel M. Weinreich
Associate Professor of Ecology and Evolutionary Biology

Marion F. Winkler
Associate Professor of Surgery

Edward G. Wittels
Associate Professor of Medicine; Associate Professor of Medical Science

Albert S. Woo
Associate Professor of Surgery (Plastic Surgery); Associate Professor of Pediatrics

Wen-Chih Wu
Associate Professor of Medicine; Associate Professor of Epidemiology

Israel Yaar
Associate Professor Emeritus of Neurology

Evgeny Yakirevich
Associate Professor of Pathology and Laboratory Medicine

Wentian Yang
Associate Professor of Medicine; Associate Professor of Orthopaedics

Cunxian Zhang
Associate Professor of Pathology and Laboratory Medicine

Richard J. Zienowicz
Associate Professor of Surgery (Plastic Surgery)

Mark Robert Zonfrillo
Associate Professor of Emergency Medicine; Associate Professor of Pediatrics

Associate Professor Clinical

Brian G. Abbott
Associate Professor of Medicine (Clinical)

Geoffrey Allen
Associate Professor of Pediatrics (Clinical)

Angela C. Anderson
Associate Professor of Pediatrics (Clinical); Associate Professor of Emergency Medicine (Clinical)

James Barbeau
Associate Professor of Pathology and Laboratory Medicine (Clinical)

Christine E. Barron
Associate Professor of Pediatrics (Clinical)

William D. Binder
Associate Professor of Emergency Medicine (Clinical)

Julie Boergers
Associate Professor of Psychiatry and Human Behavior (Clinical); Associate Professor of Pediatrics (Clinical)

Alice E. Bonitati
Associate Professor of Medicine (Clinical)

David B. Bouslough
Associate Professor of Emergency Medicine (Clinical)

Jeffrey M. Brody
Associate Professor of Diagnostic Imaging (Clinical)

David Bullard
Associate Professor of Emergency Medicine (Clinical)

Robert T. Burke
Associate Professor of Pediatrics (Clinical)

Gerardo P. Carino
Associate Professor of Medicine (Clinical)

Carolina S. Cerezo
Associate Professor of Pediatrics (Clinical)

Anjulika Chawla
Associate Professor of Pediatrics (Clinical)

Sybil Cineas
Associate Professor of Medicine (Clinical); Associate Professor of Medical Science (Clinical); Associate Professor of Pediatrics (Clinical)

Jennifer D. Davis
Associate Professor of Psychiatry and Human Behavior (Clinical)

Susan Day
Associate Professor of Pediatrics (Clinical)

Viren A. D'Sa
Associate Professor of Pediatrics (Clinical)

Gregory J. Dubel
Associate Professor of Diagnostic Imaging (Clinical)

Christine M. Duffy
Associate Professor of Medicine (Clinical)

Annmarie Dunican
Associate Professor of Surgery (Clinical)

Kevin M. Dushay
Associate Professor of Medicine (Clinical)

Afshin Ehsan
Associate Professor of Surgery (Clinical)(Cardiothoracic Surgery)

Peter T. Evangelista
Associate Professor of Diagnostic Imaging (Clinical)

M. Khurram Faizan
Associate Professor of Pediatrics (Clinical)

Jeffrey Feden
Associate Professor of Emergency Medicine (Clinical)

John L. Foggle
Associate Professor of Emergency Medicine (Clinical)

Sara R. Ford
Associate Professor of Pediatrics (Clinical)

John A. Froehlich
Associate Professor of Orthopaedics (Clinical)

Ana Tuya Fulton
Associate Professor of Medicine (Clinical); Associate Professor of Psychiatry and Human Behavior (Clinical)

Sarah A. Gaines
Associate Professor of Emergency Medicine (Clinical)

Melissa M. Gaitanis
Associate Professor of Medicine (Clinical)

Abbe L. Garcia
Associate Professor of Psychiatry and Human Behavior (Clinical)

Maria Luisa Garcia-Moliner
Associate Professor of Pathology and Laboratory Medicine (Clinical)

Bethany J. Gentilesco
Associate Professor of Medicine (Clinical); Associate Professor of Medical Science (Clinical)

Frantz J. Gibbs
Associate Professor of Emergency Medicine (Clinical)

Richard L. Gold
Associate Professor of Diagnostic Imaging (Clinical)

Amy P. Goldberg
Associate Professor of Pediatrics (Clinical)

Tracey Guthrie
Associate Professor of Psychiatry and Human Behavior (Clinical); Associate Professor of Medical Science (Clinical)

Richard Alan Haas
Associate Professor of Diagnostic Imaging (Clinical); Associate Professor of Neurology (Clinical); Associate Professor of Neurosurgery (Clinical)

Fadlallah G. Habr
Associate Professor of Medicine (Clinical)

Elisabeth D. Howard
Associate Professor of Obstetrics and Gynecology (Clinical)

Susie Lee Hu
Associate Professor of Medicine (Clinical); Associate Professor of Medical Science (Clinical)

Robert M. Insoft
Associate Professor of Pediatrics (Clinical)

Elizabeth S. Jacobs
Associate Professor of Emergency Medicine (Clinical)

Jennifer Jeremiah
Associate Professor of Medicine (Clinical)

Gary D. Johnson
Associate Professor of Neurology (Clinical)

Emily R. Katz
Associate Professor of Psychiatry and Human Behavior (Clinical); Associate Professor of Pediatrics (Clinical)

Karen L. Kerman
Associate Professor of Neurology (Clinical); Associate Professor of Pediatrics (Clinical)

Noubar Kessimian
Associate Professor of Pathology and Laboratory Medicine (Clinical)

Unab I. Khan
Associate Professor of Pediatrics (Clinical)

Michael P. Koster
Associate Professor of Pediatrics (Clinical)

Jerome M. Larkin
Associate Professor of Medicine (Clinical)

Robert Legare
Associate Professor of Obstetrics and Gynecology (Clinical); Associate Professor of Medicine (Clinical)

Carol T. Lewis
Associate Professor of Pediatrics (Clinical)

Victoria Leytin
Associate Professor of Emergency Medicine (Clinical)

David G. Lindquist
Associate Professor of Emergency Medicine (Clinical)

Gregory R. Lockhart
Associate Professor of Emergency Medicine (Clinical); Associate Professor of Pediatrics (Clinical)

Michele M. Lomme
Associate Professor of Pathology and Laboratory Medicine (Clinical)

Elizabeth A. Lowenhaupt
Associate Professor of Psychiatry and Human Behavior (Clinical); Associate Professor of Medical Science (Clinical); Associate Professor of Pediatrics (Clinical)

Edward W. Martin
Associate Professor of Medicine (Clinical)

Peter J. Mazzaglia
Associate Professor of Surgery (Clinical)

Anthony E. Mega
Associate Professor of Medicine (Clinical); Associate Professor of Surgery (Clinical)(Urology)

Niharika D. Mehta
Associate Professor of Medicine (Clinical); Associate Professor of Obstetrics and Gynecology (Clinical)

Stephen T. Mernoff
Associate Professor of Neurology (Clinical)

Jennie J. Muglia
Associate Professor of Dermatology (Clinical)

Gerard Nau
Associate Professor of Medicine (Clinical)

Ezequiel Neimark
Associate Professor of Pediatrics (Clinical)

John W. O'Bell
Associate Professor of Medicine (Clinical)

James A. O'Brien
Associate Professor of Obstetrics and Gynecology (Clinical)

Adetokunbo A. Oyelese
Associate Professor of Neurosurgery (Clinical)

Adam D. Pallant
Associate Professor of Pediatrics (Clinical)

John A. Pezzullo
Associate Professor of Diagnostic Imaging (Clinical)

Latha R. Pisharodi
Associate Professor of Pathology and Laboratory Medicine (Clinical)

Michael J. Price
Associate Professor of Radiation Oncology (Clinical)

Mark S. Ridlen
Associate Professor of Diagnostic Imaging (Clinical)

Syed A. Rizvi
Associate Professor of Neurology (Clinical)

Kathleen M. Rotondo
Associate Professor of Pediatrics (Clinical)

G. Dean Roye
Associate Professor of Surgery (Clinical)

George M. Sachs
Associate Professor of Neurology (Clinical)

Frank John Schaberg Jr
Associate Professor Emeritus of Surgery (Clinical)

Justin Schleifer
Associate Professor of Psychiatry and Human Behavior (Clinical)

Linda B. Shalon
Associate Professor of Pediatrics (Clinical)

Uzma Sharif
Associate Professor of Neurology (Clinical); Associate Professor of Pediatrics (Clinical)

William M. Sikov
Associate Professor of Medicine (Clinical); Associate Professor of Obstetrics and Gynecology (Clinical)

Jessica L. Smith
Associate Professor of Emergency Medicine (Clinical)

Julie H. Song
Associate Professor of Diagnostic Imaging (Clinical)

Daniel B. Spencer
Associate Professor of Psychiatry and Human Behavior (Clinical)

Philippa Gabrielle Sprinz
Associate Professor of Pediatrics (Clinical)

Jie Tang
Associate Professor of Medicine (Clinical)

Umadevi Tantravahi
Associate Professor of Pathology and Laboratory Medicine (Clinical)

Albert Edward Telfeian
Associate Professor of Neurosurgery (Clinical)

Marcia W. Vanvleet
Associate Professor of Pediatrics (Clinical)

Sivamainthan Vithiananthan
Associate Professor of Surgery (Clinical)

Michael T. Wallach
Associate Professor of Diagnostic Imaging (Clinical); Associate Professor of Pediatrics (Clinical)

Shunping Wang
Associate Professor of Obstetrics and Gynecology (Clinical)

Joseph Benjamin Weiss
Associate Professor of Medicine (Clinical)

Carol A. Wheeler
Associate Professor of Obstetrics and Gynecology (Clinical)

Kenneth Williams
Associate Professor of Emergency Medicine (Clinical)

Karen L. Woolfall-Quinn
Associate Professor of Medicine (Clinical)

Ali Yalcindag
Associate Professor of Pediatrics (Clinical)

Yvette E. Yatchmink
Associate Professor of Pediatrics (Clinical)

Don C. Yoo
Associate Professor of Diagnostic Imaging (Clinical)

James W. Ziegler
Associate Professor of Pediatrics (Clinical)

Clinical Associate Professor

Khaja Ahmed
Clinical Associate Professor of Medicine

Paul E. Alexander
Clinical Associate Professor of Psychiatry and Human Behavior

Marla Cathleen Angermeier
Clinical Associate Professor of Dermatology

Genie L. Bailey
Clinical Associate Professor of Psychiatry and Human Behavior

Steven Barreto
Clinical Associate Professor of Psychiatry and Human Behavior

Jacques Benun
Clinical Associate Professor of Pediatrics

Karyn K. Blane
Clinical Associate Professor of Psychiatry and Human Behavior

John R. Boekamp
Clinical Associate Professor of Psychiatry and Human Behavior

Kathleen Cote Bowling
Clinical Associate Professor of Obstetrics and Gynecology

Edward M. Brown
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior

Steven A. Brown
Clinical Associate Professor of Surgery (Oral & Maxillofacial Surgery)

Maria L. Buckley
Clinical Associate Professor of Psychiatry and Human Behavior

James Butera
Clinical Associate Professor of Medicine

Terrence F. Cahill
Clinical Associate Professor of Obstetrics and Gynecology

Nicholas A. Califano
Clinical Associate Professor of Medicine

Lauralyn B. Cannistra
Clinical Associate Professor of Medicine

Carmine J. Capalbo
Clinical Associate Professor Emeritus of Surgery

David P. Carter
Clinical Associate Professor of Family Medicine

Amos Charles
Clinical Associate Professor of Medicine

Reid W. Coleman
Clinical Associate Professor of Medicine

Francis Connor
Clinical Associate Professor of Surgery (Oral & Maxillofacial Surgery)

George Cooper
Clinical Associate Professor Emeritus of Surgery

Dominic Corrigan
Clinical Associate Professor of Medicine

Catherine A. Cummings
Clinical Associate Professor of Emergency Medicine

Karen Daigle
Clinical Associate Professor of Pediatrics

Arthur D. Daily
Clinical Associate Professor Emeritus of Dermatology

Kwame Dapaah-Afriyie
Clinical Associate Professor of Medicine

Thomas D. Denucci
Clinical Associate Professor of Medicine

Allan Mark Deutsch
Clinical Associate Professor Emeritus of Diagnostic Imaging

Elizabeth R. Didie
Clinical Associate Professor of Psychiatry and Human Behavior

Amy Bach Dilello
Clinical Associate Professor of Psychiatry and Human Behavior

Walter E. Donat
Clinical Associate Professor of Medicine

Joseph L. Dowling Jr
Clinical Associate Professor Emeritus of Surgery

Thomas Drew
Clinical Associate Professor Emeritus of Medicine

Brian E. Duff
Clinical Associate Professor of Surgery (Otolaryngology)

Charles Eil
Clinical Associate Professor of Medicine

Yul D. Ejnes
Clinical Associate Professor of Medicine

Jonathan Elion
Clinical Associate Professor of Medicine

Yoash R. Enzer
Clinical Associate Professor of Surgery; Clinical Associate Professor of Dermatology

David Ettensohn
Clinical Associate Professor of Medicine

Stephen Falkenberry
Clinical Associate Professor of Obstetrics and Gynecology

Glenn G. Fort
Clinical Associate Professor of Medicine

Arthur Frazzano
Clinical Associate Professor Emeritus of Family Medicine

Michael A. Friedman
Clinical Associate Professor of Psychiatry and Human Behavior; Clinical Associate Professor of Neurology

Martin Furman
Clinical Associate Professor of Psychiatry and Human Behavior

Prerna Ganjoo
Clinical Associate Professor of Medicine

Arthur I. Geltzer
Clinical Associate Professor of Surgery (Ophthalmology)

Thomas Germano
Clinical Associate Professor of Emergency Medicine

Sharon E. Gibson
Clinical Associate Professor of Surgery (Otolaryngology)

David R. Gifford
Clinical Associate Professor of Medicine; Clinical Associate Professor of Health Services, Policy and Practice

Michael F. Gilson
Clinical Associate Professor of Medicine

Stephen E. Glinick
Clinical Associate Professor of Dermatology

Alan L. Gordon
Clinical Associate Professor of Psychiatry and Human Behavior

Norman M. Gordon
Clinical Associate Professor of Neurology

Lisa Gould
Clinical Associate Professor of Medicine

Rod A. Gragg
Clinical Associate Professor of Psychiatry and Human Behavior

Neil R. Greenspan
Clinical Associate Professor of Medicine

Mark W. Greve
Clinical Associate Professor of Emergency Medicine

Robert T. Griffith
Clinical Associate Professor of Pediatrics

Thomas J. Haronian
Clinical Associate Professor of Emergency Medicine

Tawfik F. Hawwa
Clinical Associate Professor of Obstetrics and Gynecology

Arnold Herman
Clinical Associate Professor of Surgery

Horacio B. Hojman
Clinical Associate Professor of Psychiatry and Human Behavior

Karen A. Holler
Clinical Associate Professor of Psychiatry and Human Behavior

Karyn J. Horowitz
Clinical Associate Professor of Psychiatry and Human Behavior

Robert H. Janigian
Clinical Associate Professor of Surgery (Ophthalmology)

Ilse M. Jenouri
Clinical Associate Professor of Emergency Medicine

Robert G. M. Johnston
Clinical Associate Professor of Psychiatry and Human Behavior

Achyut Kamat
Clinical Associate Professor of Emergency Medicine

Edward Keating
Clinical Associate Professor of Medicine; Clinical Associate Professor of Medical Science

Susan M. Kelly
Clinical Associate Professor of Psychiatry and Human Behavior

Brian Andrew Kimble
Clinical Associate Professor of Medicine; Clinical Associate Professor of Medical Science

Boyd P. King
Clinical Associate Professor Emeritus of Medicine

David L. Kitzes
Clinical Associate Professor of Medicine

Robert E. Knisley
Clinical Associate Professor Emeritus of Medicine

Donna M. Lafontaine
Clinical Associate Professor of Obstetrics and Gynecology

Elizabeth Lange
Clinical Associate Professor of Pediatrics

Elizabeth A. Laposata
Clinical Associate Professor of Pathology and Laboratory Medicine

Phillip R. Lucas
Clinical Associate Professor of Orthopaedics

Susanna R. Magee
Clinical Associate Professor of Family Medicine

Henry G. Magendantz
Clinical Associate Professor Emeritus of Obstetrics and Gynecology

Tariq K. Malik
Clinical Associate Professor of Medicine

James Manis
Clinical Associate Professor Emeritus of Medicine

Louis J. Marino
Clinical Associate Professor of Psychiatry and Human Behavior

Charles E. McCoy
Clinical Associate Professor of Medicine

Paul N. McMillan
Clinical Associate Professor Emeritus of Pathology and Laboratory Medicine

A. Sattar N. Memon
Clinical Associate Professor Emeritus of Medicine

Robert D. Meringolo
Clinical Associate Professor of Medicine

Martin M. Miner
Clinical Associate Professor of Family Medicine; Clinical Associate Professor of Surgery (Urology)

Samir G. Moubayed
Clinical Associate Professor Emeritus of Obstetrics and Gynecology

Brian L. Murphy
Clinical Associate Professor of Diagnostic Imaging

James R. Myers
Clinical Associate Professor of Medicine

Judith Nudelman
Clinical Associate Professor of Family Medicine

Richard K. Ohnmacht
Clinical Associate Professor of Pediatrics

Nicklas B. E. Oldenburg
Clinical Associate Professor of Radiation Oncology

Karen Oliver
Clinical Associate Professor of Psychiatry and Human Behavior

Jay M. Orson
Clinical Associate Professor Emeritus of Pediatrics

Calvin E. Oyer
Clinical Associate Professor Emeritus of Pathology and Laboratory Medicine

Frank A. Pensa
Clinical Associate Professor of Obstetrics and Gynecology

Elliot M. Perlman
Clinical Associate Professor of Surgery (Ophthalmology)

Francis J. Pescosolido
Clinical Associate Professor of Psychiatry and Human Behavior

Barry A. Plummer
Clinical Associate Professor of Psychiatry and Human Behavior

David C. Portelli
Clinical Associate Professor of Emergency Medicine

Roy M. Poses
Clinical Associate Professor of Medicine

M. Yakub A. Puthawala
Clinical Associate Professor of Radiation Oncology

Lucille A. Rathier
Clinical Associate Professor of Psychiatry and Human Behavior

Barbara H. Roberts
Clinical Associate Professor of Medicine

Henry J. Robidoux
Clinical Associate Professor Emeritus of Surgery

Pablo Rodriguez
Clinical Associate Professor of Obstetrics and Gynecology

Albert M. Ross
Clinical Associate Professor of Pediatrics

Ara Sadaniantz
Clinical Associate Professor of Medicine

Steven Schechter
Clinical Associate Professor of Surgery

Stuart T. Schwartz
Clinical Associate Professor of Medicine

Thomas E. Sepe
Clinical Associate Professor of Medicine

Reza Shah-Hosseini
Clinical Associate Professor Emeritus of Obstetrics and Gynecology

Thomas K. Shahinian
Clinical Associate Professor of Surgery

Lisa B. Shea
Clinical Associate Professor of Psychiatry and Human Behavior

Latha Sivaprasad
Clinical Associate Professor of Medicine

Lory Snady-Mccoy
Clinical Associate Professor of Surgery (Ophthalmology)

Clarence H. Soderberg Jr
Clinical Associate Professor Emeritus of Surgery

Shirley A. Spater
Clinical Associate Professor of Surgery (Oral & Maxillofacial Surgery)

Patricia K. Spencer
Clinical Associate Professor Emerita of Diagnostic Imaging

Pamela Steadman-Wood
Clinical Associate Professor of Psychiatry and Human Behavior

Francis M. Sullivan
Clinical Associate Professor of Emergency Medicine

Frank W. Sullivan
Clinical Associate Professor Emeritus of Psychiatry and Human Behavior

James K. Sullivan
Clinical Associate Professor of Psychiatry and Human Behavior

Ghulam M. Surti
Clinical Associate Professor of Psychiatry and Human Behavior

Linda S. Tartell
Clinical Associate Professor of Pediatrics

Charlene A. Tate
Clinical Associate Professor of Neurology

David Robbins Tien
Clinical Associate Professor of Surgery (Ophthalmology)

Philip A. Torgan
Clinical Associate Professor Emeritus of Medicine

Jody Ann Underwood
Clinical Associate Professor of Psychiatry and Human Behavior

Wilson F. Utter
Clinical Associate Professor Emeritus of Pediatrics

Rendueles Villalba
Clinical Associate Professor of Psychiatry and Human Behavior

Hedy S. Wald
Clinical Associate Professor of Family Medicine

Anne S. Walters
Clinical Associate Professor of Psychiatry and Human Behavior

Banice M. Webber
Clinical Associate Professor Emeritus of Radiation Oncology

Marilyn Weigner
Clinical Associate Professor of Medicine

Robert R. Williams
Clinical Associate Professor of Obstetrics and Gynecology

Harold A. Woodcome
Clinical Associate Professor of Surgery (Ophthalmology)

Tony C. Wu
Clinical Associate Professor of Medicine

Associate Professor Research

Ana M. Abrantes
Associate Professor of Psychiatry and Human Behavior (Research)

Janette Baird
Associate Professor of Emergency Medicine (Research)

Dale S. Bond
Associate Professor of Psychiatry and Human Behavior (Research)

Linda A. Bradley
Associate Professor of Pathology and Laboratory Medicine (Research)

Chung-Ja M. Cha
Associate Professor Emerita of Pediatrics (Research)

Adam Chodobski
Associate Professor of Emergency Medicine (Research)

Bum-Rak Choi
Associate Professor of Medicine (Research)

Deborah Mck Ciombor
Associate Professor of Surgery (Research); Associate Professor of Molecular Pharmacology, Physiology and Biotechnology (Research)

Robbert J. Creton
Associate Professor of Medical Science (Research); Associate Professor of Pathology and Laboratory Medicine (Research)

Scott J. Cruikshank
Associate Professor of Neuroscience (Research)

Loren D. Fast
Associate Professor of Medicine (Research)

Jun Feng
Associate Professor of Surgery (Research)

Brandon A. Gaudiano
Associate Professor of Psychiatry and Human Behavior (Research)

Stephen H. Gregory
Associate Professor of Medicine (Research)

Kate M. Guthrie
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Behavioral and Social Sciences (Research)

Christopher D. Houck
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Pediatrics (Research)

Stephanie R. Jones
Associate Professor of Neuroscience (Research)

Miran Kim
Associate Professor of Medicine (Research)

Daphne Koinis-Mitchell
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Pediatrics (Research)

Linda L. LaGasse
Associate Professor of Pediatrics (Research); Associate Professor of Psychiatry and Human Behavior (Research)

Ji Su Li
Associate Professor of Medicine (Research)

Qing Lu
Associate Professor of Medicine (Research)

Jason T. Machan
Associate Professor of Orthopaedics (Research); Associate Professor of Surgery (Research)

Jeanne McCaffery
Associate Professor of Psychiatry and Human Behavior (Research)

Robert Miranda
Associate Professor of Psychiatry and Human Behavior (Research)

Nicole R. Nugent
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Pediatrics (Research)

Donna R. Parker
Associate Professor of Family Medicine (Research); Associate Professor of Epidemiology (Research)

Susan Ramsey
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Medicine (Research)

Carl Saab
Associate Professor of Neurosurgery (Research); Associate Professor of Neuroscience (Research)

Lori Scott-Sheldon
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Behavioral and Social Sciences (Research)

Michael F. Sheff
Associate Professor Emeritus of Pathology and Laboratory Medicine (Research)

Laura Stroud
Associate Professor of Psychiatry and Human Behavior (Research); Associate Professor of Behavioral and Social Sciences (Research)

Dmitry Terentyev
Associate Professor of Medicine (Research)

Graham Thomas
Associate Professor of Psychiatry and Human Behavior (Research)

Yi-Tang Tseng
Associate Professor of Pediatrics (Research)

Anny Usheva
Associate Professor of Surgery (Research)(Cardiothoracic Surgery)

Lei Wei
Associate Professor of Orthopaedics (Research)

Lauren M. Weinstock
Associate Professor of Psychiatry and Human Behavior (Research)

Hongwei Yao
Associate Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Shirley Yen
Associate Professor of Psychiatry and Human Behavior (Research)

Associate Professor MBL

Irina Arkhipova
Associate Professor of Molecular Biology, Cell Biology and Biochemistry (MBL)

Marko Horb
Associate Professor of Molecular Biology, Cell Biology and Biochemistry (MBL)

Julie A. Huber
Associate Professor of Ecology and Evolutionary Biology (MBL)

David Mark Welch
Associate Professor of Molecular Biology, Cell Biology and Biochemistry (MBL)

Christopher Neill
Associate Professor of Ecology and Evolutionary Biology (MBL)

Assistant Professor

Liza M. Aguiar
Assistant Professor of Surgery (Urology); Assistant Professor of Pediatrics

Jason M. Aliotta
Assistant Professor of Medicine

Valentin Antoci
Assistant Professor of Orthopaedics

Eirini Apostolidou
Assistant Professor of Medicine

Herbert D. Aronow
Assistant Professor of Medicine

Maroun Azar
Assistant Professor of Medicine

Thomas B. Bartnikas
Assistant Professor of Pathology and Laboratory Medicine

Francesca L. Beaudoin
Assistant Professor of Emergency Medicine

Peter A. Belenky
Assistant Professor of Molecular Microbiology and Immunology

Brad D. Blankenhorn
Assistant Professor of Orthopaedics

James M. Brindle
Assistant Professor of Radiation Oncology

E. Christine Brousseau
Assistant Professor of Obstetrics and Gynecology

Amy Cameron
Assistant Professor of Psychiatry and Human Behavior

Christy Capone
Assistant Professor of Psychiatry and Human Behavior

Geoffrey Capraro
Assistant Professor of Emergency Medicine

Philip A. Chan
Assistant Professor of Medicine

Kenneth K. L. Chen
Assistant Professor of Medicine; Assistant Professor of Obstetrics and Gynecology

Won-Kyung Cho
Assistant Professor of Medicine

Antony Chu
Assistant Professor of Medicine

Michael Connolly
Assistant Professor of Surgery (Trauma); Assistant Professor of Medical Science

Aristides Cruz
Assistant Professor of Orthopaedics

Shawna M. Cutting
Assistant Professor of Neurology

Alan H. Daniels
Assistant Professor of Orthopaedics

Manuel F. Dasilva
Assistant Professor of Orthopaedics

Michelle Rose Dawson
Assistant Professor of Molecular Pharmacology, Physiology and Biotechnology

Alexandra M. Deaconescu
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Christopher L. de Graffenried
Assistant Professor of Molecular Microbiology and Immunology

Theresa M. Desrochers
Assistant Professor of Neuroscience; Assistant Professor of Psychiatry and Human Behavior

Kristin H. Dwyer
Assistant Professor of Emergency Medicine

Erica Eaton
Assistant Professor of Psychiatry and Human Behavior

Staci R. Eisenberg
Assistant Professor of Pediatrics; Assistant Professor of Medicine

Christopher Elco
Assistant Professor of Pathology and Laboratory Medicine; Assistant Professor of Dermatology

Heinrich Elinzano
Assistant Professor of Neurology

Gary P. Epstein-Lubow
Assistant Professor of Psychiatry and Human Behavior; Assistant Professor of Health Services, Policy and Practice; Assistant Professor of Medical Science

Andrew R. Evans
Assistant Professor of Orthopaedics

Katherine E. Faricy-Anderson
Assistant Professor of Medicine

Nicolas Lux Fawzi
Assistant Professor of Molecular Pharmacology, Physiology and Biotechnology

Dilber E. Gamsiz
Assistant Professor of Pathology and Laboratory Medicine

Elizabeth M. Goldberg
Assistant Professor of Emergency Medicine

Christopher J. Got
Assistant Professor of Orthopaedics

Ricky Grisson
Assistant Professor of Pathology and Laboratory Medicine

Philip Haines
Assistant Professor of Medicine

Daithi J. Heffernan
Assistant Professor of Surgery

Jaroslaw T. Hepel
Assistant Professor of Radiation Oncology; Assistant Professor of Neurosurgery

Thaddeus W. Herliczek
Assistant Professor of Diagnostic Imaging

Amanda M. Jamieson
Assistant Professor of Molecular Microbiology and Immunology

Matthew D. Jankowich
Assistant Professor of Medicine

Alexander Jaworski
Assistant Professor of Neuroscience

Derek Randall Jenkins
Assistant Professor of Orthopaedics

Jennifer E. Johnson
Assistant Professor of History; Adjunct Associate Professor of Psychiatry and Human Behavior

Elizabeth Kalife
Assistant Professor of Pathology and Laboratory Medicine

Naz Karim
Assistant Professor of Emergency Medicine

Tyler R. Kartzinel
Assistant Professor of Ecology and Evolutionary Biology

Karla Kaun
Assistant Professor of Neuroscience

James R. Kellner
Assistant Professor of Ecology and Evolutionary Biology and Environment and Society

Louis R. Lapierre
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Erica N. Larschan
Richard and Edna Salomon Assistant Professor of Molecular Biology, Cell Biology and Biochemistry

Jodi L. Layton
Assistant Professor of Medicine

Beatrice E. Lechner
Assistant Professor of Pediatrics

Andrew Burgess Leslie
Assistant Professor of Ecology and Evolutionary Biology

Adam Levine
Assistant Professor of Emergency Medicine

Fenghong Liu
Assistant Professor of Radiation Oncology

Kalyan Mantripragada
Assistant Professor of Medicine

Lauren Massingham
Assistant Professor of Pediatrics

Thais P. Mather
Assistant Professor of Medical Science; Assistant Professor of Molecular Microbiology and Immunology

Cara A. Mathews
Assistant Professor of Obstetrics and Gynecology

Ryan A. McTaggart
Assistant Professor of Diagnostic Imaging; Assistant Professor of Neurology; Assistant Professor of Neurosurgery

Michelle L. Mellion
Assistant Professor of Neurology

Lisa H. Merck
Assistant Professor of Emergency Medicine; Assistant Professor of Diagnostic Imaging

Alexander Mohler
Assistant Professor of Neurology

Kristina A. Monteiro
Assistant Professor of Medical Science

Mari Mori
Assistant Professor of Pediatrics

Alan R. Morrison
Assistant Professor of Medicine

Nicola Neretti
Assistant Professor of Biology

Amanda Noska
Assistant Professor of Medicine

Adam Olszewski
Assistant Professor of Medicine

Jin O-Uchi
Assistant Professor of Medicine

Nikhil Panda
Assistant Professor of Medicine

Cherie Paquette
Assistant Professor of Pathology and Laboratory Medicine

Nimesh R. Patel
Assistant Professor of Pathology and Laboratory Medicine

E. Scott Paxton
Assistant Professor of Orthopaedics

Noah S. Philip
Assistant Professor of Psychiatry and Human Behavior

Jessica S. Plavicki
Assistant Professor of Pathology and Laboratory Medicine

N. Stevenson Potter
Assistant Professor of Neurology; Assistant Professor of Neurosurgery

Ethan A. Prince
Assistant Professor of Diagnostic Imaging

Matthew I. Quesenberry
Assistant Professor of Medicine

Aadia I. Rana
Assistant Professor of Medicine

John L. Reagan
Assistant Professor of Medicine

Katina M. Robison
Assistant Professor of Obstetrics and Gynecology

Dario R. Roque
Assistant Professor of Obstetrics and Gynecology

Julie L. Roth
Assistant Professor of Neurology; Assistant Professor of Medical Science

Susan A. Rudders
Assistant Professor of Pediatrics

Beth A. Ryder
Assistant Professor of Surgery; Assistant Professor of Medical Science

Keith Scarfo
Assistant Professor of Neurosurgery

Jonathan R. Schiller
Assistant Professor of Orthopaedics

Nicole E. Alexander-Scott
Assistant Professor of Health Services, Policy and Practice; Assistant Professor of Pediatrics; Assistant Professor of Medicine

Sadhak Sengupta
Assistant Professor of Neurosurgery

Nishant R. Shah
Assistant Professor of Health Services, Policy and Practice; Assistant Professor of Medicine

Nishit Shah
Assistant Professor of Surgery

Robert M. Shalvoy
Assistant Professor of Orthopaedics

Matthew S. Siket
Assistant Professor of Emergency Medicine

Kamaljeet Singh
Assistant Professor of Pathology and Laboratory Medicine

Shahrzad Ehdaivand Slater
Assistant Professor of Pathology and Laboratory Medicine

Michael A. Smit
Assistant Professor of Pediatrics

Neel R. Sodha
Assistant Professor of Surgery

Ashley R. Stuckey
Assistant Professor of Obstetrics and Gynecology; Assistant Professor of Medical Science

Lisa Michelle Swartz Topor
Assistant Professor of Pediatrics

Joanne P. Szczygiel
Assistant Professor of Medicine

Bradford B. Thompson
Assistant Professor of Neurology; Assistant Professor of Neurosurgery

Iris L. Tong
Assistant Professor of Medicine

Wilson Truccolo
Assistant Professor of Neuroscience

Shipra Vaishnava
Assistant Professor of Molecular Microbiology and Immunology

Corey E. Ventetuolo
Assistant Professor of Medicine; Assistant Professor of Health Services, Policy and Practice

Roxanne Vrees
Assistant Professor of Obstetrics and Gynecology

Yihong Wang
Assistant Professor of Pathology and Laboratory Medicine

Sarita S. Warrier
Assistant Professor of Medicine; Assistant Professor of Medical Science

Ashley E. Webb
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry; Assistant Professor of Neurology

Jennifer Greene Welch
Assistant Professor of Pediatrics

Linda C. Wendell
Assistant Professor of Neurology; Assistant Professor of Medical Science; Assistant Professor of Neurosurgery

Erika Werner
Assistant Professor of Obstetrics and Gynecology

Jordan C. White
Assistant Professor of Family Medicine; Assistant Professor of Medical Science

Joanne E. Wilkinson
Assistant Professor of Family Medicine

Hale E. Wills
Assistant Professor of Surgery; Assistant Professor of Pediatrics

Robyn Wing
Assistant Professor of Emergency Medicine; Assistant Professor of Pediatrics

Diana Ying Wu
Assistant Professor of Medicine

Shadi Yaghi
Assistant Professor of Neurology

Assistant Professor Clinical

Cathleen M. Adams
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Saurabh Agarwal
Assistant Professor of Diagnostic Imaging (Clinical)

Khalid A. Alhourani
Assistant Professor of Medicine (Clinical)

Tanya Ali
Assistant Professor of Medicine (Clinical)

Srividya Anandan
Assistant Professor of Diagnostic Imaging (Clinical)

Michelle Stozek Anvar
Assistant Professor of Medicine (Clinical)

David Ashley
Assistant Professor of Family Medicine (Clinical)

Nicholas Asselin
Assistant Professor of Emergency Medicine (Clinical)

Kimberly S. Babb
Assistant Professor of Medicine (Clinical); Assistant Professor of Pediatrics (Clinical)

Christina A. Bandera
Assistant Professor of Surgery (Clinical)

Amy Bass
Assistant Professor of Emergency Medicine (Clinical)

Jacob R. Berger
Assistant Professor of Neurology (Clinical)

Kristina Berglund
Assistant Professor of Medicine (Clinical)

Harikrashna Bhatt
Assistant Professor of Medicine (Clinical)

Courtney C. Bilodeau
Assistant Professor of Medicine (Clinical)

Ariel Birnbaum
Assistant Professor of Medicine (Clinical)

Kenneth D. Bishop
Assistant Professor of Medicine (Clinical)

Laura Celeste Black
Assistant Professor of Medicine (Clinical)

Christina Alexandra Blum
Assistant Professor of Neurology (Clinical)

Elizabeth Brannan
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Rimini Breakstone
Assistant Professor of Medicine (Clinical)

Dayna Burrell
Assistant Professor of Obstetrics and Gynecology (Clinical)

Jonathan F. Cahill
Assistant Professor of Neurology (Clinical)

Rachael L. Caiati
Assistant Professor of Diagnostic Imaging (Clinical)

Luisa Cala Cala
Assistant Professor of Pediatrics (Clinical)

Cassandra L. Carberry
Assistant Professor of Obstetrics and Gynecology (Clinical)

Stephanie Cardarella
Assistant Professor of Medicine (Clinical)

Eden R. Cardozo
Assistant Professor of Obstetrics and Gynecology (Clinical)

Chelsy Caren
Assistant Professor of Obstetrics and Gynecology (Clinical)

John A. Cassese
Assistant Professor of Diagnostic Imaging (Clinical); Assistant Professor of Pediatrics (Clinical)

Tamuna Chadashvili
Assistant Professor of Diagnostic Imaging (Clinical)

Abdullah Chahin
Assistant Professor of Medicine (Clinical)

Sandy Chai
Assistant Professor of Dermatology (Clinical)

Laura L. Chapman
Assistant Professor of Emergency Medicine (Clinical)

Vicky Cheng
Assistant Professor of Medicine (Clinical)

Lydia R. Ciarallo
Assistant Professor of Emergency Medicine (Clinical); Assistant Professor of Pediatrics (Clinical)

Daniel T. Coghlin
Assistant Professor of Pediatrics (Clinical)

Erika Constantine
Assistant Professor of Emergency Medicine (Clinical); Assistant Professor of Pediatrics (Clinical)

Delaney Conway
Assistant Professor of Medicine (Clinical)

Ricardo Correa
Assistant Professor of Medicine (Clinical)

Thomas F. Crain
Assistant Professor of Medicine (Clinical)

Patricia A. Cristofaro
Assistant Professor of Medicine (Clinical)

Beth Cronin
Assistant Professor of Obstetrics and Gynecology (Clinical)

Adam J. Czynski
Assistant Professor of Pediatrics (Clinical)

Tanya Dailey
Assistant Professor of Obstetrics and Gynecology (Clinical)

Deepan Dalal
Assistant Professor of Medicine (Clinical)

Lawrence M. Davis
Assistant Professor of Diagnostic Imaging (Clinical)

Sarah M. Davis
Assistant Professor of Obstetrics and Gynecology (Clinical)

Gina M. Deck
Assistant Professor of Neurology (Clinical)

Diane DerMarderosian
Assistant Professor of Pediatrics (Clinical)

John Bartlett Dickey
Assistant Professor of Medicine (Clinical)

Christopher Dimarco
Assistant Professor of Dermatology (Clinical)

Kimberly A. Dodd
Assistant Professor of Medicine (Clinical)

Abigail Donaldson
Assistant Professor of Pediatrics (Clinical)

Linda L. Donegan
Assistant Professor of Diagnostic Imaging (Clinical)

Kathleen R. Donise
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Kristina M. Duarte
Assistant Professor of Family Medicine (Clinical)

Luba Dumenco
Assistant Professor of Medical Science (Clinical); Assistant Professor of Pathology and Laboratory Medicine (Clinical)

Laura M. Edmonds
Assistant Professor of Medicine (Clinical)

David Edmonson
Assistant Professor of Surgery (Clinical); Assistant Professor of Obstetrics and Gynecology (Clinical)

Renee Ross Eger
Assistant Professor of Obstetrics and Gynecology (Clinical)

Sadie Elisseou
Assistant Professor of Medicine (Clinical)

Fadya El Rayess
Assistant Professor of Family Medicine (Clinical)

Shannon Marie Erisman
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Matthew A. Esposito
Assistant Professor of Obstetrics and Gynecology (Clinical)

Samuel K. Evans
Assistant Professor of Medicine (Clinical)

Dimitrios Farmakiotis
Assistant Professor of Medicine (Clinical)

Anna Filip
Assistant Professor of Family Medicine (Clinical)

Whitney Charles Fisher
Assistant Professor of Emergency Medicine (Clinical)

Rachel L. Fowler
Assistant Professor of Emergency Medicine (Clinical)

Sarah Freeman
Assistant Professor of Medicine (Clinical)

Katharine French
Assistant Professor of Medicine (Clinical)

Christopher M. Furey
Assistant Professor of Family Medicine (Clinical)

Heather Gale
Assistant Professor of Diagnostic Imaging (Clinical)

Joseph Garland
Assistant Professor of Medicine (Clinical)

Ana C. Garnecho
Assistant Professor of Pediatrics (Clinical)

Seth Gemme
Assistant Professor of Emergency Medicine (Clinical)

Boris Gershman
Assistant Professor of Surgery (Clinical)(Urology)

Arnoldas Giedrimas
Assistant Professor of Medicine (Clinical)

Jeanette G. Giedrimas
Assistant Professor of Medicine (Clinical)

Holly C. Gil
Assistant Professor of Diagnostic Imaging (Clinical)

Courtney N. Gleason
Assistant Professor of Orthopaedics (Clinical)

Bracha Goldsweig
Assistant Professor of Pediatrics (Clinical)

Natalia Golova
Assistant Professor of Pediatrics (Clinical)

Dina Gozman
Assistant Professor of Emergency Medicine (Clinical)

Theresa A. Graves
Assistant Professor of Surgery (Clinical)

Nicole L. Grenier
Assistant Professor of Dermatology (Clinical)

Jan C. Groblewski
Assistant Professor of Surgery (Clinical)(Otolaryngology); Assistant Professor of Pediatrics (Clinical)

Geetika Gupta
Assistant Professor of Medicine (Clinical)

Preeti Gupta
Assistant Professor of Neurology (Clinical); Assistant Professor of Medical Science (Clinical)

Sarah E. Hagin
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Mohamad Hamdi
Assistant Professor of Obstetrics and Gynecology (Clinical)

Erica J. Hardy
Assistant Professor of Medicine (Clinical)

Ross Hilliard
Assistant Professor of Medicine (Clinical)

Mary M. Hillstrom
Assistant Professor of Diagnostic Imaging (Clinical)

Nadine T. Himelfarb
Assistant Professor of Emergency Medicine (Clinical)

Dawn B. Hogan
Assistant Professor of Family Medicine (Clinical)

Mary H. Hohenhaus
Assistant Professor of Medicine (Clinical)

Nathan Hudepohl
Assistant Professor of Emergency Medicine (Clinical)

Neha Hudepohl
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Zilla H. Hussain
Assistant Professor of Medicine (Clinical)

Omar N. Hyder
Assistant Professor of Medicine (Clinical)

Sarah Marie Hyder
Assistant Professor of Medicine (Clinical)

Hafiz Muhammad Imran
Assistant Professor of Medicine (Clinical)

Randall Ingham
Assistant Professor of Medicine (Clinical)

Judy L. Jang
Assistant Professor of Medicine (Clinical)

Thomas W. Jean
Assistant Professor of Medicine (Clinical)

Gaurav Jindal
Assistant Professor of Diagnostic Imaging (Clinical)

Shuba Kamath
Assistant Professor of Pediatrics (Clinical)

Vania Kasper
Assistant Professor of Pediatrics (Clinical)

May M. Kassem
Assistant Professor of Diagnostic Imaging (Clinical)

Alexis Kearney
Assistant Professor of Emergency Medicine (Clinical)

Bridget Kelly
Assistant Professor of Obstetrics and Gynecology (Clinical)

Hanan I. Khalil
Assistant Professor of Diagnostic Imaging (Clinical)

Humera Khurshid
Assistant Professor of Medicine (Clinical)

Susan L. Koelliker
Assistant Professor of Diagnostic Imaging (Clinical)

Nicole Korbly
Assistant Professor of Obstetrics and Gynecology (Clinical)

Margaret K. Kozel
Assistant Professor of Pediatrics (Clinical)

Robin A. Kremsdorf
Assistant Professor of Pediatrics (Clinical)

Peter K. Kriz
Assistant Professor of Orthopaedics (Clinical); Assistant Professor of Pediatrics (Clinical)

Donald Labbe
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Ashley Lakin
Assistant Professor of Family Medicine (Clinical)

Vita Lamberson
Assistant Professor of Pediatrics (Clinical)

Candace S. Lapidus
Assistant Professor of Medical Science (Clinical)

Joseph R. Lauro
Assistant Professor of Emergency Medicine (Clinical)

Athene Ka Wing Lee
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Kachiu Lee
Assistant Professor of Dermatology (Clinical)

Mark P. Legolvan
Assistant Professor of Pathology and Laboratory Medicine (Clinical)

Emily A. Leveen
Assistant Professor of Medicine (Clinical)

Scott M. Levine
Assistant Professor of Diagnostic Imaging (Clinical)

Melanie J. Lippmann
Assistant Professor of Emergency Medicine (Clinical)

Kristin Lombardi
Assistant Professor of Pediatrics (Clinical)

Stephanie N. Lueckel
Assistant Professor of Surgery (Clinical)

Natalia Lukankina
Assistant Professor of Pathology and Laboratory Medicine (Clinical)

Jessica C. Manyan
Assistant Professor of Family Medicine (Clinical)

Kamran Manzoor
Assistant Professor of Medicine (Clinical)

Douglas W. Martin
Assistant Professor of Medicine (Clinical)

Augusto Rafael Martinez
Assistant Professor of Pediatrics (Clinical)

Donnah Mathews
Assistant Professor of Medicine (Clinical)

Kristina McAteer
Assistant Professor of Emergency Medicine (Clinical)

Robin L. McKinney
Assistant Professor of Pediatrics (Clinical)

Joseph M. McNamara
Assistant Professor of Pediatrics (Clinical)

Laura H. McPeake
Assistant Professor of Emergency Medicine (Clinical)

Basma Merhi
Assistant Professor of Medicine (Clinical)

Stephen J. Migliori
Assistant Professor of Surgery (Clinical)

Srilakshmi Mitta
Assistant Professor of Medicine (Clinical)

Debra J. Moorhead
Assistant Professor of Family Medicine (Clinical)

Jonathan S. Movson
Assistant Professor of Diagnostic Imaging (Clinical)

Nadia Mujahid
Assistant Professor of Medicine (Clinical)

Judy I. Murphy
Assistant Professor of Medicine (Clinical)

Nicholas Musisca
Assistant Professor of Emergency Medicine (Clinical)

Drew Nagle
Assistant Professor of Medicine (Clinical)

Deepa Nankani
Assistant Professor of Medicine (Clinical)

David P. Neumann
Assistant Professor of Diagnostic Imaging (Clinical)

Van T. Nguyen
Assistant Professor of Diagnostic Imaging (Clinical)

Julia E. Noguchi
Assistant Professor of Medical Science (Clinical)

Kara Nugent
Assistant Professor of Medicine (Clinical)

Zsolt Orban
Assistant Professor of Medicine (Clinical)

So Yeon Paek
Assistant Professor of Dermatology (Clinical)

Suzanne M. Palinski
Assistant Professor of Pediatrics (Clinical)

Marcelle L. Piccolello
Assistant Professor of Diagnostic Imaging (Clinical)

Francine Pingitore
Assistant Professor of Pediatrics (Clinical); Assistant Professor of Psychiatry and Human Behavior (Clinical)

Lee Alexander Polikoff
Assistant Professor of Pediatrics (Clinical)

Jane Preotle
Assistant Professor of Emergency Medicine (Clinical)

Sakeena Raza
Assistant Professor of Medicine (Clinical)

Rebecca Reece
Assistant Professor of Medicine (Clinical)

Steven K. Reich
Assistant Professor of Pediatrics (Clinical)

Katherine Richman
Assistant Professor of Medicine (Clinical)

Jeffrey S. Riese
Assistant Professor of Pediatrics (Clinical)

Tina Rizack
Assistant Professor of Medicine (Clinical); Assistant Professor of Obstetrics and Gynecology (Clinical)

Jael Rodriguez
Assistant Professor of Medicine (Clinical)

Jamison E. Rogers
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Tanya Rogo
Assistant Professor of Pediatrics (Clinical)

Noah K. Rosenberg
Assistant Professor of Emergency Medicine (Clinical)

Fred A. Rotenberg
Assistant Professor of Surgery (Clinical)(Anesthesiology)

Leslie A. Roth
Assistant Professor of Surgery (Clinical)

Amity Rubeor
Assistant Professor of Family Medicine (Clinical)

Maureen Ryall
Assistant Professor of Pediatrics (Clinical)

Natasha R. Rybak
Assistant Professor of Medicine (Clinical)

Noha Sadek
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Martha Cristina Sanchez
Assistant Professor of Medicine (Clinical)

Janice Santos Cortes
Assistant Professor of Surgery (Clinical)(Urology)

Andrea Sartori
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Zabe R. Sayeed
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Albert A. Scappaticci
Assistant Professor of Diagnostic Imaging (Clinical)

Margaret Scheffler
Assistant Professor of Pediatrics (Clinical)

Lauren K. Schlanger
Assistant Professor of Medicine (Clinical)

Carl Schwartz
Assistant Professor of Surgery (Clinical)(Anesthesiology)

Jonathan Hale Season
Assistant Professor of Emergency Medicine (Clinical)

Sairah Sharif
Assistant Professor of Medicine (Clinical)

Laura Shawhughes
Assistant Professor of Medicine (Clinical)

Jared A. Silverstein
Assistant Professor of Pediatrics (Clinical)

Michael Sisitsky
Assistant Professor of Obstetrics and Gynecology (Clinical)

Jeffrey M. Slaiby
Assistant Professor of Surgery (Clinical)

Brett Slingsby
Assistant Professor of Pediatrics (Clinical)

Victoria Snegovskikh
Assistant Professor of Obstetrics and Gynecology (Clinical)

Amy L. Snyder
Assistant Professor of Obstetrics and Gynecology (Clinical)

Christopher Song
Assistant Professor of Medicine (Clinical)

Peter Soukas
Assistant Professor of Medicine (Clinical)

Jensy Stafford
Assistant Professor of Medicine (Clinical); Assistant Professor of Family Medicine (Clinical)

Lars Stangenberg
Assistant Professor of Surgery (Clinical)

Kara Stavros
Assistant Professor of Neurology (Clinical)

Andrew H. Stephen
Assistant Professor of Surgery (Clinical)

Ryan Stevenson
Assistant Professor of Medicine (Clinical)

Mariah Stump
Assistant Professor of Medicine (Clinical)

Patrick Sullivan
Assistant Professor of Emergency Medicine (Clinical)

Rachel A. Sullivan
Assistant Professor of Surgery (Clinical)(Plastic Surgery)

David W. Swenson
Assistant Professor of Diagnostic Imaging (Clinical)

Angela M. Taber
Assistant Professor of Medicine (Clinical); Assistant Professor of Medical Science (Clinical)

Lynn E. Taylor
Assistant Professor of Medicine (Clinical)

Christopher Tessier
Assistant Professor of Medicine (Clinical)

Francine Touzard Romo
Assistant Professor of Medicine (Clinical)

Gregory J. Towne
Assistant Professor of Surgery (Clinical)(Anesthesiology)

Andrew S. Triebwasser
Assistant Professor of Surgery (Clinical)(Anesthesiology)

Tovah Tripp
Assistant Professor of Medicine (Clinical)

Margaret Tryforos
Assistant Professor of Family Medicine (Clinical)

Robert Y. Velasco
Assistant Professor of Medicine (Clinical)

Alice-Lee Vestner
Assistant Professor of Psychiatry and Human Behavior (Clinical); Assistant Professor of Medical Science (Clinical)

Lidia A. Vognar
Assistant Professor of Medicine (Clinical)

Joanna Walker
Assistant Professor of Dermatology (Clinical)

Robert Ward
Assistant Professor of Diagnostic Imaging (Clinical)

Otis U. Warren
Assistant Professor of Emergency Medicine (Clinical)

Matthew D. Willis
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Kyle J. Wohlrab
Assistant Professor of Obstetrics and Gynecology (Clinical)

Pei-Chi Wu
Assistant Professor of Pediatrics (Clinical)

Shirley Wu
Assistant Professor of Emergency Medicine (Clinical)

Rayan Yousefzai
Assistant Professor of Medicine (Clinical)

Kate M. Zaluski
Assistant Professor of Obstetrics and Gynecology (Clinical)

Natalie A. Zervas
Assistant Professor of Psychiatry and Human Behavior (Clinical)

Clinical Assistant Professor

Christopher A. Abadi
Clinical Assistant Professor of Medicine

Alyn L. Adrain
Clinical Assistant Professor of Medicine

David C. Ahern
Clinical Assistant Professor of Psychiatry and Human Behavior

Irfan Ahmad
Clinical Assistant Professor of Medicine

Paul A. Akerman
Clinical Assistant Professor of Medicine; Clinical Assistant Professor of Surgery

Ali Akhtar
Clinical Assistant Professor of Medicine

Holly L. Alexandre
Clinical Assistant Professor of Family Medicine

Ahmad Al-Raqqad
Clinical Assistant Professor of Medicine

Elizabeth L. Altenhein
Clinical Assistant Professor of Surgery

Meredith Amaya-Hodges
Clinical Assistant Professor of Psychiatry and Human Behavior

Seema Amin
Clinical Assistant Professor of Medicine

Jennifer L. Anthony
Clinical Assistant Professor of Psychiatry and Human Behavior

Rex W. Appenfeller
Clinical Assistant Professor of Medicine

Denise M. Arcand
Clinical Assistant Professor of Family Medicine

Michel A. Arcand
Clinical Assistant Professor of Orthopaedics

Richard C. Archambault
Clinical Assistant Professor of Psychiatry and Human Behavior

Andrea E. Arena
Clinical Assistant Professor of Family Medicine

Naureen Attiullah
Clinical Assistant Professor of Psychiatry and Human Behavior

James M. Badger
Clinical Assistant Professor of Psychiatry and Human Behavior

Robert L. Bahr
Clinical Assistant Professor of Surgery (Ophthalmology)

Kevin E. Baill
Clinical Assistant Professor of Psychiatry and Human Behavior

Priya Banerjee
Clinical Assistant Professor of Pathology and Laboratory Medicine

Mohammad Banki
Clinical Assistant Professor of Surgery

Jonathan Barker
Clinical Assistant Professor of Psychiatry and Human Behavior

Keith Richard Bartolomei
Clinical Assistant Professor of Medicine

Orazio J. Basile
Clinical Assistant Professor Emeritus of Dermatology

Josef Bautista
Clinical Assistant Professor of Medicine

Solmaz Behtash
Clinical Assistant Professor of Family Medicine

M. David Beitle
Clinical Assistant Professor of Obstetrics and Gynecology

Michael J. Belanger
Clinical Assistant Professor of Orthopaedics

Christina Bellanti
Clinical Assistant Professor of Psychiatry and Human Behavior

Matthew D. Belley
Clinical Assistant Professor of Radiation Oncology

Natalia Belosloudtseva
Clinical Assistant Professor of Medicine

Irene Belsky
Clinical Assistant Professor of Family Medicine

Talya Benoff
Clinical Assistant Professor of Psychiatry and Human Behavior

Elizabeth Benz
Clinical Assistant Professor of Surgery

Scott E. Benzuly
Clinical Assistant Professor of Surgery (Anesthesiology)

Nathan Barry Beraha
Clinical Assistant Professor of Pediatrics

Arnaldo Berges
Clinical Assistant Professor of Psychiatry and Human Behavior

Douglas B. Bernon
Clinical Assistant Professor of Psychiatry and Human Behavior

Richard G. Bertini
Clinical Assistant Professor Emeritus of Orthopaedics

Jonathan M. Bertman
Clinical Assistant Professor of Family Medicine

Michael A. Bharier
Clinical Assistant Professor of Dermatology

Reena A. Bhatt
Clinical Assistant Professor of Surgery (Plastic Surgery)

David Bica
Clinical Assistant Professor of Family Medicine

Karen Blackmer
Clinical Assistant Professor of Family Medicine

Dawna A. Blake
Clinical Assistant Professor of Medicine

Steven L. Blazar
Clinical Assistant Professor of Orthopaedics

Sheenagh M. Bodkin
Clinical Assistant Professor of Medicine

Paul J. Botelho
Clinical Assistant Professor of Surgery (Ophthalmology)

Lisa D. Bowie
Clinical Assistant Professor of Family Medicine

Amy M. Boyer
Clinical Assistant Professor of Medicine

Lisa D. Boyle
Clinical Assistant Professor of Obstetrics and Gynecology

Michelle A. Boyle
Clinical Assistant Professor of Family Medicine

Nancy H. Brand
Clinical Assistant Professor of Psychiatry and Human Behavior

Peter C. Brasch
Clinical Assistant Professor of Surgery (Ophthalmology)

Allyson D. Brathwaite Gardner
Clinical Assistant Professor of Psychiatry and Human Behavior

James A. Brcak
Clinical Assistant Professor of Psychiatry and Human Behavior

Paul Breiding
Clinical Assistant Professor of Medicine

Patricia Bresky
Clinical Assistant Professor of Psychiatry and Human Behavior

Cheryl Brodsky
Clinical Assistant Professor of Obstetrics and Gynecology

Rajvi Broker-Sen
Clinical Assistant Professor of Psychiatry and Human Behavior

Daniel C. Brown
Clinical Assistant Professor of Family Medicine

Joanna D. Brown
Clinical Assistant Professor of Family Medicine

Matthew A. Brumbaugh
Clinical Assistant Professor of Medicine

Nicholas P. Bruno
Clinical Assistant Professor of Dermatology

Richard G. Bryan
Clinical Assistant Professor of Surgery (Ophthalmology)

Andrew E. Burchard
Clinical Assistant Professor of Surgery (Otolaryngology)

Michael D. Burgard
Clinical Assistant Professor of Psychiatry and Human Behavior

Jeffrey M. Burock
Clinical Assistant Professor of Psychiatry and Human Behavior

Douglas M. Burtt
Clinical Assistant Professor of Medicine

Melissa A. Buttaro
Clinical Assistant Professor of Psychiatry and Human Behavior

Joseph M. Caiati
Clinical Assistant Professor of Surgery (Anesthesiology)

Joseph F. Callaghan
Clinical Assistant Professor Emeritus of Surgery (Urology)

Karen T. Cammuso
Clinical Assistant Professor of Psychiatry and Human Behavior

Christopher A. Campagnari
Clinical Assistant Professor of Family Medicine

Anthony J. Cannistra
Clinical Assistant Professor of Medicine

Joseph E. Caruolo
Clinical Assistant Professor Emeritus of Surgery

Colleen Cavanaugh
Clinical Assistant Professor of Obstetrics and Gynecology

Timothy Cavanaugh
Clinical Assistant Professor of Family Medicine

Joseph Centofanti
Clinical Assistant Professor of Neurology

Stephen L. Chabot
Clinical Assistant Professor of Family Medicine

John A. Chaffey
Clinical Assistant Professor of Medicine

Karmela Kim Chan
Clinical Assistant Professor of Medicine

Manchin Chang
Clinical Assistant Professor of Medicine

Heather A. Chapman
Clinical Assistant Professor of Pediatrics

Yael Chatav Schonbrun
Clinical Assistant Professor of Psychiatry and Human Behavior

Elizabeth Chattillion
Clinical Assistant Professor of Psychiatry and Human Behavior

Ahmed A. Chaudhary
Clinical Assistant Professor of Radiation Oncology

Nicole Christian-Brathwaite
Clinical Assistant Professor of Psychiatry and Human Behavior

David J. Chronley
Clinical Assistant Professor of Pediatrics

Mark Andrew Clark
Clinical Assistant Professor of Family Medicine

David R. Cloutier
Clinical Assistant Professor of Surgery

Steven Ira Cohen
Clinical Assistant Professor of Surgery

Steven Irvin Cohen
Clinical Assistant Professor of Surgery (Urology)

Mauro Colavita
Clinical Assistant Professor of Obstetrics and Gynecology

Bradley J. Collins
Clinical Assistant Professor of Medicine

Samantha Collum
Clinical Assistant Professor of Psychiatry and Human Behavior

Jeannine M. Connolly
Clinical Assistant Professor of Obstetrics and Gynecology

Maria Constantinou
Clinical Assistant Professor of Medicine

John M. Conte
Clinical Assistant Professor of Medicine

Alfredo C. Cordova
Clinical Assistant Professor of Surgery

Giselle A. Corre
Clinical Assistant Professor of Psychiatry and Human Behavior

Joseph A. Costa
Clinical Assistant Professor of Family Medicine

Geanina O. Costea
Clinical Assistant Professor of Psychiatry and Human Behavior

Philippe Cote
Clinical Assistant Professor of Family Medicine

Travis Cotton
Clinical Assistant Professor of Surgery

Maria Coutinho
Clinical Assistant Professor of Psychiatry and Human Behavior

Hugh P. Cowdin
Clinical Assistant Professor of Surgery (Anesthesiology)

Norbert J. Craybas
Clinical Assistant Professor of Surgery

Marjorie L. Crozier
Clinical Assistant Professor of Psychiatry and Human Behavior

Dragos Cucu
Clinical Assistant Professor of Medicine

Michael L. Cummings
Clinical Assistant Professor of Pediatrics

Stephen Cummings
Clinical Assistant Professor of Family Medicine

Marlene Cutitar
Clinical Assistant Professor of Surgery

Stephanie Jo Czech
Clinical Assistant Professor of Psychiatry and Human Behavior

John K. Czerwein
Clinical Assistant Professor of Orthopaedics

Catherine D'Avanzato
Clinical Assistant Professor of Psychiatry and Human Behavior

Sarah Davenport
Clinical Assistant Professor of Family Medicine

Kristopher Davignon
Clinical Assistant Professor of Surgery (Anesthesiology)

Laura Deihl
Clinical Assistant Professor of Psychiatry and Human Behavior

Paul V. Del Guercio
Clinical Assistant Professor of Family Medicine

Thomas D. Della Torre
Clinical Assistant Professor of Family Medicine

Allen M. Dennison
Clinical Assistant Professor of Medicine

Judith Depue
Clinical Professor Emerita of Psychiatry and Human Behavior

Christy Dibble
Clinical Assistant Professor of Medicine

Joseph Di Benedetto
Clinical Assistant Professor of Medicine

Margaret A. Dicarlo
Clinical Assistant Professor of Psychiatry and Human Behavior

Dinusha W. Dietrich
Clinical Assistant Professor of Pediatrics

Joseph Dimase
Clinical Assistant Professor Emeritus of Medicine

John Diorio
Clinical Assistant Professor of Obstetrics and Gynecology

Stephen Dizio
Clinical Assistant Professor of Psychiatry and Human Behavior

Lisa Ruth Domagalski
Clinical Assistant Professor of Obstetrics and Gynecology

John P. Donahue
Clinical Assistant Professor of Surgery (Ophthalmology)

Lindsay E. Donat
Clinical Assistant Professor of Obstetrics and Gynecology

Edmund T. Dosremedios
Clinical Assistant Professor of Orthopaedics

Richard J. Doyle
Clinical Assistant Professor of Medical Science; Clinical Assistant Professor of Family Medicine

Melanie Dubard
Clinical Assistant Professor of Psychiatry and Human Behavior

George J. Du Pont
Clinical Assistant Professor of Surgery (Oral & Maxillofacial Surgery)

Robert K. Dyer
Clinical Assistant Professor of Dermatology

Robert Eden
Clinical Assistant Professor of Pediatrics

Joely V. Edwards
Clinical Assistant Professor of Medical Science

Lana G. Elhalabi
Clinical Assistant Professor of Psychiatry and Human Behavior

Salaheldin Elhamamsy
Clinical Assistant Professor of Medicine

Peter M. Eller
Clinical Assistant Professor of Medicine

Martin Elson
Clinical Assistant Professor of Surgery

Joseph J. England
Clinical Assistant Professor of Family Medicine

Cynthia Esteban
Clinical Assistant Professor of Pediatrics

Elkin O. Estrada
Clinical Assistant Professor of Medicine

Frank Faltus
Clinical Assistant Professor of Psychiatry and Human Behavior

Elisabeth K. Farnum
Clinical Assistant Professor of Family Medicine

Leslie Ann Feil
Clinical Assistant Professor of Psychiatry and Human Behavior

Sarah J. Fessler
Clinical Assistant Professor of Family Medicine

John W. Finigan
Clinical Assistant Professor of Pediatrics

Joshua E. Fischer
Clinical Assistant Professor of Medicine

Walter D. Fitzhugh
Clinical Assistant Professor of Psychiatry and Human Behavior

Ellen Flynn
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Medicine

Julie Forrest
Clinical Assistant Professor of Psychiatry and Human Behavior

Gregory A. Fox
Clinical Assistant Professor of Pediatrics

Laura L. Frakey
Clinical Assistant Professor of Psychiatry and Human Behavior

Caren L. Francione
Clinical Assistant Professor of Psychiatry and Human Behavior

Heather M. Frank
Clinical Assistant Professor of Psychiatry and Human Behavior

Steven Frank
Clinical Assistant Professor of Family Medicine

Lisa M. Freda
Clinical Assistant Professor of Psychiatry and Human Behavior

Katherine E. Frias
Clinical Assistant Professor of Medical Science

David L. Fried
Clinical Assistant Professor of Medicine

Joseph R. Gaeta
Clinical Assistant Professor Emeritus of Medicine

Harris M. Galkin
Clinical Assistant Professor of Obstetrics and Gynecology

Michele R. Gange
Clinical Assistant Professor of Obstetrics and Gynecology

John A. Gelzhiser
Clinical Assistant Professor of Medicine

Emily Gentes
Clinical Assistant Professor of Psychiatry and Human Behavior

Jonathan Gershon
Clinical Assistant Professor of Psychiatry and Human Behavior

Richard J. Gilbert
Clinical Assistant Professor of Family Medicine

Peter S. Gill
Clinical Assistant Professor of Surgery

Peter Joseph Gillen
Clinical Assistant Professor of Psychiatry and Human Behavior

Athina Giovanis
Clinical Assistant Professor of Family Medicine

William Arthur Golden
Clinical Assistant Professor of Medicine

Ariel Goldschmidt
Clinical Assistant Professor of Pathology and Laboratory Medicine

Lisa J. Goldstein
Clinical Assistant Professor of Pathology and Laboratory Medicine

Vinay Goyal
Clinical Assistant Professor of Family Medicine

Robyn A. Gray
Clinical Assistant Professor of Obstetrics and Gynecology

David M. Greenberg
Clinical Assistant Professor of Surgery

Nicole E. Greenwood
Clinical Assistant Professor of Family Medicine

Nicholas Grumbach
Clinical Assistant Professor of Medicine; Clinical Assistant Professor of Pediatrics

Gaurav Gupta
Clinical Assistant Professor of Surgery (Ophthalmology)

Joshua D. Gutman
Clinical Assistant Professor of Family Medicine

Thomas E. Guttmacher
Clinical Assistant Professor of Family Medicine

Amy R. Halt
Clinical Assistant Professor of Psychiatry and Human Behavior

Geoffrey R. Hamilton
Clinical Assistant Professor of Medicine

Cynthia M. Hanna
Clinical Assistant Professor of Obstetrics and Gynecology

Uma Harinarayanan
Clinical Assistant Professor of Medical Science

Emily C. Harrison
Clinical Assistant Professor of Family Medicine

Terry E. Harrison-Goldman
Clinical Assistant Professor of Psychiatry and Human Behavior

Pamela A. Harrop
Clinical Assistant Professor of Medicine

Stephanie Hartselle
Clinical Assistant Professor of Psychiatry and Human Behavior

Sadaf Hashmi
Clinical Assistant Professor of Psychiatry and Human Behavior

Eugene H. Healey
Clinical Assistant Professor Emeritus of Surgery

David Herec
Clinical Assistant Professor of Medicine

Jennifer Herren
Clinical Assistant Professor of Psychiatry and Human Behavior

Melvin Hershkowitz
Clinical Assistant Professor Emeritus of Medicine

Ellen B. Hight
Clinical Assistant Professor of Family Medicine

Laurence M. Hirshberg
Clinical Assistant Professor of Psychiatry and Human Behavior

Lynn T. Ho
Clinical Assistant Professor of Family Medicine

Laurie Hoffman
Clinical Assistant Professor of Pediatrics

Erin Hogan
Clinical Assistant Professor of Medicine

Jamie Hollenbeck
Clinical Assistant Professor of Psychiatry and Human Behavior

Peter Amberg Hollmann
Clinical Assistant Professor of Family Medicine

Darren Holowka
Clinical Assistant Professor of Psychiatry and Human Behavior

Cathleen S. Hood
Clinical Assistant Professor of Family Medicine

John F. X. Horan
Clinical Assistant Professor of Family Medicine

James P. Hoye
Clinical Assistant Professor of Family Medicine

Kelly L. Hoye
Clinical Assistant Professor of Family Medicine

Heather L. Hunter
Clinical Assistant Professor of Psychiatry and Human Behavior

Brett Hurteau
Clinical Assistant Professor of Family Medicine

Harry Iannotti
Clinical Assistant Professor of Surgery (Urology)

Lynn E. Iler
Clinical Assistant Professor of Dermatology

James Ingraham
Clinical Assistant Professor of Psychiatry and Human Behavior

Marc A. Jaffe
Clinical Assistant Professor of Obstetrics and Gynecology

Bryan S. Jay
Clinical Assistant Professor of Diagnostic Imaging

Tarek Jazaerly
Clinical Assistant Professor of Pathology and Laboratory Medicine

Nathaniel J. Jellinek
Clinical Assistant Professor of Dermatology

Carol J. Jensen
Clinical Assistant Professor of Psychiatry and Human Behavior

Sharon Mullane Jezard
Clinical Assistant Professor of Family Medicine

Eileen G. Johnson
Clinical Assistant Professor of Psychiatry and Human Behavior

Michael P. Johnson
Clinical Assistant Professor of Medicine

Plakyil J. Joseph
Clinical Assistant Professor of Medicine

Annmarie Jurczak
Clinical Assistant Professor of Obstetrics and Gynecology

Amr Kader
Clinical Assistant Professor of Obstetrics and Gynecology

David Kadmon
Clinical Assistant Professor of Family Medicine

Samuel Kagan
Clinical Assistant Professor of Family Medicine

Sewell I. Kahn
Clinical Assistant Professor Emeritus of Medicine

Brett D. Kalmowitz
Clinical Assistant Professor of Medicine

Amber Kaltenstein
Clinical Assistant Professor of Psychiatry and Human Behavior

Aziza Azadali Kamani
Clinical Assistant Professor of Medicine

Elena Kamenetsky
Clinical Assistant Professor Emerita of Neurology

Joshua S. Kane
Clinical Assistant Professor of Psychiatry and Human Behavior

Barbara T. Kao
Clinical Assistant Professor of Pediatrics

David Kaplan
Clinical Assistant Professor of Emergency Medicine

Donald G. Kaufman
Clinical Assistant Professor Emeritus of Medicine

Craig Kaufmann
Clinical Assistant Professor of Psychiatry and Human Behavior

W. Scott Keigwin
Clinical Assistant Professor of Family Medicine

Martin J. Kerzer
Clinical Assistant Professor of Family Medicine

Marianna Kessimian
Clinical Assistant Professor of Psychiatry and Human Behavior

Anita Susan Kestin
Clinical Assistant Professor of Family Medicine

Pervez A. Khatib
Clinical Assistant Professor of Family Medicine

Razib Khaund
Clinical Assistant Professor of Medicine

Kerri Kim
Clinical Assistant Professor of Psychiatry and Human Behavior

Malcolm M. Kirk
Clinical Assistant Professor of Medicine

Jennifer E. Kittler
Clinical Assistant Professor of Psychiatry and Human Behavior

Dale Keith Klatzker
Clinical Assistant Professor of Psychiatry and Human Behavior

Christopher Ryan Klaus
Clinical Assistant Professor of Family Medicine

Erika Klein
Clinical Assistant Professor of Obstetrics and Gynecology

Michael D. Klein
Clinical Assistant Professor of Family Medicine

Adam Klipfel
Clinical Assistant Professor of Surgery

Cindy E. Klipfel
Clinical Assistant Professor of Pediatrics

Margaret G. Klitzke
Clinical Assistant Professor of Psychiatry and Human Behavior

Uma Kolli
Clinical Assistant Professor of Family Medicine

R. James Koness
Clinical Assistant Professor of Surgery

Renu O. Kothari
Clinical Assistant Professor of Psychiatry and Human Behavior

Altug Koymen
Clinical Assistant Professor of Obstetrics and Gynecology

Thomas A. Krahn
Clinical Assistant Professor of Medicine

Naomi R. Kramer
Clinical Assistant Professor of Medicine

Andrea Kretzschmar
Clinical Assistant Professor of Psychiatry and Human Behavior

Marina Kuperman-Beade
Clinical Assistant Professor of Dermatology

Jayne A. Kurkjian-Siegel
Clinical Assistant Professor of Psychiatry and Human Behavior

Daniel K. Kwan
Clinical Assistant Professor of Surgery

John V. Ladetto
Clinical Assistant Professor of Medicine

Alexis Nicole Lamb
Clinical Assistant Professor of Psychiatry and Human Behavior

Jennifer F. Lambert
Clinical Assistant Professor of Psychiatry and Human Behavior

Chad E. Lamendola
Clinical Assistant Professor of Family Medicine

Lesley Landau
Clinical Assistant Professor of Medical Science

Thomas P. Lang
Clinical Assistant Professor of Surgery (Ophthalmology)

Gina La Prova
Clinical Assistant Professor of Family Medicine

Scott D. Lauermann
Clinical Assistant Professor of Family Medicine

Benjamin Lederer
Clinical Assistant Professor of Psychiatry and Human Behavior

George Lee
Clinical Assistant Professor of Medicine

Jonathan K. Lee
Clinical Assistant Professor of Psychiatry and Human Behavior

Lawrence W. Lee
Clinical Assistant Professor of Family Medicine

Ruby Lee
Clinical Assistant Professor of Psychiatry and Human Behavior

Sung-Hee Rhim Lee
Clinical Assistant Professor of Obstetrics and Gynecology

Michelle J. Lefebvre
Clinical Assistant Professor of Pediatrics

Carole Lengyel
Clinical Assistant Professor of Pediatrics

Laura Levine
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Medical Science

Todd P. Levine
Clinical Assistant Professor of Psychiatry and Human Behavior

Sarah Simone Levy
Clinical Assistant Professor of Surgery (Ophthalmology)

Jonah M. Licht
Clinical Assistant Professor of Medicine

Rashmi Shetty Licht
Clinical Assistant Professor of Surgery (Urology)

Kristin N. Lichtenberg
Clinical Assistant Professor of Family Medicine

David P. Lichtenstein
Clinical Assistant Professor of Psychiatry and Human Behavior

Stacey P. Lievense
Clinical Assistant Professor of Obstetrics and Gynecology

James Lippincott
Clinical Assistant Professor of Family Medicine

Nancy T. Littell
Clinical Assistant Professor of Medicine

Salvatore J. Loporchio
Clinical Assistant Professor of Surgery (Ophthalmology)

Maranatha E. Lougheed
Clinical Assistant Professor of Family Medicine

Melissa Ludwig
Clinical Assistant Professor of Psychiatry and Human Behavior

Krista Lussier
Clinical Assistant Professor of Psychiatry and Human Behavior

Joanna V. Maclean
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Medicine

James F. Maguire
Clinical Assistant Professor of Pediatrics

James O. Maher
Clinical Assistant Professor of Orthopaedics

Marta B. Majczak
Clinical Assistant Professor of Psychiatry and Human Behavior

Sterling L. Malish
Clinical Assistant Professor of Medicine

Pierre R. Manzo
Clinical Assistant Professor of Family Medicine

Beth G. Marcaccio
Clinical Assistant Professor of Obstetrics and Gynecology

John R. Marcaccio
Clinical Assistant Professor Emeritus of Surgery (Urology)

David A. Marcoux
Clinical Assistant Professor of Medicine

Saul A. Martin
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Mario I. Martinez
Clinical Assistant Professor of Pediatrics

Christopher M. Matkovic
Clinical Assistant Professor of Psychiatry and Human Behavior

Thomas McCauley
Clinical Assistant Professor of Surgery (Ophthalmology)

Katherine Donovan McCleary
Clinical Assistant Professor of Family Medicine

Karen L. Mcgoldrick
Clinical Assistant Professor of Obstetrics and Gynecology

John McGonigle
Clinical Assistant Professor of Family Medicine

Kevin M. McKay
Clinical Assistant Professor of Psychiatry and Human Behavior

Rebecca McLean
Clinical Assistant Professor of Psychiatry and Human Behavior

Robert G. McRae
Clinical Assistant Professor of Surgery (Otolaryngology)

Carroll A. Medeiros
Clinical Assistant Professor of Obstetrics and Gynecology

Ildiko Medve
Clinical Assistant Professor of Medicine

Alisa J. Merolli
Clinical Assistant Professor of Medicine

E. Miller
Clinical Assistant Professor of Surgery (Urology)

Kimberly V. Miller
Clinical Assistant Professor of Surgery (Ophthalmology)

Marsha Miller
Clinical Assistant Professor of Family Medicine

Steven Milman
Clinical Assistant Professor of Surgery

Diane E. Minasian
Clinical Assistant Professor of Family Medicine

Jane Miniutti
Clinical Assistant Professor of Family Medicine

John P. Miskovsky
Clinical Assistant Professor of Medicine

Cristina L. Mitchell
Clinical Assistant Professor of Family Medicine

Jennifer Adelson Mitty
Clinical Assistant Professor of Medicine

Sean F. Monaghan
Clinical Assistant Professor of Surgery

Douglas C. Moore
Clinical Assistant Professor of Orthopaedics

Carla C. Moreira
Clinical Assistant Professor of Surgery

Julie Morris
Clinical Assistant Professor of Psychiatry and Human Behavior

John Rollin Morton
Clinical Assistant Professor of Family Medicine

Susannah L. Mozley
Clinical Assistant Professor of Psychiatry and Human Behavior

Jon A. Mukand
Clinical Assistant Professor of Orthopaedics

Donald R. Murphy
Clinical Assistant Professor of Family Medicine

Emily E. Murphy
Clinical Assistant Professor of Psychiatry and Human Behavior

John A. Murphy
Clinical Assistant Professor Emeritus of Obstetrics and Gynecology

Aye Myint-u
Clinical Assistant Professor of Psychiatry and Human Behavior

Jennifer Judd Nadeau
Clinical Assistant Professor of Psychiatry and Human Behavior

Yehia Mazen Naga
Clinical Assistant Professor of Medicine

Christopher Shawn Nasin
Clinical Assistant Professor of Family Medicine

Dawn K. Nero
Clinical Assistant Professor of Psychiatry and Human Behavior

Laura S. Nevel
Clinical Assistant Professor of Obstetrics and Gynecology

Chad P. Nevola
Clinical Assistant Professor of Pediatrics

Christopher Newton
Clinical Assistant Professor of Surgery (Ophthalmology)

Peter T. Nigri
Clinical Assistant Professor of Surgery (Otolaryngology)

Arthur W. Noel
Clinical Assistant Professor of Diagnostic Imaging

Jill A. O'Brien
Clinical Assistant Professor of Medicine

Patricia M. Ogera
Clinical Assistant Professor of Pathology and Laboratory Medicine

Kathleen A. O'Heelan
Clinical Assistant Professor of Family Medicine

Alvaro Olivares
Clinical Assistant Professor of Psychiatry and Human Behavior

Warren Ong
Clinical Assistant Professor of Psychiatry and Human Behavior

Roberto Ortiz
Clinical Assistant Professor of Medicine; Clinical Assistant Professor of Pathology and Laboratory Medicine

Wendy A. Ossman
Clinical Assistant Professor of Psychiatry and Human Behavior

Margaret Paccione-Dyszlewski
Clinical Assistant Professor of Psychiatry and Human Behavior

Rafael E. Padilla
Clinical Assistant Professor of Surgery (Anesthesiology)

Frank Paletta
Clinical Assistant Professor of Surgery (Oral & Maxillofacial Surgery)

Stephen V. Panaro
Clinical Assistant Professor of Surgery (Anesthesiology)

Altin Pani
Clinical Assistant Professor of Surgery (Ophthalmology)

Stacey Pappas
Clinical Assistant Professor of Family Medicine

George Pasquarello
Clinical Assistant Professor of Family Medicine

Khyati Patel
Clinical Assistant Professor of Medicine

Samir M. Patel
Clinical Assistant Professor of Psychiatry and Human Behavior

Mark Raymond Paulos
Clinical Assistant Professor of Medicine

David W. Pearson
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Heather Pelletier
Clinical Assistant Professor of Psychiatry and Human Behavior

Michael A. Pepi
Clinical Assistant Professor of Obstetrics and Gynecology

Daniel Perli
Clinical Assistant Professor of Medicine

Heidi H. Peterson
Clinical Assistant Professor of Family Medicine

Victor A. Pinkes
Clinical Assistant Professor of Emergency Medicine

Martha A. Pizzarello
Clinical Assistant Professor of Obstetrics and Gynecology

Peter A. Pizzarello
Clinical Assistant Professor of Family Medicine

Jose R. Polanco
Clinical Assistant Professor of Medicine

David S. Pomerantz
Clinical Assistant Professor of Dermatology

Michael A. Pomerantz
Clinical Assistant Professor of Medicine

Michael T. Poshkus
Clinical Assistant Professor of Medicine

Marylin M. Powers
Clinical Assistant Professor of Obstetrics and Gynecology

Ann Back Price
Clinical Assistant Professor of Psychiatry and Human Behavior

Sherri Provencal
Clinical Assistant Professor of Psychiatry and Human Behavior

Albert J. Puerini
Clinical Assistant Professor of Family Medicine

Sharath Puttichanda
Clinical Assistant Professor of Psychiatry and Human Behavior

C. Brandon Qualls
Clinical Assistant Professor of Psychiatry and Human Behavior

Dale F. Radka
Clinical Assistant Professor of Psychiatry and Human Behavior

Keith W. L. Rafal
Clinical Assistant Professor of Family Medicine

Richmond Ramirez
Clinical Assistant Professor of Family Medicine

Donald A. Ramos
Clinical Assistant Professor of Obstetrics and Gynecology

Roger D. Raymond
Clinical Assistant Professor of Medicine

Christine E. Rayner
Clinical Assistant Professor of Psychiatry and Human Behavior

James R. Rayner
Clinical Assistant Professor of Emergency Medicine

Jana S. Rice
Clinical Assistant Professor of Diagnostic Imaging

John Riedel
Clinical Assistant Professor of Emergency Medicine

Mark A. Ringiewicz
Clinical Assistant Professor of Family Medicine

Dannie Ritchie
Clinical Assistant Professor of Family Medicine

David Rivera
Clinical Assistant Professor of Surgery (Ophthalmology)

Jorge Rivera
Clinical Assistant Professor of Surgery (Ophthalmology)

James F. Robertson
Clinical Assistant Professor of Medicine

Marcia B. Robitaille
Clinical Assistant Professor of Emergency Medicine

Jodi Roque
Clinical Assistant Professor of Family Medicine

Karen J. Rosen
Clinical Assistant Professor of Psychiatry and Human Behavior

Wilma S. F. Rosen
Clinical Assistant Professor Emerita of Psychiatry and Human Behavior

Mindy S. Rosenbloom
Clinical Assistant Professor of Psychiatry and Human Behavior

Michael G. Ross
Clinical Assistant Professor of Medicine

Steven C. Roth
Clinical Assistant Professor of Family Medicine

Richard J. Ruggieri
Clinical Assistant Professor of Medicine

Charles M. Ruhl
Clinical Assistant Professor of Surgery (Otolaryngology)

Lorna B. Russell
Clinical Assistant Professor of Medicine

Philip Christian Russell
Clinical Assistant Professor of Psychiatry and Human Behavior

Patricia M. Russo-Magno
Clinical Assistant Professor of Medicine

Walid S. Saber
Clinical Assistant Professor of Medicine

Henry T. Sachs
Clinical Assistant Professor of Psychiatry and Human Behavior

Caitlin Saint-Aubin
Clinical Assistant Professor of Obstetrics and Gynecology

Matthew T. Salisbury
Clinical Assistant Professor of Family Medicine

Robert A. Salk
Clinical Assistant Professor of Obstetrics and Gynecology

Philip Salko
Clinical Assistant Professor of Family Medicine

Jason Salter
Clinical Assistant Professor of Family Medicine

David A. Sam
Clinical Assistant Professor of Family Medicine

Kelley Sanzen
Clinical Assistant Professor of Medicine

Paul E. Sapir
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Thomas J. Scaramella
Clinical Assistant Professor Emeritus of Psychiatry and Human Behavior

Monica J. Schaberg
Clinical Assistant Professor Emerita of Pediatrics

Jay S. Schachne
Clinical Assistant Professor of Medicine

Scott T. Schmidt
Clinical Assistant Professor of Surgery (Plastic Surgery)

Ronald Schneider
Clinical Assistant Professor of Dermatology

Howard E. Schulman
Clinical Assistant Professor of Medicine

Eric B. Schwam
Clinical Assistant Professor of Emergency Medicine

James M. Schwartz
Clinical Assistant Professor of Family Medicine

Gregg J. Selke
Clinical Assistant Professor of Psychiatry and Human Behavior

Bahram Shah-Hosseini
Clinical Assistant Professor of Obstetrics and Gynecology

Edward P. Sheehan
Clinical Assistant Professor of Psychiatry and Human Behavior

Thomas Sheeran
Clinical Assistant Professor of Psychiatry and Human Behavior

Cullen Shipman
Clinical Assistant Professor of Family Medicine

Priscilla Shube
Clinical Assistant Professor of Family Medicine

Afreen Siddiqui
Clinical Assistant Professor of Surgery (Anesthesiology)

Santina L. Siena
Clinical Assistant Professor Emerita of Obstetrics and Gynecology

Michael A. Silver
Clinical Assistant Professor of Psychiatry and Human Behavior

Joanne M. Silvia
Clinical Assistant Professor of Family Medicine

Louis R. Simeone
Clinical Assistant Professor of Orthopaedics

Meera Varma Simoes
Clinical Assistant Professor of Obstetrics and Gynecology

Caroline Skudlarek-Prete
Clinical Assistant Professor of Obstetrics and Gynecology

John G. Slattery
Clinical Assistant Professor of Family Medicine

Matthew Joseph Smith
Clinical Assistant Professor of Family Medicine

Richard M. Smith
Clinical Assistant Professor of Psychiatry and Human Behavior

Romina P. Smulever
Clinical Assistant Professor of Psychiatry and Human Behavior

Shalini Solanki
Clinical Assistant Professor of Family Medicine

Patricia M. Solga
Clinical Assistant Professor of Orthopaedics

Jon S. Solis
Clinical Assistant Professor of Dermatology

Joel B. Solomon
Clinical Assistant Professor of Psychiatry and Human Behavior

John J. Solomon
Clinical Assistant Professor of Family Medicine

Sukanya Somasundar
Clinical Assistant Professor of Family Medicine

Jennifer M. Souza
Clinical Assistant Professor of Family Medicine

Dana Sparhawk
Clinical Assistant Professor of Emergency Medicine

Joel S. Spellun
Clinical Assistant Professor of Medicine

Megan B. Spencer
Clinical Assistant Professor of Psychiatry and Human Behavior

Kenneth F. Sperber
Clinical Assistant Professor of Family Medicine

Emily B. Spurrell
Clinical Assistant Professor of Psychiatry and Human Behavior

Laura Stanton
Clinical Assistant Professor of Psychiatry and Human Behavior

Brett S. Stecker
Clinical Assistant Professor of Family Medicine

Steven H. Stein
Clinical Assistant Professor of Family Medicine

Robin Lisa Stern
Clinical Assistant Professor of Psychiatry and Human Behavior

Angela Stewart
Clinical Assistant Professor of Psychiatry and Human Behavior

Elizabeth Loring Sullivan
Clinical Assistant Professor of Obstetrics and Gynecology

Margaret A. Sun
Clinical Assistant Professor of Family Medicine

Jeffrey B. Syme
Clinical Assistant Professor of Family Medicine

Erica M. Szabados
Clinical Assistant Professor of Neurology

Ramin Tabaddor
Clinical Assistant Professor of Family Medicine

Pedro F. Tactacan
Clinical Assistant Professor of Psychiatry and Human Behavior

John M. Tarro
Clinical Assistant Professor of Surgery (Otolaryngology)

Anthony G. Thomas
Clinical Assistant Professor of Medicine

Edward S. Thomas
Clinical Assistant Professor of Medicine

Tenny J. Thomas
Clinical Assistant Professor of Emergency Medicine

Valerie A. Thomas
Clinical Assistant Professor of Medicine

Alysha D. Thompson
Clinical Assistant Professor of Psychiatry and Human Behavior

Sarah Thurman
Clinical Assistant Professor of Radiation Oncology

Robert Tilton
Clinical Assistant Professor of Psychiatry and Human Behavior

Cristina Toba
Clinical Assistant Professor of Psychiatry and Human Behavior

John F. Todaro
Clinical Assistant Professor of Psychiatry and Human Behavior

Lyudmil Todorov
Clinical Assistant Professor of Family Medicine

David R. Tomlinson
Clinical Assistant Professor of Family Medicine

Tanya B. C. Tran
Clinical Assistant Professor of Psychiatry and Human Behavior

Erica R. Tsai
Clinical Assistant Professor of Psychiatry and Human Behavior

Jean E. Twomey
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Pediatrics

Jamsheed Vakharia
Clinical Assistant Professor of Surgery

Tanya Vishnevsky
Clinical Assistant Professor of Psychiatry and Human Behavior

Matthew D. Vrees
Clinical Assistant Professor of Surgery

Safa F. Wagdi
Clinical Assistant Professor of Surgery (Ophthalmology)

Eric Ferguson Walsh
Clinical Assistant Professor of Orthopaedics

Michael R. Walther
Clinical Assistant Professor of Psychiatry and Human Behavior

Susan D. Weinman
Clinical Assistant Professor of Psychiatry and Human Behavior

Elizabeth A. Welch
Clinical Assistant Professor of Dermatology

Ryan J. Welter
Clinical Assistant Professor of Family Medicine

Lee V. Wesner
Clinical Assistant Professor of Surgery (Anesthesiology)

Judith B. Westrick
Clinical Assistant Professor of Pediatrics

Gary G. Wharton
Clinical Assistant Professor of Obstetrics and Gynecology

Doreen L. Wiggins
Clinical Assistant Professor of Surgery; Clinical Assistant Professor of Obstetrics and Gynecology

Caroline S. Wilkel
Clinical Assistant Professor of Dermatology

Jeffrey M. Wilson
Clinical Assistant Professor of Family Medicine

Julie M. Wilson
Clinical Assistant Professor of Psychiatry and Human Behavior

Patricia N. Wold
Clinical Assistant Professor Emerita of Psychiatry and Human Behavior

Hugh Woolverton
Clinical Assistant Professor of Family Medicine

Claudia F. Yellin
Clinical Assistant Professor of Psychiatry and Human Behavior

Agustin Yip
Clinical Assistant Professor of Psychiatry and Human Behavior; Clinical Assistant Professor of Health Services, Policy and Practice

Carolyn Te Young
Clinical Assistant Professor of Pathology and Laboratory Medicine

Mark Young
Clinical Assistant Professor of Family Medicine

Najam A. Zaidi
Clinical Assistant Professor of Medicine

Assistant Professor Research

Youngwook Ahn
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Nagib Ahsan
Assistant Professor of Medicine (Research)

Sarah A. Arias
Assistant Professor of Psychiatry and Human Behavior (Research)

Michael F. Armey
Assistant Professor of Psychiatry and Human Behavior (Research)

Grayson L. Baird
Assistant Professor of Diagnostic Imaging (Research)

David H. Barker
Assistant Professor of Psychiatry and Human Behavior (Research)

Kristen Benito
Assistant Professor of Psychiatry and Human Behavior (Research)

Bahar Bilgen
Assistant Professor of Orthopaedics (Research)

Erika L. Bloom
Assistant Professor of Psychiatry and Human Behavior (Research); Assistant Professor of Medicine (Research)

Christina L. Boisseau
Assistant Professor of Psychiatry and Human Behavior (Research)

Devasis Chatterjee
Assistant Professor of Medicine (Research)

Xiaodi Chen
Assistant Professor of Pediatrics (Research)

Paul P. Christopher
Assistant Professor of Psychiatry and Human Behavior (Research)

Chun-Shiang Chung
Assistant Professor of Surgery (Research)

Kasey R. Claborn
Assistant Professor of Medicine (Research); Assistant Professor of Psychiatry and Human Behavior (Research)

Richard Clements
Assistant Professor of Surgery (Research)

Mia Coetzer
Assistant Professor of Medicine (Research)

Lori Daiello
Assistant Professor of Neurology (Research); Assistant Professor of Health Services, Policy and Practice (Research)

Valery A. Danilack
Assistant Professor of Epidemiology (Research); Assistant Professor of Obstetrics and Gynecology (Research)

Aaron Mark Drucker
Assistant Professor of Dermatology (Research)

Yovanska Duarte-Velez
Assistant Professor of Psychiatry and Human Behavior (Research)

Patrycja Dubielecka-Szczerba
Assistant Professor of Medicine (Research)

Cara Fuchs
Assistant Professor of Psychiatry and Human Behavior (Research)

Helen Fuchs
Assistant Professor of Medicine (Research)

Senthil Kumar Ganesan
Assistant Professor of Molecular Pharmacology, Physiology and Biotechnology (Research)

Gretchen V. Gee
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Fizza S. Gillani
Assistant Professor of Medicine (Research)

Andrea Goldschmidt
Assistant Professor of Psychiatry and Human Behavior (Research)

Carolina L. Haass-Koffler
Assistant Professor of Behavioral and Social Sciences (Research); Assistant Professor of Psychiatry and Human Behavior (Research)

Wendy S. Hadley
Assistant Professor of Psychiatry and Human Behavior (Research)

Sheila A. Haley
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Chiung-Kuei Huang
Assistant Professor of Medicine (Research)

Sandra Japuntich
Assistant Professor of Psychiatry and Human Behavior (Research)

Chathuraka T. Jayasuriya
Assistant Professor of Orthopaedics (Research)

Euy-Myoung Jeong
Assistant Professor of Medicine (Research)

Beth A. Jerskey
Assistant Professor of Psychiatry and Human Behavior (Research)

Anatoli Kabakov
Assistant Professor of Medicine (Research)

Jerome Lionel Kalifa
Assistant Professor of Medicine (Research)

Rebecca Y. Kartzinel
Assistant Professor of Ecology and Evolutionary Biology (Research)

Kathleen Kemp
Assistant Professor of Psychiatry and Human Behavior (Research)

Kyu Kwang Kim
Assistant Professor of Obstetrics and Gynecology (Research)

Jill A. Kreiling
Assistant Professor of Molecular Biology, Cell Biology and Biochemistry (Research)

Han-Kyu Lee
Assistant Professor of Neurology (Research)

Craig T. Lefort
Assistant Professor of Surgery (Research)

Ming Li
Assistant Professor of Medicine (Research)

Jason Lillis
Assistant Professor of Psychiatry and Human Behavior (Research)

Hong Liu
Assistant Professor of Medicine (Research)

Man Liu
Assistant Professor of Medicine (Research)

Joanne L. Lomas-Neira
Assistant Professor of Surgery (Research)

Maria C. Mancebo
Assistant Professor of Psychiatry and Human Behavior (Research)

Abigail Marcaccio
Assistant Professor of Psychiatry and Human Behavior (Research)

Emily McDonald
Assistant Professor of Pediatrics (Research)

Nicole McLaughlin
Assistant Professor of Psychiatry and Human Behavior (Research)

Derek Merck
Assistant Professor of Diagnostic Imaging (Research); Assistant Professor of Engineering (Research)

Mandar T. Naik
Assistant Professor of Molecular Pharmacology, Physiology and Biotechnology (Research)

Kimberly M. Nelson
Assistant Professor of Psychiatry and Human Behavior (Research)

Rebecca Newland
Assistant Professor of Psychiatry and Human Behavior (Research)

Lindsay M. Orchowski
Assistant Professor of Psychiatry and Human Behavior (Research)

Rohan Palmer
Assistant Professor of Psychiatry and Human Behavior (Research)

Rebecca K. Papas
Assistant Professor of Psychiatry and Human Behavior (Research)

Stephanie Parade
Assistant Professor of Psychiatry and Human Behavior (Research)

William Keun Chan Park
Assistant Professor of Diagnostic Imaging (Research)

Megan Pinkston-Camp
Assistant Professor of Psychiatry and Human Behavior (Research)

Barbara Porton
Assistant Professor of Psychiatry and Human Behavior (Research)

Jared M. Saletin
Assistant Professor of Psychiatry and Human Behavior (Research)

Jennifer A. Sanders
Assistant Professor of Pediatrics (Research); Assistant Professor of Pathology and Laboratory Medicine (Research)

Heather T. Schatten
Assistant Professor of Psychiatry and Human Behavior (Research)

Christoph Schorl
Assistant Professor of Biology (Research)

Sunil K. Shaw
Assistant Professor of Pediatrics (Research)

Stephanie A. Shepard-Umaschi
Assistant Professor of Psychiatry and Human Behavior (Research)

Rebecca B. Silver
Assistant Professor of Psychiatry and Human Behavior (Research); Assistant Professor of Pediatrics (Research)

Katherine F. Smith
Assistant Professor of Ecology and Evolutionary Biology (Research)

Joanna Szmydynger-Chodobska
Assistant Professor of Emergency Medicine (Research)

Xiaoli Tang
Assistant Professor of Medicine (Research)

Lori A. Underhill
Assistant Professor of Pediatrics (Research)

Jacob Vandenberg
Assistant Professor of Behavioral and Social Sciences (Research); Assistant Professor of Medicine (Research)

Eliza Van Reen
Assistant Professor of Psychiatry and Human Behavior (Research)

Mascha van 't Wout
Assistant Professor of Psychiatry and Human Behavior (Research)

Sara Vargas
Assistant Professor of Psychiatry and Human Behavior (Research)

Carlos E. Vargas-Irwin
Assistant Professor of Neuroscience (Research)

Annette E. Von Dem Bussche
Assistant Professor of Pathology and Laboratory Medicine (Research)

Edward G. Walsh
Assistant Professor of Neuroscience (Research)

Xiaotian Wang
Assistant Professor of Surgery (Research)(Plastic Surgery)

Sicheng Wen
Assistant Professor of Medicine (Research)

Timothy J. Whitfeld
Assistant Professor of Ecology and Evolutionary Biology (Research)

Jennifer C. Wolff
Assistant Professor of Psychiatry and Human Behavior (Research)

Michael S. Worden
Assistant Professor of Neuroscience (Research)

An Xie
Assistant Professor of Medicine (Research)

Peng Zhang
Assistant Professor of Medicine (Research)

Anyu Zhou
Assistant Professor of Medicine (Research)

Assistant Professor MBL

Sangshin Park
Assistant Professor of Pediatrics (MBL)

Senior Lecturer

Richard D. Bungiro
Senior Lecturer in Molecular Microbiology and Immunology

Monica Linden
Senior Lecturer in Neuroscience

Dale A. Ritter
Senior Lecturer in Biology

John J. Stein
Senior Lecturer in Neuroscience

Lecturer

Diana M. Horrigan
Lecturer in Molecular Pharmacology, Physiology and Biotechnology

Babette Taylor Stewart
Lecturer Emerita in Medical Science

Instructor

Muhammad Baig
Instructor in Medicine

Debasree Banerjee
Instructor in Medicine

Prakash Kandel
Instructor in Medicine

Brian Kavanaugh
Instructor in Psychiatry and Human Behavior

Vijairam Selvaraj
Instructor in Medicine

Instructor Clinical

Amer AlSamman
Instructor in Medicine (Clinical)

Lauren Michele Catalano
Instructor in Medicine (Clinical)

Sabina Holland
Instructor in Pediatrics (Clinical)

Ghazi G. Hussain
Instructor in Medicine (Clinical)

Aravind Kokkirala
Instructor in Medicine (Clinical)

Erin McKnight
Instructor in Medicine (Clinical)

Jharendra Rijal
Instructor in Medicine (Clinical)

Sylvester Sarfraz
Instructor in Medicine (Clinical)

Clinical Instructor

Sarah Anis
Clinical Instructor in Surgery (Ophthalmology)

Daniel K. Asiedu
Clinical Instructor in Medicine

Muhammad Bajwa
Clinical Instructor in Medicine

William C. Bastan
Clinical Instructor in Pathology and Laboratory Medicine

Kim A. Basu
Clinical Instructor in Medicine

Geoffrey H. Berg
Clinical Instructor in Medicine

Navaneel Biswas
Clinical Instructor in Medicine

Douglas L. Blecker
Clinical Instructor in Medicine

Cheryl Elizabeth Bline
Clinical Instructor in Surgery (Anesthesiology)

Cynthia Bliss
Clinical Instructor in Pediatrics

Evan Burke
Clinical Instructor in Surgery (Anesthesiology)

Danica Buzniak
Clinical Instructor in Medicine

John Cava
Clinical Instructor in Medicine

Dana Chofay
Clinical Instructor in Medicine

Sapna Chowdhry
Clinical Instructor in Medicine

Nicole A. Coleman
Clinical Instructor in Emergency Medicine

Douglas J. Colson
Clinical Instructor in Surgery (Otolaryngology)

Kathryn Coolidge
Clinical Instructor in Medicine

A. Michael Coppa
Clinical Instructor in Obstetrics and Gynecology

Suzanne C. Dambek
Clinical Instructor in Obstetrics and Gynecology

Laura H. Dawson
Clinical Instructor in Pediatrics

Ashok Raj Devkota
Clinical Instructor in Medicine

Paula Deyoung
Clinical Instructor in Obstetrics and Gynecology

Sara DiNardo
Clinical Instructor in Surgery

Ibrahim Elgabry
Clinical Instructor in Medicine

Tina M. Ellis
Clinical Instructor in Surgery (Anesthesiology)

Rinchen-Tzo Emgushov
Clinical Instructor in Medicine

Martin P. Feldman
Clinical Instructor Emeritus in Surgery

Bruce E. Fischer
Clinical Instructor in Medicine

Barbara Miller Gaines
Clinical Instructor in Pediatrics

Veenu Garewal
Clinical Instructor in Medicine

Lauren A. Goddard
Clinical Instructor in Family Medicine

Amy Goldfarb
Clinical Instructor in Medicine

Emily Goodwin
Clinical Instructor in Surgery (Anesthesiology)

Carmen Goojha
Clinical Instructor in Family Medicine

Carin A. Got
Clinical Instructor in Surgery (Anesthesiology)

Michael Gottlieb
Clinical Instructor in Obstetrics and Gynecology

Samantha Noel Greenberg
Clinical Instructor in Family Medicine

Barbara J. Guillette
Clinical Instructor in Surgery (Otolaryngology)

Katherine Hart
Clinical Instructor in Pediatrics

Jeffrey D. Horowitz
Clinical Instructor in Medicine

Adriana M. Hosu
Clinical Instructor in Medicine

Raneem Islam
Clinical Instructor in Emergency Medicine

Katherine Alexandra Jarrell
Clinical Instructor in Family Medicine

Maen Kamal
Clinical Instructor in Medicine

Kabita Karki
Clinical Instructor in Medicine

Elias Khawam
Clinical Instructor in Medicine

Suvasini Lakshmanan
Clinical Instructor in Medicine

Gary K. Lundstrom
Clinical Instructor in Diagnostic Imaging

Kerri Luzzo
Clinical Instructor in Family Medicine

Nayana Manjunath
Clinical Instructor in Obstetrics and Gynecology

Vincent F. Marcaccio
Clinical Instructor in Medicine

Daniel Toby Marwil
Clinical Instructor in Pediatrics

Elizabeth Mayhall
Clinical Instructor in Obstetrics and Gynecology

Andrew P. McGrath
Clinical Instructor in Pediatrics

Mary Minn
Clinical Instructor in Surgery (Anesthesiology)

Howard M. Mintz
Clinical Instructor in Pediatrics

Michael A. Moffat
Clinical Instructor in Pediatrics; Clinical Instructor in Medical Science

Leslie C. Mohlman
Clinical Instructor in Pediatrics

Namrata Nandakumar
Clinical Instructor in Surgery (Ophthalmology)

Christopher D. Palmer
Clinical Instructor in Medicine

Sonali Pandya
Clinical Instructor in Surgery

Robert K. Parker
Clinical Instructor in Surgery

Dhiren Patel
Clinical Instructor in Family Medicine

Alfred A. Paul
Clinical Instructor in Surgery (Ophthalmology)

Jessica Pineda
Clinical Instructor in Family Medicine

Gabriel J. Pleasants
Clinical Instructor in Family Medicine

Richard A. Regnante
Clinical Instructor in Medicine

Anthony R. Ricci
Clinical Instructor in Medicine

Zachary Robbins
Clinical Instructor in Surgery (Anesthesiology)

Marina Rodriguez
Clinical Instructor in Medicine

Katherine Rotker
Clinical Instructor in Surgery (Urology)

Hayley Ryan
Clinical Instructor in Family Medicine

Ammar Saati
Clinical Instructor in Medicine

David Schreiber
Clinical Instructor in Medicine

Holly Schroeder
Clinical Instructor in Pediatrics

Kapil Sharma
Clinical Instructor in Medicine

Rajesh Shrestha
Clinical Instructor in Medicine

Suvash Shrestha
Clinical Instructor in Medicine

Jeffrey Alan Sorokin
Clinical Instructor in Family Medicine

Sylvester C. Sviokla
Clinical Instructor in Medicine

Jeanne W. Swen
Clinical Instructor in Family Medicine

Vinod Thomas
Clinical Instructor in Medicine

Andrew J. Tompkins
Clinical Instructor in Surgery (Urology)

Devin Tsai
Clinical Instructor in Emergency Medicine

George A. Turini
Clinical Instructor in Medical Science

Mark Turshen
Clinical Instructor in Family Medicine

Todd D. Viccione
Clinical Instructor in Medicine

Simon Waghchoure
Clinical Instructor in Medicine

Myles Webster
Clinical Instructor in Pediatrics; Clinical Instructor in Medical Science

Jane Wen
Clinical Instructor in Medicine

Megan Wheelden
Clinical Instructor in Medicine

Kosuke Yasukawa
Clinical Instructor in Medicine

Instructor Research

Claire E. Blevins
Instructor in Psychiatry and Human Behavior (Research)

Silvia Chiang
Instructor in Pediatrics (Research)

Erin Whitney Evans
Instructor in Psychiatry and Human Behavior (Research)

Bong Sook Jhun
Instructor in Medicine (Research)

Karni Moshal
Instructor in Medicine (Research)

Tanbir Najrana
Instructor in Pediatrics (Research)

Christina Nixon
Instructor in Pathology and Laboratory Medicine (Research)

Adjunct Professor

Fatemeh Akhlaghi
Adjunct Professor of Medicine

Thomas F. Anders
Adjunct Professor of Psychiatry and Human Behavior

Oleg Andreev
Adjunct Professor of Radiation Oncology

Kenneth A. Arndt
Adjunct Professor of Dermatology

Andrew W. Artenstein
Adjunct Professor of Medicine; Adjunct Professor of Health Services, Policy and Practice

Seth F. Berkley
Adjunct Professor of Medicine

Charlotte M. Boney
Adjunct Professor of Pediatrics

Phil Brown
Adjunct Professor of Pathology and Laboratory Medicine

Richard A. Brown
Adjunct Professor of Psychiatry and Human Behavior

Carlos Camargo
Adjunct Professor of Pediatrics

Giovanni Camussi
Adjunct Professor of Medicine

Richard Dollase
Adjunct Professor of Medical Science

Robert M. Dowben
Adjunct Professor Emeritus of Molecular Pharmacology, Physiology and Biotechnology

Jane L. Eisen
Adjunct Professor of Psychiatry and Human Behavior

Lynne A. Fontaine
Adjunct Professor of Pediatrics

Michael G. Goldstein
Adjunct Professor of Psychiatry and Human Behavior

Pamina Gorbach
Adjunct Professor of Medicine

Risa Weisberg
Adjunct Professor of Psychiatry and Human Behavior; Adjunct Professor of Family Medicine

Anthony Richard Hayward
Adjunct Professor of Pediatrics

Jiman He
Adjunct Professor of Medicine

Anne L. Hume
Adjunct Professor of Family Medicine

Kerry L. Laplante
Adjunct Professor of Medicine

Arnold M. Ludwig
Adjunct Professor of Psychiatry and Human Behavior

Kenneth Hugh Mayer
Adjunct Professor of Medicine; Adjunct Professor of Epidemiology

Stefan McDonough
Adjunct Professor of Molecular Pharmacology, Physiology and Biotechnology

Francisco Amado Mejia Ortiz
Adjunct Professor of Medicine

Richard G. Moore
Adjunct Professor of Obstetrics and Gynecology

G. Richard Olds
Adjunct Professor of Medicine

Lawrence E. Reinstein
Adjunct Professor of Radiation Oncology

Shmuel Reis
Adjunct Professor of Family Medicine

Yana K. Reshetnyak
Adjunct Professor of Radiation Oncology

Mark J. Rivard
Adjunct Professor of Radiation Oncology

Alan Rothman
Adjunct Professor of Molecular Microbiology and Immunology

Lynn Rothschild
Adjunct Professor of Molecular Biology, Cell Biology and Biochemistry

Gail Skowron
Adjunct Professor of Medicine

Michael D. Stein
Adjunct Professor of Medicine

Robert L. Stout
Adjunct Professor of Psychiatry and Human Behavior

Gregory L. Stuart
Adjunct Professor of Psychiatry and Human Behavior

Mary C. Sullivan
Adjunct Professor of Pediatrics

Lawrence H. Sweet
Adjunct Professor of Psychiatry and Human Behavior

Tracey Taveira
Adjunct Professor of Medicine

Julie S. Taylor
Adjunct Professor of Family Medicine

Peter L. Tilkemeier
Adjunct Professor of Medicine

Thomas F. Tracy
Adjunct Professor of Surgery

Stephen H. Zinner
Adjunct Professor of Medicine

Adjunct Professor Research

Djuro Josic
Adjunct Professor of Medicine (Research)

Adjunct Associate Professor

Rachel Altura
Adjunct Associate Professor of Pediatrics

David B. Baier
Adjunct Associate Professor of Ecology and Evolutionary Biology

Marilyn M. Barbour
Adjunct Associate Professor of Medicine

Esther Choo
Adjunct Associate Professor of Emergency Medicine

Francis J. Cummings
Adjunct Associate Professor of Medicine

R. Davenport
Adjunct Associate Professor of Neuroscience

David Denofrio
Adjunct Associate Professor of Medicine

Jeffrey Dover
Adjunct Associate Professor of Dermatology

Traci C. Green
Adjunct Associate Professor of Emergency Medicine

Thomas J. Guilmette
Adjunct Associate Professor of Psychiatry and Human Behavior

Brian W. Jack
Adjunct Associate Professor of Family Medicine

Yow-Pin Lim
Adjunct Associate Professor of Pathology and Laboratory Medicine

Albert C. Lo
Adjunct Associate Professor of Neurology

Richard Long
Adjunct Associate Professor of Family Medicine

Celia P. MacDonnell
Adjunct Associate Professor of Family Medicine

Keith A. Metzger
Adjunct Associate Professor of Ecology and Evolutionary Biology

Martha Murray
Adjunct Associate Professor of Orthopaedics

Henry Querfurth
Adjunct Associate Professor of Neurology

Thomas E. Rohrer
Adjunct Associate Professor of Dermatology

Marina Tolou-Shams
Adjunct Associate Professor of Psychiatry and Human Behavior

Rajan Wadhawan
Adjunct Associate Professor of Pediatrics

William H. Zywiak
Adjunct Associate Professor of Psychiatry and Human Behavior

Adjunct Associate Professor Research

A. Raymond Frackelton
Adjunct Associate Professor of Pathology and Laboratory Medicine (Research)

Nicola M. Kouttab
Adjunct Associate Professor of Pathology and Laboratory Medicine (Research)

Adjunct Assistant Professor

Joyce A. Alves
Adjunct Assistant Professor of Family Medicine

Motasem A. Al-Yacoub
Adjunct Assistant Professor of Neurology

Melvin Andersen
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Kathleen Askland
Adjunct Assistant Professor of Psychiatry and Human Behavior

Sasha H. Bakhru
Adjunct Assistant Professor of Molecular Pharmacology, Physiology and Biotechnology

Julia G. Brody
Adjunct Assistant Professor of Pathology and Laboratory Medicine

John D. Cahill
Adjunct Assistant Professor of Emergency Medicine

Ayesha Cheema-Hasan
Adjunct Assistant Professor of Pediatrics

Rocco A. Chiappini
Adjunct Assistant Professor of Neurology

Doreen M. Ciancaglini
Adjunct Assistant Professor of Pediatrics

Matthew J. Collins
Adjunct Assistant Professor of Family Medicine

Catherine Degood
Adjunct Assistant Professor of Family Medicine

Antoniu L. Fantana
Adjunct Assistant Professor of Neurology

Adrian Gardner
Adjunct Assistant Professor of Medicine

Jessica Gokee La Rose
Adjunct Assistant Professor of Psychiatry and Human Behavior

Matthew S. Goodwin
Adjunct Assistant Professor of Psychiatry and Human Behavior

Katheleen Hawes
Adjunct Assistant Professor of Pediatrics

Rachel Herz
Adjunct Assistant Professor of Psychiatry and Human Behavior

David Hughes
Adjunct Assistant Professor of Pediatrics

Michael S. Kaminer
Adjunct Assistant Professor of Dermatology

Min-Jong Kang
Adjunct Assistant Professor of Molecular Microbiology and Immunology

Megan M. Kelly
Adjunct Assistant Professor of Psychiatry and Human Behavior

George B. Knight
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Bruce A. Lazarus
Adjunct Assistant Professor of Family Medicine

Collin M. Lines
Adjunct Assistant Professor of Emergency Medicine

William H. McQuade
Adjunct Assistant Professor of Family Medicine

Jeffrey S. Moffit
Adjunct Assistant Professor of Pathology and Laboratory Medicine

Michael Newstein
Adjunct Assistant Professor of Medicine

Anthony Pinto
Adjunct Assistant Professor of Psychiatry and Human Behavior

Brandy Roane
Adjunct Assistant Professor of Psychiatry and Human Behavior

Elizabeth Samuels
Adjunct Assistant Professor of Emergency Medicine

Linda C. Shafer
Adjunct Assistant Professor of Psychiatry and Human Behavior

Timothy D. Shafman
Adjunct Assistant Professor of Obstetrics and Gynecology

James David Sullivan
Adjunct Assistant Professor of Emergency Medicine

Daria A. Szkwarko
Adjunct Assistant Professor of Family Medicine

Golfo Tzilos
Adjunct Assistant Professor of Psychiatry and Human Behavior

Nico Vehse
Adjunct Assistant Professor of Pediatrics

Peter M. Yeracaris
Adjunct Assistant Professor of Family Medicine

Adjunct Instructor

Priya Bansal
Adjunct Instructor in Medicine

Briannan E. Bintz
Adjunct Instructor in Pathology and Laboratory Medicine

Courtney Beard Elias
Adjunct Instructor in Psychiatry and Human Behavior

Laura C. Parajon
Adjunct Instructor in Family Medicine

Zoe Tseng
Adjunct Instructor in Medicine

Visiting Scholar

Laura A. M. Vares
Visiting Scholar in Anthropology

Research Associate

Bin Feng
Research Associate in Epidemiology

Biology

The Biology concentration invites students to study, in depth and in breadth, the science of life and living matter. Whether pursuing the Bachelor of Arts (A.B.) or Science (Sc.B.) in biology, students can expect to learn broadly in the discipline through a selection of courses in three areas: cell and molecular biology, structure and function, and organismal biology. In addition, students pursuing the Sc.B. complete a thematic track through which they gain an in-depth understanding of a particular subfield (such as, Immunopathology; Ecology and Evolutionary Biology; Physiology/Biotechnology; Cell and Molecular Biology; Physical Sciences. The concentration also emphasizes practical skills and experimental design. Concentrators are required to take at least 3 courses with a laboratory or fieldwork component. Within all of these requirements, students have a high degree of flexibility and choice. Broad research opportunities are also available across several departments within the basic sciences as well.

Standard program for the A.B. Biology

The concentration program for the A.B. in Biology consists of four prerequisite courses in math, chemistry, and a statistics course as well as ten courses in biological sciences, including  at least one course in each of the following three areas: Area 1: Cell/Molecular Biology, Area 2: Structure/Function, and Area 3: Organismal Biology.

Prerequisites: 1
Equilibrium, Rate, and Structure
Organic Chemistry
Introductory Calculus, Part I (or placement. MATH 0050/MATH 0060 may be substituted for MATH 0090.)
One of the following:
Introductory Calculus, Part II (or placement)
Advanced Placement Calculus (or equivalent placement)
Or a statistics course, to be approved by the concentration advisor.
Ten Core Courses: 2,4
BIOL 0200The Foundation of Living Systems (Required course; AP credit or similar IB or A-levels accepted, placement test available.)1
The Area requirement must be fulfilled by taking at least one course in each of these groups: 33
Area 1 (Cell/Molecular Biology)
Introductory Biochemistry
Genetics
Cell and Molecular Biology
Introductory Microbiology
Principles of Immunology
Biology of the Eukaryotic Cell
Developmental Biology
Principles of Neurobiology
Area 2 (Structure/Function)
Biological Design: Structural Architecture of Organisms
Invertebrate Zoology
Inquiry in Plant Biology: Analysis of Plant Growth, Reproduction and Adaptive Responses
Principles of Physiology
Biomaterials
Developmental Biology
Biology of Reproduction
Comparative Biology of the Vertebrates
The Brain: An Introduction to Neuroscience
Area 3 (Organismal Biology)
Conservation Medicine
Diversity of Life
The Fossil Record: Life through Time on Earth
The Ecology and Evolution of Infectious Disease
Invertebrate Zoology
Microbes in the Environment
Principles of Ecology
The Evolution of Plant Diversity
Evolutionary Biology
Comparative Biology of the Vertebrates
Environmental Science in a Changing World
Six additional courses chosen from BIOL and/or NEUR offerings for concentrators. At least two at the advanced (1000-2000) level. The Core may include up to two related sciences, with advisor approval.6
Total Credits10
1

 AP scores of 4 or above may substitute Math courses.

2

 At least two biology and/or neuroscience courses must be at the advanced level (between 1000-2999).  EXCLUSIONS: BIOL 0920 series courses, BIOL 1070, & BIOL 1920 series courses.  *Courses numbered below BIOL 0100 do not carry concentration credit.  At least three of the Biology and/or Neuroscience courses must include laboratory or fieldworkBIOL 1950/BIOL 1960, (Directed Research) may be included, but is not required. If a lab project, this can count for ONE of the three lab course requirements, and one advanced course.

3

 No substitutions per above Area list. If a course is listed in more than one area, it may be used to fulfill one of those, the other must be fulfilled by a different course.

4

Biology courses for concentration credit include those numbered greater than 0100 with some exceptions noted within the course descriptions. Courses numbered over 3000 do not count towards Undergraduate requirements either quantity or for concentration.


Honors: Honors in biology requires a thesis and presentation based on a research project (conducted via BIOL 1950/BIOL 1960), and quality grades in the concentration. Guidelines and information on faculty research are available in the Office of Biology Undergraduate Education or found at http://www.brown.edu/academics/biology/undergraduate-education/.

Standard Program for the Sc.B. Biology

The concentration program for the Sc.B. in Biology consists of seven prerequisite courses in math, chemistry, and physics as well as thirteen to fourteen courses in biological sciences, including courses in each of the following three areas: Area 1: Cell/Molecular Biology, Area 2: Structure/Function, and Area 3: Organismal Biology, and the three-course Track.  The biological sciences requirement also requires research (BIOL 1950/BIOL 1960), which should reflect the advanced cluster.

Students pursuing a ScB in Biology have the option to substitute a course for CHEM 0360 (Organic Chemistry) in their background core. For students pursuing the Marine Biology track, an upper level course in Geological Sciences may replace CHEM 0360. For students pursuing all other tracks, BIOL 0280 (Introductory Biochemistry) may serve as the replacement course.  Please note that approval from the concentration advisor is required for these background course substitutions. If the student has already declared, then a revised concentration plan must be submitted and approved via the ASK system. If BIOL 0280 is used as a substitute for CHEM 0360, it cannot be counted as a core course, as a laboratory course, or as an Area 1 course. Students planning to apply to medical or graduate school should seek additional advising (such as from the Health Careers Office) in crafting their course plan.

Prerequisites: 1
Introductory Calculus, Part I (or placement. MATH 0050/MATH 0060 may be substituted for MATH 0090)
Introductory Calculus, Part II
Advanced Placement Calculus
Equilibrium, Rate, and Structure (or IB credit)
Organic Chemistry
Organic Chemistry
Introductory Biochemistry
Basic Physics (or equivalent. PHYS 0050 or ENGN 0030 may be substituted for PHYS 0030.)
Basic Physics (or equivalent. PHYS 0060 or ENGN 0040 may be substituted for PHYS 0040.)
Core Courses: 2, 3
BIOL 0200The Foundation of Living Systems (or placement)1
The Area requirement must be fulfilled by taking at least one course in each of these groups:3
Area 1 (Cell/Molecular Biology)
Introductory Biochemistry
Genetics
Cell and Molecular Biology
Introductory Microbiology
Principles of Immunology
Biology of the Eukaryotic Cell
Developmental Biology
Principles of Neurobiology
Area 2 (Structure/Function)
Biological Design: Structural Architecture of Organisms
Invertebrate Zoology
Inquiry in Plant Biology: Analysis of Plant Growth, Reproduction and Adaptive Responses
Principles of Physiology
Biomaterials
Developmental Biology
Biology of Reproduction
Comparative Biology of the Vertebrates
The Brain: An Introduction to Neuroscience
Area 3 (Organismal Biology)
Conservation Medicine
Diversity of Life
The Fossil Record: Life through Time on Earth
BIOL 0370 - Experimental Evolution
Invertebrate Zoology
Microbes in the Environment
Principles of Ecology
The Evolution of Plant Diversity
Evolutionary Biology
Comparative Biology of the Vertebrates
Environmental Science in a Changing World
Five additional courses chosen from BIOL and/or NEUR offerings for concentrators. Alternatively, students may may include up to two related (non-BIOL/NEUR) sciences suitable for science concentrators. 45
RESEARCH: 5
Typically, two courses in Track is advanced level research (BIOL 1950,1960).1-2
TRACK:
The Track consists of three additional biological sciences courses (not including BIOL 1950/1960 research) that form a Track. Tracks include: Immuno/Pathobiology; Ecology and Evolutionary Biology; Physiology and Biotechnology; Neurobiology; Physical Sciences; Marine Biology; Cell and Molecular Biology. 63
Total Credits13-14
1

 AP scores of 4 or 5 may substitute Math courses.

2

 At least two biology and/or neuroscience courses numbered at the 1000 or 2000 level.  EXCLUSIONS: BIOL 0920 series courses, BIOL 1070,  or BIOL 1920 series courses. *Courses numbered below BIOL 0100 do not carry concentration credit.  At least three of the biology and/or neuroscience courses must include laboratory or fieldwork.  No substitutions per above Area lists. If a course is listed in more than one area, it may be used to fulfill one of those; the other must be fulfilled by a different course

3

Biology courses for concentration credit include those numbered greater than 0100 with some exceptions noted within the course descriptions. Courses numbered over 3000 do not count towards Undergraduate requirements either quantity or for concentration.

4

 See listing at http://biology.brown.edu/bug/ for options.  Related sciences must be above prerequisite level, and suitable for science concentrators.

5

 If substantial research is carried out away from Brown, it must be approved by an appropriate Brown BioMed faculty member but does not carry course credit toward the Core program.

6

At least two, and preferably all three, must be above 1000-level.  Courses used for the cluster, must be approved by an advisor and/or Associate Dean of Biology, Katherine Smith.

Honors: Honors in biology requires a thesis and presentation based on a research project (usually conducted via BIOL 1950/BIOL 1960), and quality grades in the concentration. Guidelines and information on faculty research are available in the Office of Biology Undergraduate Education or at http://www.brown.edu/academics/biology/undergraduate-education/.

Stipulations for Biology Programs:

  1. For double concentrations, no more than two courses may overlap (i.e., be used to meet requirements of both) programs, prerequisites excepted.
  2. No more than two semesters of directed research may be used as concentration credits. Each does count as an individual core towards the program, but only carry one lab credit towards the three required.
  3. A limited number of transfer or study abroad courses may be used within the program, subject to approval of advisor, and Associate Dean of Biology, Katherine Smith.

Health & Human Biology

 

Health and Human Biology is an interdisciplinary concentration that provides a rigorous foundation in the biological sciences with substantive course work in humanities and social sciences within a subfield of Human Health and Disease. The program includes: background courses, biology core courses, a set of theme courses, and a Senior Capstone activity. Background courses provide the essential foundations in chemistry, mathematics, methods, and basic biology. These support the Biology core, which is comprised of a flexible menu of intermediate and advanced courses. A required portion of the Biology core is Genetics, a cornerstone of human biology and its interface with other fields. The Biology core underscores the related coursework within the Health and Disease Theme. The Theme courses are social science and humanities courses that form a cohesive, thoughtful grouping. Theme groupings must be approved by the advisor. A required senior capstone course or activity builds on the program's focus.

 

Program Requirements

REQUIRED BACKGROUND:
Four (4) courses including:
MATH 0090Introductory Calculus, Part I (or equivalent placement)1
OR
Analytic Geometry and Calculus
and Analytic Geometry and Calculus
OR
Introductory Calculus, Part II
Advanced Placement Calculus
CHEM 0330Equilibrium, Rate, and Structure1
BIOL 0200The Foundation of Living Systems1
Statistics course chosen with advisor's help.1
CORE PROGRAM:
In addition to the stated background in Chemistry, Math, Biology and Statistics, five (5) Biology plus four (4) coherently-grouped Theme courses, plus a Senior-Year Capstone course or project. (See description of Capstone at link below this table).
BIOLOGY:
Five (5) courses, including:5
Genetics, which can be fulfilled in the following ways:
Genetics
-OR-
Evolutionary Biology
and Cell and Molecular Biology
-OR-
Evolutionary Biology
and Introductory Microbiology
-OR-
Evolutionary Biology
and Introductory Biochemistry
Select one course in structure/function/development such as:
Biological Design: Structural Architecture of Organisms
Principles of Physiology
Developmental Biology
Animal Locomotion
Comparative Biology of the Vertebrates
The Brain: An Introduction to Neuroscience
One course in organismal/population biology such as:
BIOL 0370 - Experimental Evolution
The Ecology and Evolution of Infectious Disease
Vertebrate Evolution and Diversity
Biological Design: Structural Architecture of Organisms
Invertebrate Zoology
Microbes in the Environment
Principles of Ecology
Evolutionary Biology
Conservation Biology
Comparative Biology of the Vertebrates
Environmental Science in a Changing World
Or a course from the NEUR 1940 series
c) Two Biology or Neuroscience courses that relate to and support the chosen theme course grouping. At least one must be at the advanced level.
THEME: With the advisor's assistance, a theme is chosen and a cohesive set of courses ae selected from ourside of Biology. See Notes below:4
SENIOR CAPSTONE ACTIVITY: Must be conducted during the senior year, fulfilled by one of the following, and related to the student's chosen theme: 1
1) Advisor approved senior seminar or advanced course related to the theme
2) One semester of independent research/independent study (BIOL 1950 or BIOL 1960); in the case of a senior honors thesis, both BIOL 1950 and BIOL 1960 can be used as the capstone.
3) An appropriate internship with a scholarly context can be used if coupled with a semester of indpendent study mentored by a Brown faculty member.
Total Credits14

THEME: 

  • Approved courses must be above the introductory level and at least one must be 1000-level or above. 
  • No more than TWO courses from a given department may be included in the theme portion.
  • Students can choose from SIX theme options: 1) Health Behavior, 2) Environmental Health, 3) Global/International Health, 4) Social Context of Health and Disease, 5) Health Systems, Structure and Policy, and 6) Women's/Children Health.
  • NOTE: Beginning with the Class of 2020: Health Systems, Structure and Policy and Women's/Children Health will be eliminated.
  • Students will then select from FOUR theme options: 1) Health Behavior, 2) Environmental Health, 3) Global/International Health, 4) Social Context of Health and Disease. 

CAPSTONE: See http://www.brown.edu/academics/biology/undergraduate-education/ for more information on the Capstone Activity.

HONORS:  See more information about Honors at http://www.brown.edu/academics/biology/undergraduate-education/.

Applied Mathematics-Biology

The Applied Math - Biology concentration recognizes that mathematics is essential to address many modern biological problems in the post genomic era. Specifically, high throughput technologies have rendered vast new biological data sets that require novel analytical skills for the most basic analyses. These technologies are spawning a new "data-driven" paradigm in the biological sciences and the fields of bioinformatics and systems biology. The foundations of these new fields are inherently mathematical, with a focus on probability, statistical inference, and systems dynamics. These mathematical methods apply very broadly in many biological fields including some like population growth, spread of disease, that predate the genomics revolution. Nevertheless, the application of these methods in areas of biology from molecular genetics to evolutionary biology has grown very rapidly in with the availability of vast amounts of genomic sequence data. Required coursework in this program aims at ensuring expertise in mathematical and statistical sciences, and their application in biology. The students will focus in particular areas of biology. The program culminates in a senior capstone experience that pairs student and faculty in creative research collaborations. 

Standard program for the Sc.B. degree

Required coursework in this program aims at ensuring expertise in mathematical and statistical sciences, and their application in biology. The students will focus in particular areas of biology. The program culminates in a senior capstone experience that pairs student and faculty in creative research collaborations. Applied Math – Biology concentrators are prepared for careers in medicine, public health, industry and academic research.

Required Courses:

Students are required to take all of the following courses.

MATH 0090Introductory Calculus, Part I1
MATH 0100Introductory Calculus, Part II1
or MATH 0170 Advanced Placement Calculus
MATH 0180Intermediate Calculus (or equivalent placement)1
MATH 0520Linear Algebra1
or MATH 0540 Honors Linear Algebra
CHEM 0330Equilibrium, Rate, and Structure 11
PHYS 0030Basic Physics1
or PHYS 0050 Foundations of Mechanics
Select one of the following sequences:2
Applied Ordinary Differential Equations
and Applied Partial Differential Equations I
Methods of Applied Mathematics I, II
and Methods of Applied Mathematics I, II
APMA 1650Statistical Inference I1
or APMA 1655 Statistical Inference I
APMA 1070Quantitative Models of Biological Systems1
APMA 1080Inference in Genomics and Molecular Biology1
BIOL 0200The Foundation of Living Systems (or equivalent)1
Additional Courses
In addition to required courses listed above, students must take the following:
Two additional courses in Applied Math or Biology. At least one of these must be a directed research course, e.g. a senior seminar or independent study in Applied Math or a directed research/independent study in Biology. For example: 12
A course from the APMA 1930 series
A course from the APMA 1940 series
Independent Study
Directed Research/Independent Study
Directed Research/Independent Study
Four classes in the biological sciences agreed upon by the student and advisor. These four courses should form a cohesive grouping ina specific area of emphasis, at least two of which should be at the 1000-level. Some example groupings are below: 4
Areas of Emphasis and Suggested Courses:
Some areas of possible emphasis for focusing of elective courses are listed below. Given the large number of course offerings in the biosciences and neuroscience, students are free to explore classes in these areas that are not listed below. However, all classes must be approved by the concentration advisor.
Biochemistry
Introductory Biochemistry
Advanced Biochemistry
Organic Chemistry
Chemical Biology
Biotechnology and Physiology
Principles of Physiology
Cell Physiology and Biophysics
and/or appropriate bioengineering courses, such as:
Polymer Science for Biomaterials
Biomaterials
Tissue Engineering
Stem Cell Engineering
Synthetic Biological Systems
Ecology, Evolution, and Genetics
Invertebrate Zoology
and Evolutionary Biology
Principles of Ecology
and The Evolution of Plant Diversity
Genetics
Experimental Design in Ecology
Population Genetics
Human Population Genomics
Molecular Genetics
Neuroscience
Mathematical Methods in the Brain Sciences
Neurosciences courses: See https://www.brown.edu/academics/neuroscience/undergraduate/neuroscience-concentration-requirements
Cell Physiology and Biophysics
Topics in Signal Transduction
Synaptic Transmission and Plasticity
Total Credits18
1

Students whose independent study is expected to be in an experimental field are strongly encouraged to take APMA 1660, which covers experimental design and the analysis of variance (ANOVA), a method commonly used in the analysis of experimental data.

Honors

Requirements and Process: Honors in the Applied Math-Biology concentration is based primarily upon an in-depth, original research project carried out under the guidance of a Brown (and usually Applied Math or BioMed) affiliated faculty advisor.  Projects must be conducted for no less than two full semesters, and student smust register for credit for the project via APMA 1970 or BIOL 1950/BIOL 1960 or similar independent study courses.  The project culminates in the writing of a thesis which is reviewed by the thesis advisor and a second reader.  It is essential that the student have one advisor from the biological sciences and one in Applied Mathematics.  The thesis work must be presented in the form of an oral presentation (arranged with the primary thesis advisor) or posted at the annual Undergraduate Research Day in either Applied Mathematics or Biology.  For information on registering for BIOL 1950/BIOL 1960, please see https://www.brown.edu/academics/biology/undergraduate-education/undergraduate-research

Excellence in grades within the concentration as well as a satisfactory evaluation by the advsors are also required for Honors.  The student's grades must place them within the upper 20% of their cohort, in accordance with the university policy on honors.  Honors recipients typically maintain a Grade Point Average of 3.4 or higher in the concentration.  However, in the case of outstanding independent research as demonstrated in the thesis and supported by the Thesis Committee, candidates with a GPA between 3.0 an 3.4 will be considered and are encouraged to apply.

The deadline for applying to graduate with honors in the concentration are the same as those of the biology concentrations.  However, students in the joint concentration must inform the undergraduate chair in Applied Mathematics of their intention to apply for honors by these dates.

Biochemistry & Molecular Biology

How does life work at the molecular level? This question is at the core of the concentration program Biochemistry and Molecular Biology. In earlier years of this discipline, the focus was on structure and function of proteins, nucleic acids, lipids, carbohydrates and small molecules such as vitamins. Today the logical approach and tools of biochemical science are being expanded to new areas in neuroscience, developmental biology, immunology, pharmacology and synthetic biology (the design of analogs of biological systems). Training in biochemistry begins with a foundation in mathematics, physics, chemistry and biology. Some courses offered in other departments, including engineering, geology and computer science, are also useful. A key component of this program is the year of hands-on research carried out in collaboration with a faculty member here at Brown. Faculty sponsors are drawn from both the Chemistry Department and the Division of Biology and Medicine, and include basic science and clinical faculty.

Standard program for the Sc.B. degree

Students must take twenty courses in biology, chemistry, mathematics, and physics, including the following core requirements, some of these may be fulfilled with AP credits. Students are expected to take courses that will count toward the concentration ABC/NC.  Students should discuss the S/NC option with their concentration advisor if circumstances warrant consideration.  Students should not register S/NC for a concentration course without advisor pre-approval.

Three courses in mathematics, statistics and/or computer science, typically including MATH 0090, MATH 0100, or equivalent) 13
Two courses in physics, typically: 12
Basic Physics
Foundations of Mechanics
Introduction to Engineering
Basic Physics
Foundations of Electromagnetism and Modern Physics
Dynamics and Vibrations
Three courses in physical and organic chemistry:3
Equilibrium, Rate, and Structure
Organic Chemistry
One course in biophysical or related chemistry:1
Biophysical and Bioinorganic Chemistry
Inorganic Chemistry
Instrumental Analysis with Environmental Applications
-or-
Instrumental Analysis with Environmental Applications
Three courses in biochemistry:3
Introductory Biochemistry
Advanced Biochemistry
Chemical Biology
Biochemistry
Select two semester courses of independent research approved by a concentration advisor:2
Directed Research/Independent Study
-or-
Undergraduate Research
Suggested Elective Courses:
Students are required to take six (6) elective courses: four (4) taken from the chart below and two (2) from any science or mathematics course relevant to biochemistry, cell and molecular biology from the suggested courses below:4
Biology Electives:
The Foundation of Living Systems
The Ecology and Evolution of Infectious Disease
Microbes in the Environment
Genetics
Cell and Molecular Biology
Principles of Immunology
Principles of Physiology
Biology of the Eukaryotic Cell
Polymer Science for Biomaterials
Cell Physiology and Biophysics
Topics in Signal Transduction
Protein Biophysics and Structure
Synthetic Biological Systems
Physiological Pharmacology
Cancer Biology
Developmental Biology
Biology of Reproduction
Innate Immunity
Molecular Genetics
Virology
Development of Vaccines to Infectious Diseases
Drug and Gene Delivery
Neuroscience Electives: 2
The Brain: An Introduction to Neuroscience
Biology of Hearing
Principles of Neurobiology
Introduction to Neurogenetics
Neuropharmacology and Synaptic Transmission
Chemistry Electives:
Inorganic Chemistry
Physical Chemistry: Quantum Chemistry
Physical Chemistry: Thermodynamics and Statistical Mechanics
Computational Tools in Biochemistry and Chemical Biology
Chemical Biology
Biochemistry
Advanced Organic Chemistry
Computer Science Electives:
Computational Molecular Biology
Quantitative Science or Mathematics Electives:2
Select two electives from any quantitative science or mathematics course relevant to biochemistry (including courses on the preceding list) and approved by a concentration advisor.
Total Credits20
1

Note that the mathematics and physics requirements may be satisfied by Advanced Placement credit.

2

or any NEUR course in Cell, Genetics, Molecular Biology, or Development.

Honors Requirements for Biochemistry

All ScB Biochemistry concentrators are candidates for Honors; no separate application is necessary.

The requirements for Honors in Biochemistry are:

* A strong grade record in concentration courses. This means a grade point average for the concentration that is higher than 3.25.

* Two semesters of Independent Study (CHEM 0970, CHEM 0980 or equivalent. Guidelines and requirements associated with Independent Study are in the Undergraduate Concentration Handbook which can be found at the department website.

* A Thesis in a form approved by the research advisor, and recommended by the research advisor.  Additional information about thesis guidelines will be provided by the Concentration Advisor in the first half of the fall semester.

Biomedical Engineering

The Sc.B. program in Biomedical Engineering is accredited by the Engineering Accreditation Commission of ABET, http://www.abet.org/. It is jointly offered by the School of Engineering and the Division of Biology and Medicine as an interdisciplinary concentration designed for students interested in applying the methods and tools of engineering to the subject matter of biology and the life sciences. The education objectives of the Biomedical Engineering program are to prepare graduates: (1) to be employed in careers of useful service to society, including scientific and technical areas within medicine, industry, and health care delivery; (2) to demonstrate the ability to apply the basic principles of engineering and science, as well as problem solving skills and critical thinking, to a broad spectrum of biomedical engineering problems; (3) to demonstrate their ability to work in teams, and to effectively communicate and understand the broad social, ethical, economic and environmental consequences of their lifelong education. The student outcomes of this program are the (a) - (k) Student Outcomes as defined by the "ABET Criteria for Accrediting Engineering Programs (available online at http://www.abet.org/accreditation-criteria-policies-documents/)." The Biomedical Engineering concentration shares much of the core with the other engineering programs, but is structured to include more courses in biology and chemistry, and a somewhat different emphasis in mathematics.

The requirements regarding Mathematics, Advanced Placement, Transfer Credit, Substitutions for Required Courses, and Humanities and Social Science Courses are identical to those of the Sc.B. degree programs in Engineering. Please refer to the Engineering section of the University Bulletin for explicit guidelines.  

The Biomedical Engineering concentration shares much of the core with the other engineering programs, but is structured to include more courses in biology and chemistry, and a somewhat different emphasis in mathematics. 

 Standard program for the Sc.B. degree

1. Core Courses
ENGN 0030Introduction to Engineering1
ENGN 0040Dynamics and Vibrations1
ENGN 0510Electricity and Magnetism1
ENGN 0720Thermodynamics1
ENGN 0810Fluid Mechanics1
BIOL 0200The Foundation of Living Systems 11
or NEUR 0010 The Brain: An Introduction to Neuroscience
BIOL 0800Principles of Physiology1
CHEM 0330Equilibrium, Rate, and Structure1
CHEM 0350Organic Chemistry1
MATH 0190Advanced Placement Calculus (Physics/Engineering) 21
or MATH 0170 Advanced Placement Calculus
MATH 0200Intermediate Calculus (Physics/Engineering)1
or MATH 0180 Intermediate Calculus
or MATH 0350 Honors Calculus
APMA 0330Methods of Applied Mathematics I, II1
or APMA 0350 Applied Ordinary Differential Equations
APMA 0650Essential Statistics1
or APMA 1650 Statistical Inference I
2. Upper Level Biomedical Engineering Curriculum
ENGN 1110Transport and Biotransport Processes1
ENGN 1210Biomechanics1
ENGN 1230Instrumentation Design1
ENGN 1490Biomaterials1
Three Additional upper Level Biomedical Engeinnering Courses3
Select at least one of the following: 3
Tissue Engineering
Neuroengineering
Analytical Methods in Biomaterials
Biomedical Optics
Molecular and Cell Biology for Engineers
Cancer Nanotechnology
Select at most two of the following: 4
Stem Cell Engineering
Synthetic Biological Systems
Animal Locomotion
Drug and Gene Delivery
Techniques in Molecular and Cell Science
3. Capstone Design 5
ENGN 1930LBiomedical Engineering Design, Research and Modeling1
* In addition to program requirements above, students must take four courses in the humanities and social sciences.
Total Credits21
1

Students with advanced Biology backgrounds may replaced with BIOL 0470, BIOL 0530, or other Biology courses.

2

Students with advanced math backgrounds may replace with CHEM 0360.

3

 Or other advanced bioengineering courses (e.g., ENGN 1510 and ENGN 1520), subject to concentration advisor approval.

4

Or other advanced bioengineering courses, subject to concentration advisor approval.

5

Biomedical engineering students are also encouraged to pursue independent research with faculty members in the School of Engineering or the Division of Biology & Medicine.

Biophysics

Biophysics is that it is a quantitative science that requires a significant level of competence in physics, chemistry, mathematics, and biology. These areas therefore form the required background coursework for this program, and serve as a springboard to an advanced focus, developed in consultation with a concentration advisor. Advanced foci may include structure-function relations of macromolecules, biomechanics of cell cytoskeleton, biotechnology for drug and gene delivery, molecular mechanisms of membrane transport, sensory signal transduction, for examples. The program also requires a capstone research project that reflects this focus and may be drawn from collaborative research opportunities offered by faculty in biology, chemistry, or physics departments. 

Additional detailed information about the field of Biophysics may be found at: http://www.biophysics.org/AboutUs/Biophysics/tabid/517/Default.aspx .

Standard program for the Sc.B. degree

Requirements

Select one of the following Series:2
Foundations of Mechanics
and Foundations of Electromagnetism and Modern Physics
Analytical Mechanics
and Introduction to Relativity and Quantum Physics
PHYS 0470Electricity and Magnetism1
CHEM 0330Equilibrium, Rate, and Structure1
CHEM 0350Organic Chemistry1
CHEM 0360Organic Chemistry1
Select one of the following:1
Biophysical and Bioinorganic Chemistry
Physical Chemistry: Quantum Chemistry
Thermodynamics and Statistical Mechanics
Biological Physics
MATH 0100Introductory Calculus, Part II (or equivalent)1
MATH 0180Intermediate Calculus (or equivalent)1
BIOL 0200The Foundation of Living Systems1
Select two additional biology courses chosen with approval of the advisor. Examples include courses in:2
Cell Biology
Cell and Molecular Biology
Biology of the Eukaryotic Cell
Protein Biophysics and Structure
Physiology
Principles of Physiology
Cell Physiology and Biophysics
Synaptic Transmission and Plasticity
Principles of Neurobiology
Pharmacology
Physiological Pharmacology
Biotechnology
Polymer Science for Biomaterials
Biomaterials
Tissue Engineering
Select six additional intermediate or advanced level courses, chosen from biology (e.g., biochemistry, genetics, physiology, physics, chemistry, and/or computer sciences and mathematics). Examples include:6
Biology
Introductory Biochemistry
Genetics
Principles of Physiology
Synaptic Transmission and Plasticity
Physics
Advanced Classical Mechanics
Experiments in Modern Physics
Quantum Mechanics A
Quantum Mechanics B
Biological Physics
Mathematics
Linear Algebra
Applied Mathematics
Methods of Applied Mathematics I, II
Methods of Applied Mathematics I, II
Applied Ordinary Differential Equations
Applied Partial Differential Equations I
Chemistry
Chemical Biology
Advanced Organic Chemistry
A course from the CHEM 1560 series.
Select at least one semester (two are recommended) of Directed Research 1
Biology
Directed Research/Independent Study
Chemistry
Undergraduate Research
Physics
Senior Conference Course
Total Credits19

Computational Biology

Computational biology involves the analysis and discovery of biological phenomena using computational tools, and the algorithmic design and analysis of such tools. The field is widely defined and includes foundations in computer science, applied mathematics, statistics, biochemistry, molecular biology, genetics, ecology, evolution, anatomy, neuroscience, and visualization. 

Students may pursue a Bachelor of Arts or a Bachelor of Science. Students pursuing the ScB have the option of electing a concentration in Computational Biology with one of three focus areas: Computer Sciences, Biological Sciences, or Applied Mathematics & Statistics. Both programs require a senior capstone experience that pairs students and faculty in creative research collaborations.

Standard program for the A.B. degree

Prerequisites:2
Introductory Calculus, Part II
Advanced Placement Calculus
The Foundation of Living Systems
General Core Requirements: Biology2
Genetics
Introductory Biochemistry
Cell and Molecular Biology
General Core Requirements: Chemistry1
Equilibrium, Rate, and Structure
Organic Chemistry
General Core Requirements: Computer Science2
Introduction to Object-Oriented Programming and Computer Science
and Introduction to Algorithms and Data Structures
OR
Computer Science: An Integrated Introduction
and Computer Science: An Integrated Introduction
OR
Accelerated Introduction to Computer Science
and Computer Science: An Integrated Introduction
and Introduction to Software Engineering
and Introduction to Computer Systems
and Theory of Computation
General Core Requirements: Probability & Statistics1
Statistical Inference I
OR
Probability and Computing
OR
Probability
Comp Bio Core Course Requirements4
Computational Molecular Biology
Inference in Genomics and Molecular Biology
AND two of the following:
Algorithmic Foundations of Computational Biology
Population Genetics
Human Population Genomics
Machine Learning
Computational Probability and Statistics
Statistical Inference II
Additional course with Director approval
Total Credits12

University Writing Requirement:

As part of Brown’s writing requirement, all students must demonstrate that they have worked on their writing both in their general studies and their concentration. There are a number of ways for Computational Biology concentrators to fulfill these requirements:

  • Enrolling in an independent study: CSCI 1970, BIOL 1950, APMA 1970
  • Writing an Honors Thesis
  • Taking a “WRIT” course in the final two years

Capstone Experience

Students enrolled in the computational biology concentration will complete a research project in their senior year under faculty supervision.  The themes of such projects evolve with the field and the technology, but should represent a synthesis of the various specialties of the program. The requirements are either one semester of reading and research with a CCMB Faculty member or approved advisor, or a 2000-level Computational Biology course.

Standard program for the Sc.B. degree

Prerequisites
MATH 0100Introductory Calculus, Part II (or equivalent)1
or MATH 0170 Advanced Placement Calculus
BIOL 0200The Foundation of Living Systems (or equivalent)1
General Core Course Requirements: Biology
BIOL 0470Genetics (prerequisite BIOL 0200 or equivalent)1
BIOL 0280Introductory Biochemistry1
or BIOL 0500 Cell and Molecular Biology
General Core Requirements: Chemisty
CHEM 0330Equilibrium, Rate, and Structure1
or CHEM 0350 Organic Chemistry
General Core Requirements: Computer Science2-4
Introduction to Object-Oriented Programming and Computer Science
and Introduction to Algorithms and Data Structures
OR
Computer Science: An Integrated Introduction
and Computer Science: An Integrated Introduction
OR
Accelerated Introduction to Computer Science
and Computer Science: An Integrated Introduction
and Introduction to Software Engineering
and Introduction to Computer Systems
CSCI 0220Introduction to Discrete Structures and Probability1
General Core Requirements: Probability & Statistics
APMA 1650Statistical Inference I1
or CSCI 1450 Probability and Computing
or MATH 1610 Probability
General Core Requirements: Computational Biology
CSCI 1810Computational Molecular Biology1
APMA 1080Inference in Genomics and Molecular Biology1
Capstone Experience1
Directed Research/Independent Study
Individual Independent Study
Six courses in one of the following three tracks:6
Computer Science Track:
Three of the following:
Introduction to Computer Graphics
Database Management Systems
Artificial Intelligence
Probabilistic Methods in Computer Science
Design and Analysis of Algorithms
or other Computer Science courses approved by the concentration advisor
Three of the following:
Introduction to Computer Systems
Introduction to Software Engineering
Algorithmic Foundations of Computational Biology
Statistical Methods in Bioinformatics, I
Statistical Inference II
Population Genetics
Human Population Genomics
Computational Probability and Statistics
Biological Sciences track
At least four courses comprising a coherent theme in one of the following areas: Biochemistry, Ecology, Evolution, or Neurobiology.
AND select two courses from the following:
Algorithmic Foundations of Computational Biology
Statistical Methods in Bioinformatics, I
Statistical Inference II
Population Genetics
Human Population Genomics
Computational Probability and Statistics
Applied Mathematics & Statistics Track:
At least three courses from the following:
Statistical Inference II
Computational Probability and Statistics
Artificial Intelligence
Methods of Applied Mathematics I, II
and Methods of Applied Mathematics I, II
OR
Applied Partial Differential Equations I
and Applied Ordinary Differential Equations
At least three of the following:
Population Genetics
Algorithmic Foundations of Computational Biology
Statistical Methods in Bioinformatics, I
Quantitative Models of Biological Systems
Human Population Genomics
Total Credits18-20

Honors:

 In order to be considered a candidate for honors, students will be expected to maintain an outstanding record, with no "C's" in concentration courses and with a minimum of an "A-" average in concentration courses. In addition, students should take at least one semester, and are strongly encouraged to take 2 semesters, of reading and research with a CCMB faculty member or approved advisor. Students must submit to a public defense of their theses to be open to the CCMB community.

  • Students seeking honors are advised to choose a Thesis Advisor prior to the end of their Junior year
  • Students must complete the Registration form for Comp Bio and submit it to CCMB@BROWN.EDU

Any deviation from these rules must be approved by the director of undergraduate studies, in consultation with the student's advisor.

Marine Biology

Note: This concentration program is being phased out for Class of 2017 students and greater in favor of a track program within the Sc.B. in Biology.

Standard program for the Sc.B. degree

CHEM 0330Equilibrium, Rate, and Structure (or equivalent)1
PHYS 0030Basic Physics (or equivalent)1
PHYS 0040Basic Physics (or equivalent)1
MATH 0090Introductory Calculus, Part I (or equivalent)1
MATH 0100Introductory Calculus, Part II (or equivalent)1
Two additional courses in physics, chemistry, mathematics, applied mathematics, computer science, engineering, or geological sciences, as approved by advisor.2
Select four of the following biology courses:4
Invertebrate Zoology
Principles of Ecology
Introductory Microbiology
Principles of Physiology
Comparative Animal Physiology
Marine Biology
Comparative Biology of the Vertebrates
Select three additional biology courses, highly recommended are:3
The Foundation of Living Systems
Genetics
Evolutionary Biology
Cell and Molecular Biology
Comparative Animal Physiology
Developmental Biology
Evolutionary Genetics
Experimental Design in Ecology
Conservation Biology
Environmental Science in a Changing World
Select two of the following group of related science courses: 12
Introduction to Scientific Computing and Problem Solving
Introduction to Oceanography
Fossil Record
Estuarine Oceanography
Paleoceanography
Ocean Biogeochemical Cycles
Stratigraphy and Sedimentation
Global Environmental Remote Sensing
Quantitative Elements of Physical Hydrology
Organic Chemistry
Organic Chemistry
An approved course in statistics
BIOL 1950/1960Directed Research/Independent Study (conducted at Brown or an approved marine lab or field station.) 21
Total Credits17
1

Or substitutions as approved by the concentration advisor.

2

A summer or semester at a field station is recommended.  Please note that some recommended courses are offered every other year; others have limited enrollment and require early sign-up.